2020 in amphibian paleontology

List of years in amphibian paleontology
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This list of fossil amphibians described in 2020 is a list of new taxa of fossil amphibians that were described during the year 2020, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2020.

New taxa

Temnospondyli

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Benthosuchus lukyanovi [1]	Sp. nov	Valid	Morkovin	Early Triassic		Russia (  Vologda Oblast)	A benthosuchid
Diploseira [2]	Gen. et comb. nov	Valid	Dilkes	Early Permian	Archer City Formation	United States (  Texas)	A dissorophid temnospondyl; a new genus for "Dissorophus" angustus.
Korkonterpeton [3]	Gen. et sp. nov	Valid	Werneburg, Štamberg & Steyer	Permian (Asselian to Artinskian)	Prosečné	Czech Republic	A stereospondylomorph temnospondyl of uncertain phylogenetic placement. The type species is K. kalnense.
Palodromeus [4]	Gen. et sp. nov	Valid	Schoch, Henrici & Hook	Late Carboniferous	Allegheny Formation	United States (  Ohio)	An olsoniform dissorophoid temnospondyl. Genus includes new species P. bairdi.
Rastosuchus [5]	Gen. et sp. nov	Valid	Dias, Dias-da-Silva & Schultz	Permian	Rio do Rasto Formation	Brazil	A rhinesuchid. The type species is R. hammeri.

Allocaudata

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Yaksha [6]	Gen. et sp. nov	Valid	Daza et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An albanerpetontid. The type species is Y. perettii.

Anurans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Calyptocephalella sabrosa [7]	Sp. nov	Valid	Muzzopappa et al.	Paleocene (Danian)	Salamanca Formation	Argentina	A frog. Originally described as a species of Calyptocephalella , but subsequently transferred to the genus Xerocephalella. [8]
Ceratophrys sagani [9]	Sp. nov	Valid	Barcelos et al.	Probably late Pleistocene–early Holocene		Brazil	A South American horned frog.
Kururubatrachus [10]	Gen. et sp. nov	In press	Agnolin et al.	Early Cretaceous (Aptian)	Crato	Brazil	A neobatrachian frog resembling extant members of Hyloidea. Genus includes new species K. gondwanicus.

Caudata

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Balveherpeton [11]	Gen. et sp. nov	In press	Skutschas, Kolchanov & Schwermann	Early Cretaceous (Barremian–Aptian)		Germany	A salamandroid salamander. Genus includes new species B. hoennetalensis.
Egoria [12]	Gen. et sp. nov	Valid	Skutschas et al.	Middle Jurassic (Bathonian)	Itat Formation	Russia (  Krasnoyarsk Krai)	A stem-salamander. The type species is E. malashichevi.
Palaeoproteus miocenicus [13]	Sp. nov	Valid	Vasilyan & Yanenko	Miocene (Vallesian)		Austria   Ukraine	A salamander belonging to the family Batrachosauroididae

Others

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Brittagnathus [14]	Gen. et sp. nov	Valid	Ahlberg & Clack	Devonian (Famennian)	Britta Dal	Greenland	A basal tetrapod. The type species is B. minutus.
Leptoropha minima [15]	Sp. nov	Valid	Bulanov	Permian		Russia (  Tatarstan)	A member of Seymouriamorpha
Microphon terminalis [16]	Sp. nov	Valid	Bulanov	Late Permian		Russia (  Vladimir Oblast)	A member of Seymouriamorpha belonging to the family Kotlassiidae.
Seroherpeton [17]	Gen. et sp. nov	Valid	Chen & Liu	Permian (Wuchiapingian)	Sunjiagou	China	A member of Embolomeri. The type species is S. yangquanensis.
Steenerpeton [18]	Gen. et sp. nov	Valid	Mann et al.	Carboniferous (Pennsylvanian)	Joggins	Canada (  Nova Scotia)	A recumbirostran "microsaur". Genus includes new species S. silvae.

