Butterflyfishes Temporal range: | |
---|---|
Various examples of butterflyfish | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Acanthuriformes |
Family: | Chaetodontidae Rafinesque, 1810 |
Genera | |
About 10, see text | |
Synonyms | |
Chaetodontinae (but see text) |
The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group. The approximately 129 species in 12 genera [2] are found mostly on the reefs of the Atlantic, Indian, and Pacific Oceans. A number of species pairs occur in the Indian and Pacific Oceans, members of the huge genus Chaetodon .
Butterflyfish look like smaller versions of angelfish (Pomacanthidae), but unlike these, lack preopercle spines at the gill covers. Some members of the genus Heniochus resemble the Moorish idol (Zanclus cornutus) of the monotypic Zanclidae. Among the paraphyletic Perciformes, the former are probably not too distantly related to butterflyfish, whereas the Zanclidae seem far less close.
Butterflyfish mostly range from 12 to 22 cm (4.7 to 8.7 in) in length. The largest species, the lined butterflyfish and the saddle butterflyfish, C. ephippium, grow to 30 cm (12 in). The common name references the brightly coloured and strikingly patterned bodies of many species, bearing shades of black, white, blue, red, orange, and yellow. Other species are dull in colour. Butterflyfish are a boundless, different group of marine percoids with delegates on practically all coral reef frameworks and in every single tropical ocean. Their bright and color patterns have drawn in much consideration, creating an abundance of data about their conduct and environment. [3] Many have eyespots on their flanks and dark bands across their eyes, not unlike the patterns seen on butterfly wings. [4] Their deep, laterally narrow bodies are easily noticed through the profusion of reef life. The conspicuous coloration of butterflyfish may be intended for interspecies communication. Butterflyfish have uninterrupted dorsal fins with tail fins that may be rounded or truncated, but are never forked.
Generally diurnal and frequenting waters less than 18 m (59 ft) deep (though some species descend to 180 m (590 ft), butterflyfish stick to particular home ranges. These corallivores are especially territorial, forming pairs and staking claim to a specific coral head. Contrastingly, the zooplankton feeders form large conspecific groups. By night, butterflyfish hide in reef crevices and exhibit markedly different coloration.
Their coloration also makes them popular aquarium fish. However, most species feed on coral polyps and sea anemones. Balancing the relative populations of prey and predator is complex, leading hobby aquarists to focus on the few generalists and specialist zooplankton feeders.
Butterflyfish are pelagic spawners; that is, they release many buoyant eggs into the water, which become part of the plankton, floating with the currents until hatching. The fry go through a tholichthys stage, wherein the body of the postlarval fish is covered in large, bony plates extending from the head. They lose their bony plates as they mature. [4] Only one other family of fish, the scats (Scatophagidae) express such an armored stage.
The Chaetodontidae can be, but are not usually, divided into two lineages that arguably are subfamilies. The subfamily name Chaetodontinae is a little-used leftover from the period when the Pomacanthidae and Chaetodontidae were united under the latter name as a single family. Hence, Chaetodontinae is today considered a junior synonym of Chaetodontidae. In any case, one lineage of Chaetodontidae (in the modern sense) contains the "typical" butterflyfish around Chaetodon, while the other unites the bannerfish and coralfish genera. As the Perciformes are highly paraphyletic, the precise relationships of the Chaetodontidae as a whole are badly resolved. [5]
Chaetodontidae is classified within the suborder Percoidei by the 5th edition of Fishes of the World , but they are placed in an unnamed clade which sits outside the superfamily Percoidea. This clade contains 7 families which appear to have some relationship to Acanthuroidei, Monodactylidae, and Priacanthidae. [6] Other authorities have paced the family in the order Chaetodontiformes alongside the family Leiognathidae. [7]
Before DNA sequencing, the taxonomy was confused about whether to treat these as species or subspecies. Also, numerous subgenera have been proposed for splitting out of Chaetodon, and it is becoming clear how to subdivide the genus if that is desired. [8]
The fossil record of this group is marginal. Their restriction to coral reefs means their carcasses are liable to be dispersed by scavengers, overgrown by corals, and any that do fossilize will not long survive erosion. However, Pygaeus , a very basal fossil from the mid- to late Eocene of Europe, dates from around the Bartonian 40–37 million years ago (Mya). Thus, the Chaetodontidae emerged probably in the early to mid-Eocene. A crude molecular clock in combination with the evidence given by Pygaeus allows placement of the initial split between the two main lineages to the middle to late Eocene, and together with the few other fossils, it allows the deduction that most living genera were probably distinct by the end of the Paleogene 23 Mya. [9]
The bannerfish-coralfish lineage can be further divided in two groups; these might be considered tribes, but have not been formally named. Genera are listed in order of the presumed phylogeny, from the most ancient to the youngest: [5] [10]
Bannerfish/coralfish lineage 1:
Bannerfish/coralfish lineage 2:
The "typical" butterflyfishes may eventually come to contain more genera; see Chaetodon :
Chaetodon is a tropical fish genus in the family Chaetodontidae. Like their relatives, they are known as "butterflyfish". This genus is by far the largest among the Chaetodontidae, with about 90 living species included here, though most might warrant recognition as distinct genera.
