Americhelydia

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Americhelydia
Temporal range:
Late Jurassic or Early Cretaceous to Holocene 149.50 Ma or 1200 Ma [1] [2] [3]
Sternotherus odoratus.jpg
Common musk turtle ( Sternotherus odoratus ), a species of the family Kinosternidae
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Testudines
Suborder: Cryptodira
Clade: Americhelydia
Crawford et al., 2014
Subclades

Americhelydia is a clade of turtles that consists of sea turtles, snapping turtles, the Central American river turtle and mud turtles, supported by several lines of molecular work. [4] [5] [6] Prior to these studies some morphological and developmental work have considered sea turtles to be basal members of Cryptodira and kinosternids related to the trionychians in the clade Trionychoidea. [7] [8] Americhelydia and Testudinoidea, both clades within Durocryptodira (hardshell turtles), split a part during the early Cretaceous. [9] [10]

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Afrotheria is a superorder of placental mammals, the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles, elephant shrews, otter shrews, tenrecs, aardvarks, hyraxes, elephants, sea cows, and several extinct clades. Most groups of afrotheres share little or no superficial resemblance, and their similarities have only become known in recent times because of genetics and molecular studies. Many afrothere groups are found mostly or exclusively in Africa, reflecting the fact that Africa was an island continent from the Cretaceous until the early Miocene around 20 million years ago, when Afro-Arabia collided with Eurasia.

<span class="mw-page-title-main">Diapsid</span> Clade of reptiles with two holes in each side of their skulls

Diapsids are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The earliest traditionally identified diapsids, the araeoscelidians, appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are often placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. All modern reptiles and birds are placed within the neodiapsid subclade Sauria. Although some diapsids have lost either one hole (lizards), or both holes, or have a heavily restructured skull, they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 snakes, lizards, tuatara, turtles, and crocodiles.

<span class="mw-page-title-main">Muroidea</span> Superfamily of rodents

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<span class="mw-page-title-main">Euarchontoglires</span> Superorder of mammals

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<span class="mw-page-title-main">Cirrina</span> Suborder of octopuses

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<span class="mw-page-title-main">Laurasiatheria</span> Clade of mammals

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<span class="mw-page-title-main">Cryptodira</span> Suborder of reptiles

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<span class="mw-page-title-main">Toxicofera</span> Proposed clade of scaled reptiles

Toxicofera is a proposed clade of scaled reptiles (squamates) that includes the Serpentes (snakes), Anguimorpha and Iguania. Toxicofera contains about 4,600 species of extant Squamata. It encompasses all venomous reptile species, as well as numerous related non-venomous species. There is little morphological evidence to support this grouping; however, it has been recovered by all molecular analyses as of 2012.

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Kinosternoidea is a superfamily of aquatic turtles, which includes two families: Dermatemydidae, and Kinosternidae.

In phylogenetics, basal is the direction of the base of a rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa.

<span class="mw-page-title-main">Alligatoroidea</span> Superfamily of reptiles

Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.

Turtles have been classified in different ways by different authors. While they were previously considered anapsids, they are now considered more derived. Recent analyses of molecular evidence have strongly suggested that they belong in the clade Archosauromorpha. Below are many of the possible classifications of Testudines and Testudinata:

<span class="mw-page-title-main">Archelosauria</span> Clade comprising turtles, birds and crocodilians

Archelosauria is a clade grouping turtles and archosaurs and their fossil relatives, to the exclusion of lepidosaurs. The majority of phylogenetic analyses based on molecular data have supported a sister-group relationship between turtles and archosaurs. On the other hand, Archelosauria had not been historically supported by most morphological analyses, which have instead found turtles to either be descendants of parareptiles, early-diverging diapsids outside of Sauria, or close relatives of lepidosaurs within the clade Ankylopoda. Some recent morphological analyses have also found support for Archelosauria.

<span class="mw-page-title-main">Pantestudines</span> Clade of reptiles

Pantestudines or Pan-Testudines is the proposed group of all reptiles more closely related to turtles than to any other living animal. It includes both modern turtles and all of their extinct relatives. Pantestudines with a complete shell are placed in the clade Testudinata.