Research

• A study on the evolution of terrestrial locomotion in early tetrapods, based on data from 40 three-dimensionally preserved humeri from extinct tetrapodomorphs spanning the fin-to-limb transition, is published by Dickson et al. (2020). ^[19]
• A study aiming to determine the potential significance of tides for the evolution of bony fish and early tetrapods from the Late Silurian to early Late Devonian is published by Byrne et al. (2020). ^[20]
• A fibula of a member of the family Crassigyrinidae (otherwise known from the Viséan) is described from the Tournaisian of Blue Beach (Nova Scotia, Canada) by Lennie, Mansky & Anderson (2020). ^[21]
• A study on the bone histology of Dvinosaurus campbelli is published by Uliakhin, Skutschas & Saburov (2020). ^[22]
• A study evaluating the effects of ontogenetic disparity of known trematopid specimens on reconstructions of the phylogenetic relationships of trematopids is published by Gee (2020). ^[23]
• Redescription of Actiobates peabodyi, including an updated description of the skull and the first description of the postcranial skeleton, is published by Gee & Reisz (2020). ^[24]
• A study on the suture pattern in the skull and on the mandible anatomy of Cacops aspidephorus is published by Anderson, Scott & Reisz (2020). ^[25]
• New amphibamiform specimen with exceptionally preserved lissamphibian-like integumentary structures, including the first evidence of toepad structures in a temnospondyl body fossil, is described from the Mazon Creek fossil beds by Mann & Gee (2020). ^[26]
• Description of the anatomy of the skull of Pasawioops mayi, and a study on the ontogeny of this taxon, is published by Atkins et al. (2020). ^[27]
• A study on growth patterns in Doleserpeton annectens, as indicated by bone histology, is published by Gee, Haridy & Reisz (2020). ^[28]
• A study on a specimen of Benthosuchus korobkovi from the Olenekian of Russia affected by a neoplastic bone lesion in its jaw, representing the earliest case of such lesion in a tetrapod reported so far, is published by Novikov et al. (2020), who propose a non-odontogenic osteoma as the most likely diagnosis. ^[29]
• Redescription and a study on the phylogenetic relationships of Aphaneramma kokeni is published by Maisch (2020), who considers A. kokeni to be a valid taxon. ^[30]
• A study on the impact of local climatic and environmental conditions on growth patterns of the skeleton of Panthasaurus maleriensis is published by Teschner et al. (2020). ^[31]
• Evidence of the presence of five metacarpals in a specimen of Metoposaurus krasiejowensis from the Upper Triassic of Poland is presented by Konietzko-Meier et al. (2020), who interpret this finding as evidence of pentadactyly of the manus of M. krasiejowensis, showing that the presence of a five-digit manus among Temnospondyli was possible. ^[32]
• New fossil material of albanerpetontids is described from the lower Campanian Aguja Formation (Texas, United States) by Wick (2020), who interprets this finding as indicating that albanerpetontids were locally abundant there and also widespread throughout much of the Western Interior of North America by early Campanian time. ^[33]
• A review of the caecilian fossil record is published by Santos, Laurin & Zaher (2020). ^[34]
• New specimen of Triassurus sixtelae is described from the Triassic of Kyrgyzstan by Schoch, Werneburg & Voigt (2020), who identify this species as the oldest known stem-group salamander. ^[35]
• A study on the skeletal anatomy and phylogenetic relationships of Chunerpeton tianyiense is published by Rong et al. (2020). ^[36]
• A study on the diversity of skull shape in extant and fossil ribbed and crocodile newts, the relationship between their skull shape and ecological and reproductive traits, and its implications for the knowledge of the ecology of Chelotriton , is published by Pogoda et al. (2020). ^[37]
• New pipimorph fossil material, providing new information on the morphological diversity among the earliest pipimorphs in South America, is described from the Aptian Crato Formation (Brazil) by Báez, Muzzopappa & Moura (2020), who also redescribe Cratopipa novaolindensis. ^[38]
• Right ilium and a skull bone of a frog belonging to the genus Calyptocephalella are reported from the Eocene (Bartonian) La Meseta Formation (Antarctica) by Mörs, Reguero & Vasilyan (2020), representing the first record of a lissamphibian in Antarctica reported so far. ^[39]
• Partial humerus of a member of the genus Eleutherodactylus is described from the Oligocene San Sebastian Formation (Puerto Rico) by Blackburn et al. (2020), representing the earliest fossil frog from any Caribbean island reported so far. ^[40]
• Redescription of the anatomy and a study on the phylogenetic relationships of Eldeceeon rolfei is published by Ruta, Clack & Smithson (2020). ^[41]
• A study on the long bone histology, growth rate and the timing of the attainment of sexual maturity in seymouriamorphs is published by Jordi Estefa et al. (2020). ^[42]
• A study on the anatomy of the braincase and otic capsule of Seymouria is published by Bazzana et al. (2020). ^[43]
• Description of new postcranial material of Seymouria from the Richards Spur locality (Oklahoma, United States), and a study on bone histology, life histories and evolution of terrestriality of seymouriamorphs, is published by Bazzana et al. (2020). ^[44]
• A study on the anatomy of the skull of Euryodus dalyae, providing new information on the anatomy of the braincase and mandible, is published by Gee, Bevitt & Reisz (2020). ^[45]
• Description of the anatomy of the braincase and stapes of Diadectes absitus is published by Klembara et al. (2020). ^[46]