The blue-cheeked butterflyfish is a marine species of ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the north-western Indian Ocean.
The raccoon butterflyfish, also known as the crescent-masked butterflyfish, lunule butterflyfish, halfmoon butterflyfish, moon butterflyfish, raccoon butterfly, raccoon, raccoon coralfish, and redstriped butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indian and Pacific Oceans.
The ornate butterflyfish, or clown butterflyfish is a species of marine ray-finned fish, a butterflyfish in the family Chaetodontidae.
Chaetodon ulietensis, the Pacific double-saddle butterflyfish or false falcula butterflyfish, is a species of butterflyfish. It flourishes in coral-rich environments in the central Indo-Pacific region. Their range extends from the Cocos-Keeling Islands to the Tuamotu Islands, and north to Japan. They are usually found from the surface to 20 m depths, and like shallow channels with high current.
The blacktail butterflyfish, also known as black-tailed butterflyfish or exquisite butterflyfish, is a species marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is native to the western Indian Ocean but has reached the Mediterranean Sea as a Lessepsian migrant through the Suez Canal.
Chaetodon citrinellus is a species of butterflyfish. It is commonly known as the speckled butterflyfish or citron butterflyfish. It is found in the Indo-Pacific: the Red Sea, East Africa to the Hawaiian, Marquesan and Tuamotu islands, north to southern Japan and south to Australia.
The melon butterflyfish or the Indian redfin butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indian Ocean from East Africa to Western Java. This is one species of a closely related group which includes the blacktail butterflyfish of the Red Sea and Gulf of Aden and the oval butterflyfish which is found in the western Pacific, from eastern coasts of the Indonesian islands to Australia.
The Indian vagabond butterflyfish, also known as the blackened butterflyfish or black-finned vagabond, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indo-West Pacific, from the Maldives via India, Sri Lanka and the Andaman Sea to the westernmost portion of the Indonesian archipelago.
The mirror butterflyfish or oval-spot butterflyfish is a species of butterflyfish. It is found in the Indo-Pacific region from Indonesia to Japan and south to the Great Barrier Reef and Papua New Guinea. The species has also been reported from Madagascar, Mauritius and Réunion.
The scrawled butterflyfish, also known as Meyer's butterflyfish or the maypole butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indian and Pacific Oceans.
The bluelashed butterflyfish, also known as the eclipse butterflyfish, archer butterflyfish or Bennett's butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae, It is found in the Indo-Pacific.
The Spot-naped Butterflyfish, also known as the pig-face butterflyfish, is a species of marine ray-finned fish. a butterflyfish belonging to the family Chaetodontidae. It is found in the Indo- West Pacific region from Sri Lanka to Queensland, north to Indonesia and the Philippines.
The latticed butterflyfish, also known as Raffles’ coralfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indo-Pacific region.
The Yellow-dotted Butterflyfish is a poorly known marine ray-finned fish species, a butterflyfish belonging to the family Chaetodontidae. It is found in the Indi-Western Pacific Region.
Chaetodon mertensii, the atoll butterflyfish, yellowback butterflyfish or Merten's butterflyfish, is a species of marine ray finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the western Pacific Ocean.
Chaetodon pelewensis, the dot dash butterflyfish, spotbanded butterflyfish or punctato butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is found in the western Pacific Ocean.
Chaetodon plebeius, the blueblotch butterflyfish, bluespot butterflyfish, bluedash butterflyfish or grey-blotched butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is native to the Indian and Pacific Oceans.
Chaetodon gardineri, Gardiner's butterflyfish, is a species of marine ray-finned fish, a butterflyfish belonging to the family Chaetodontidae. It is native to the Indian Ocean.
Chaetodon rainfordi, Rainfords's butterflyfish, also known as the gold-barred butterflyfish or the Northern butterflyfish, is a species of marine ray-finned fish, a butterflyfish, belonging to the family Chaetodontidae. It is found in the southwestern Pacific Ocean where it is associated with coral reefs.