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<span class="mw-page-title-main">Neoanguimorpha</span> Clade of lizards

Neoanguimorpha is a clade of anguimorphs comprising Monstersauria and Diploglossa. Morphological studies in the past had classified helodermatids with the varanoids in the clade Platynota, while the Chinese crocodile lizard was classified as a xenosaurid. However molecular work found no support in these groupings and instead has found the helodermatids more related to Diploglossa, while the Chinese crocodile lizard and varanoids to form the clade Paleoanguimorpha.

<span class="mw-page-title-main">Diploglossa</span> Clade of lizards

Diploglossa is a clade of neoanguimorphs represented by the families Xenosauridae, Diploglossidae, Anniellidae and Anguidae, the latter three placed in the superfamily Anguioidea. In the past the Chinese crocodile lizard was classified as a xenosaurid; current molecular work has shown evidence the species related to varanoids in the clade Paleoanguimorpha.

<span class="mw-page-title-main">Colubroides</span> Clade of snakes

The Colubroides are a clade in the suborder Serpentes (snakes). It contains over 85% of all the extant species of snakes. The largest family is Colubridae, but it also includes at least six other families, at least four of which were once classified as "Colubridae" before molecular phylogenetics helped in understanding their relationships. It has been found to be monophyletic.

<span class="mw-page-title-main">Thalassochelydia</span> Extinct clade of turtles

Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including Eurysternum, Plesiochelys and Thalassemys to the exclusion of Pelomedusa, Testudo and Protostega. While a clade uniting the families Eurysternidae, Plesiochelyidae and Thalassemydidae had been supported by phylogenetic evidence, a name was not given for the clade until 2017, when Jérémy Anquetin and colleagues coined Thalassochelydia.

References

  1. Joyce, W. G., Parham, J. F., Lyson, T. R., Warnock, R. C., & Donoghue, P. C. (2013). A divergence dating analysis of turtles using fossil calibrations: an example of best practices. Journal of Paleontology, 87(04), 612-634.
  2. "Protostegidae". The Paleobiology Database. Retrieved 27 July 2013.
  3. Edwin A. Cadena and James F. Parham (2015). "Oldest known marine turtle? A new protostegid from the Lower Cretaceous of Colombia". PaleoBios. 32 (1): 1–42.
  4. Chandler, C. H., & Janzen, F. J. (2009). The phylogenetic position of the snapping turtles (Chelydridae) based on nucleotide sequence data. Copeia, 2009(2), 209-213.
  5. Barley, A. J., Spinks, P. Q., Thomson, R. C., & Shaffer, H. B. (2010). Fourteen nuclear genes provide phylogenetic resolution for difficult nodes in the turtle tree of life. Molecular Phylogenetics and Evolution, 55(3), 1189-1194.
  6. Crawford, N. G., Parham, J. F., Sellas, A. B., Faircloth, B. C., Glenn, T. C., Papenfuss, T. J., ... & Simison, W. B. (2015). A phylogenomic analysis of turtles. Molecular Phylogenetics and Evolution, 83, 250-257.
  7. Joyce, W. G. (2007). Phylogenetic relationships of Mesozoic turtles. Bulletin of the Peabody Museum of Natural History, 48(1), 3-102.
  8. Werneburg, I., & Sánchez-Villagra, M. R. (2009). Timing of organogenesis support basal position of turtles in the amniote tree of life. BMC Evolutionary Biology, 9(1), 82.
  9. Pereira, Anieli G.; Sterli, Juliana; Moreira, Filipe R.R.; Schrago, Carlos G. (August 2017). "Multilocus phylogeny and statistical biogeography clarify the evolutionary history of major lineages of turtles". Molecular Phylogenetics and Evolution. 113: 59–66. doi:10.1016/j.ympev.2017.05.008. hdl: 11336/41137 . ISSN   1055-7903. PMID   28501611.
  10. Joyce, Walter G.; Rabi, Márton; Clark, James M.; Xu, Xing (2016-10-28). "A toothed turtle from the Late Jurassic of China and the global biogeographic history of turtles". BMC Evolutionary Biology. 16 (1): 236. doi: 10.1186/s12862-016-0762-5 . ISSN   1471-2148. PMC   5084352 . PMID   27793089.