References

1. B. I. Morkovin (2020). "A new benthosuchid (Amphibia: Temnospondyli) from the Lower Triassic of the Moscow Syncline". Paleontological Journal. 54 (4): 401–409. Bibcode:2020PalJ...54..401M. doi:10.1134/S0031030120040176. S2CID   221161848. Archived from the original on 2021-01-11. Retrieved 2021-01-09.
2. David W. Dilkes (2020). "Revision of the Early Permian Dissorophid 'Dissorophus' angustus (Temnospondyli: Dissorophoidea)". Journal of Vertebrate Paleontology. 40 (4): e1801704. Bibcode:2020JVPal..40E1704D. doi:10.1080/02724634.2020.1801704. S2CID   225148088.
3. Ralf Werneburg; Stanislav Štamberg; Jean-Sébastien Steyer (2020). "A new stereospondylomorph, Korkonterpeton kalnense gen. et sp. nov., from lower Permian of the Czech Krkonoše Piedmont Basin and a redescription of Intasuchus silvicola from the lower Permian of Russia (Temnospondyli, Amphibia)". Fossil Imprint. 76 (2): 217–242. doi: 10.37520/fi.2020.019 .
4. Rainer R. Schoch; Amy C. Henrici; Robert W. Hook (2020). "A new dissorophoid temnospondyl from the Allegheny Group (late Carboniferous) of Five Points, Mahoning County, Ohio (USA)". Journal of Paleontology. 95 (3): 638–651. doi:10.1017/jpa.2020.101. S2CID   230607719.
5. Eliseu Vieira Dias; Sérgio Dias-da-Silva; Cesar Leandro Schultz (2020). "A new short-snouted rhinesuchid from the Permian of southern Brazil". Revista Brasileira de Paleontologia. 23 (2): 98–122. doi: 10.4072/rbp.2020.2.03 . hdl: 10183/229929 .
6. Juan D. Daza; Edward L. Stanley; Arnau Bolet; Aaron M. Bauer; J. Salvador Arias; Andrej Čerňanský; Joseph J. Bevitt; Philipp Wagner; Susan E. Evans (2020). "Enigmatic amphibians in mid-Cretaceous amber were chameleon-like ballistic feeders". Science. 370 (6517): 687–691. Bibcode:2020Sci...370..687D. doi:10.1126/science.abb6005. PMID   33154135. S2CID   226254862. Archived from the original on 2021-01-11. Retrieved 2021-01-09.
7. Paula Muzzopappa; Agustín G. Martinelli; Juan P. Garderes; Guillermo W. Rougier (2020). "Exceptional avian pellet from the Paleocene of Patagonia and description of its content: a new species of calyptocephalellid (Neobatrachia) anuran". Papers in Palaeontology. 7 (2): 1133–1146. doi: 10.1002/spp2.1333 .
8. Paula Muzzopappa; M. Susana Bargo; Sergio F. Vizcaíno (2025). "Anurans from the Early–Middle Miocene Santa Cruz Formation at Río Chalía (Patagonia, Argentina), and a revision of the fossil Calyptocephalellidae (Anura: Australobatrachia)". Journal of Systematic Palaeontology. 23 (1). 2456622. doi:10.1080/14772019.2025.2456622.
9. Lucas A. Barcelos; Diego Almeida-Silva; Charles M. D. Santos; Vanessa K. Verdade (2020). "Description of a new species of fossil Ceratophrys (Anura: Ceratophryidae) from Versalles Cave, São Paulo, Brazil". Journal of Vertebrate Paleontology. 40 (3): e1811293. Bibcode:2020JVPal..40E1293B. doi:10.1080/02724634.2020.1811293. S2CID   225136948.
10. Federico Agnolin; Ismar Souza Carvalho; Alexis M. Aranciaga Rolando; Fernando E. Novas; José Xavier-Neto; José Artur Ferreira Gomes Andrade; Francisco Idalécio Freitas (2020). "Early Cretaceous neobatrachian frog (Anura) from Brazil sheds light on the origin of modern anurans". Journal of South American Earth Sciences. 101: Article 102633. Bibcode:2020JSAES.10102633A. doi:10.1016/j.jsames.2020.102633. S2CID   218957427.
11. Pavel P. Skutschas; Veniamin V. Kolchanov; Achim H. Schwermann (2020). "First salamander from the Lower Cretaceous of Germany". Cretaceous Research. 116: Article 104606. Bibcode:2020CrRes.11604606S. doi:10.1016/j.cretres.2020.104606. S2CID   225440583.
12. Pavel Skutschas; Veniamin Kolchanov; Sergey Krasnolutskii; Alexander Averianov; Rico Schellhorn; Julia Schultz; Thomas Martin (2020). "A new small-sized stem salamander from the Middle Jurassic of Western Siberia, Russia". PLOS ONE. 15 (2): e0228610. Bibcode:2020PLoSO..1528610S. doi: 10.1371/journal.pone.0228610 . PMC   7029856 . PMID   32074114.
13. Davit Vasilyan; Vadym Yanenko (2020). "The last Palaeoproteus (Urodela: Batrachosauroididae) of Europe". Scientific Reports. 10 (1): Article number 2733. Bibcode:2020NatSR..10.2733V. doi:10.1038/s41598-020-59255-1. PMC   7026125 . PMID   32066790.
14. Per E. Ahlberg; Jennifer A. Clack (2020). "The smallest known Devonian tetrapod shows unexpectedly derived features". Royal Society Open Science. 7 (4): Article ID 192117. Bibcode:2020RSOS....792117A. doi:10.1098/rsos.192117. PMC   7211834 . PMID   32431888.
15. V. V. Bulanov (2020). "A new Leptoropha (Kotlassiidae, Seymouriamorpha) species from the Upper Urzhumian of European Russia". Paleontological Journal. 54 (3): 290–296. Bibcode:2020PalJ...54..290B. doi:10.1134/S0031030120030053. S2CID   219959325. Archived from the original on 2021-01-12. Retrieved 2021-01-09.
16. V. V. Bulanov (2020). "A new kotlassiid (Amphibia, Seymouriamorpha) from the terminal Permian of Eastern Europe". Paleontological Journal. 54 (7): 757–768. Bibcode:2020PalJ...54..757B. doi:10.1134/S0031030120070035. S2CID   229292215.
17. Jianye Chen; Jun Liu (2020). "The youngest occurrence of embolomeres (Tetrapoda: Anthracosauria) from the Sunjiagou Formation (Lopingian, Permian) of North China". Fossil Record. 23 (2): 205–213. Bibcode:2020FossR..23..205C. doi: 10.5194/fr-23-205-2020 .
18. Arjan Mann; Bryan M. Gee; Jason D. Pardo; David Marjanović; Gabrielle R. Adams; Ami S. Calthorpe; Hillary C. Maddin; Jason S. Anderson (2020). "Reassessment of historic 'microsaurs' from Joggins, Nova Scotia, reveals hidden diversity in the earliest amniote ecosystem". Papers in Palaeontology. 6 (4): 605–625. Bibcode:2020PPal....6..605M. doi:10.1002/spp2.1316. S2CID   218925814.
19. Blake V. Dickson; Jennifer A. Clack; Timothy R. Smithson; Stephanie E. Pierce (2020). "Functional adaptive landscapes predict terrestrial capacity at the origin of limbs". Nature. 589 (7841): 242–245. doi:10.1038/s41586-020-2974-5. PMID   33239789. S2CID   227176144.
20. H. M. Byrne; J. A. M. Green; S. A. Balbus; P. E. Ahlberg (2020). "Tides: A key environmental driver of osteichthyan evolution and the fish-tetrapod transition?". Proceedings of the Royal Society A: Mathematical, Physical and Engineering Sciences. 476 (2242): Article ID 20200355. Bibcode:2020RSPSA.47600355B. doi:10.1098/rspa.2020.0355. PMC   7655770 . PMID   33223936.
21. Kendra I. Lennie; Chris F. Mansky; Jason S. Anderson (2020). "New Crassigyrinus-like fibula from the Tournaisian (earliest Carboniferous) of Nova Scotia". Canadian Journal of Earth Sciences. 57 (11): 1365–1369. Bibcode:2020CaJES..57.1365L. doi:10.1139/cjes-2019-0128. S2CID   225160164.
22. A. V. Uliakhin; P. P. Skutschas; P. G. Saburov (2020). "Histology of Dvinosaurus campbelli (Temnospondyli, Dvinosauria) from the Late Permian locality Gorokhovets, Vladimir Region". Paleontological Journal. 54 (6): 632–639. Bibcode:2020PalJ...54..632U. doi:10.1134/S0031030120060106. S2CID   227131135. Archived from the original on 2021-10-27. Retrieved 2021-01-09.
23. Bryan M. Gee (2020). "Size matters: the effects of ontogenetic disparity on the phylogeny of Trematopidae (Amphibia: Temnospondyli)". Zoological Journal of the Linnean Society. 190 (1): 79–113. doi:10.1093/zoolinnean/zlz170.
24. Bryan M. Gee; Robert R. Reisz (2020). "A redescription of the late Carboniferous trematopid Actiobates peabodyi from Garnett, Kansas". The Anatomical Record. 303 (11): 2821–2838. doi: 10.1002/ar.24381 . PMID   32003550. S2CID   210982434.
25. Jason S. Anderson; Diane Scott; Robert R. Reisz (2020). "The anatomy of the dermatocranium and mandible of Cacops aspidephorus Williston, 1910 (Temnospondyli: Dissorophidae), from the Lower Permian of Texas". Journal of Vertebrate Paleontology. 40 (2): e1776720. Bibcode:2020JVPal..40E6720A. doi:10.1080/02724634.2020.1776720. hdl: 1880/113830 . S2CID   222213227.
26. Arjan Mann; Bryan M. Gee (2020). "Lissamphibian-like toepads in an exceptionally preserved amphibamiform from Mazon Creek". Journal of Vertebrate Paleontology. 39 (6): e1727490. doi:10.1080/02724634.2019.1727490. S2CID   216173387.
27. Jade B. Atkins; Peter Sourges; Nadia B. Fröbisch; Robert R. Reisz; Hillary C. Maddin (2020). "Late ontogeny in the small Early Permian amphibamiform dissorophoid Pasawioops mayi". Journal of Vertebrate Paleontology. 40 (2): e1772800. Bibcode:2020JVPal..40E2800A. doi:10.1080/02724634.2020.1772800. S2CID   222210249.
28. Bryan M. Gee; Yara Haridy; Robert R. Reisz (2020). "Histological skeletochronology indicates developmental plasticity in the early Permian stem lissamphibian Doleserpeton annectens". Ecology and Evolution. 10 (4): 2153–2169. Bibcode:2020EcoEv..10.2153G. doi:10.1002/ece3.6054. PMC   7042763 . PMID   32128146.
29. I. V. Novikov; P. A. Haiduk; A. V. Gribanov; А. N. Ivanov; А. V. Novikov; I. A. Starodubtseva (2020). "The earliest case of neoplastic bone lesion in tetrapods". Paleontological Journal. 54 (1): 68–72. Bibcode:2020PalJ...54...68N. doi:10.1134/S0031030120010074. S2CID   212938753. Archived from the original on 2021-01-12. Retrieved 2021-01-09.
30. Michael W. Maisch (2020). "Aphaneramma kokeni (von Huene, 1920), a lonchorhynchine trematosaurid (Amphibia: Temnospondyli) from the Lower Triassic of Pakistan". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 295 (3): 211–241. doi:10.1127/njgpa/2020/0879. S2CID   216325073.
31. Elżbieta M. Teschner; Sanjukta Chakravorti; Dhurjati P. Sengupta; Dorota Konietzko-Meier (2020). "Climatic influence on the growth pattern of Panthasaurus maleriensis from the Late Triassic of India deduced from paleohistology". PeerJ. 8: e9868. doi: 10.7717/peerj.9868 . PMC   7485487 . PMID   33194360.
32. Dorota Konietzko-Meier; Elżbieta M. Teschner; Adam Bodzioch; P. Martin Sander (2020). "Pentadactyl manus of the Metoposaurus krasiejowensis from the Late Triassic of Poland, the first record of pentadactyly among Temnospondyli". Journal of Anatomy. 237 (6): 1151–1161. doi:10.1111/joa.13276. PMC   7704227 . PMID   32707603. S2CID   225501640.
33. Steven L. Wick (2020). "Albanerpetontids (Lissamphibia, Albanerpetontidae) from the Aguja Formation (lower Campanian) of West Texas, USA". Canadian Journal of Earth Sciences. 58 (2): 141–149. doi:10.1139/cjes-2020-0071. S2CID   225381667.
34. Rodolfo Otávio Santos; Michel Laurin; Hussam Zaher (2020). "A review of the fossil record of caecilians (Lissamphibia: Gymnophionomorpha) with comments on its use to calibrate molecular timetrees". Biological Journal of the Linnean Society. 131 (4): 737–755. doi: 10.1093/biolinnean/blaa148 .
35. Rainer R. Schoch; Ralf Werneburg; Sebastian Voigt (2020). "A Triassic stem-salamander from Kyrgyzstan and the origin of salamanders". Proceedings of the National Academy of Sciences of the United States of America. 117 (21): 11584–11588. Bibcode:2020PNAS..11711584S. doi: 10.1073/pnas.2001424117 . PMC   7261083 . PMID   32393623.
36. Yu-Fen Rong; Davit Vasilyan; Li-Ping Dong; Yuan Wang (2020). "Revision of Chunerpeton tianyiense (Lissamphibia, Caudata): is it a cryptobranchid salamander?". Palaeoworld. 30 (4): 708–723. doi: 10.1016/j.palwor.2020.12.001 .
37. Peter Pogoda; Marcus Zuber; Tilo Baumbach; Rainer R. Schoch; Alexander Kupfer (2020). "Cranial shape evolution of extant and fossil crocodile newts and its relation to reproduction and ecology". Journal of Anatomy. 237 (2): 285–300. doi:10.1111/joa.13201. PMC   7369190 . PMID   32297321.
38. Ana M. Báez; Paula Muzzopappa; Geraldo J. Barbosa de Moura (2020). "The earliest records of pipimorph frogs from South America (Aptian, Crato Formation, Brazil): A critical evaluation". Cretaceous Research. 121: Article 104728. doi:10.1016/j.cretres.2020.104728. S2CID   230581615.
39. Thomas Mörs; Marcelo Reguero; Davit Vasilyan (2020). "First fossil frog from Antarctica: implications for Eocene high latitude climate conditions and Gondwanan cosmopolitanism of Australobatrachia". Scientific Reports. 10 (1): Article number 5051. Bibcode:2020NatSR..10.5051M. doi:10.1038/s41598-020-61973-5. PMC   7181706 . PMID   32327670.
40. David C. Blackburn; Rachel M. Keeffe; María C. Vallejo-Pareja; Jorge Vélez-Juarbe (2020). "The earliest record of Caribbean frogs: a fossil coquí from Puerto Rico". Biology Letters. 16 (4): Article ID 20190947. doi:10.1098/rsbl.2019.0947. PMC   7211465 . PMID   32264782.
41. Marcello Ruta; Jennifer A. Clack; Timothy R. Smithson (2020). "A review of the stem amniote Eldeceeon rolfei from the Viséan of East Kirkton, Scotland". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 111 (3): 173–192. Bibcode:2020EESTR.111..173R. doi: 10.1017/S1755691020000079 .
42. Jordi Estefa; Jozef Klembara; Paul Tafforeau; Sophie Sanchez (2020). "Limb-bone development of seymouriamorphs: implications for the evolution of growth strategy in stem amniotes". Frontiers in Earth Science. 8: Article 97. Bibcode:2020FrEaS...8...97E. doi: 10.3389/feart.2020.00097 . S2CID   215756014.
43. Kayla D. Bazzana; Bryan M. Gee; Joseph J. Bevitt; Robert R. Reisz (2020). "Neurocranial anatomy of Seymouria from Richards Spur, Oklahoma". Journal of Vertebrate Paleontology. 39 (5): e1694535. doi:10.1080/02724634.2019.1694535. S2CID   213842156.
44. Kayla D. Bazzana; Bryan M. Gee; Joseph J. Bevitt; Robert R. Reisz (2020). "Postcranial anatomy and histology of Seymouria, and the terrestriality of seymouriamorphs". PeerJ. 8: e8698. doi: 10.7717/peerj.8698 . PMC   7069408 . PMID   32195050.
45. Bryan M. Gee; Joseph J. Bevitt; Robert R. Reisz (2020). "Computed tomographic analysis of the cranium of the early Permian recumbirostran 'microsaur' Euryodus dalyae reveals new details of the braincase and mandible". Papers in Palaeontology. 7 (2): 721–749. doi:10.1002/spp2.1304. S2CID   216359488.
46. Jozef Klembara; Miroslav Hain; Andrej Čerňanský; David S. Berman; Amy C. Henrici (2020). "Anatomy of the neural endocranium and stapes of Diadectes absitus (Diadectomorpha) from the early Permian of Germany based on the high-resolution X-ray microcomputed tomography". The Anatomical Record. 303 (12): 2977–2999. doi: 10.1002/ar.24376 . PMID   31967384. S2CID   210870508.