Dysderoidea Temporal range: | |
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Segestria florentina (Segestriidae) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Araneae |
Infraorder: | Araneomorphae |
Superfamily: | Dysderoidea |
Families [1] | |
Diversity | |
4 or 5 families |
The Dysderoidea are a clade or superfamily of araneomorph spiders. The monophyly of the group, initially consisting of the four families Dysderidae, Oonopidae, Orsolobidae and Segestriidae, [2] has consistently been recovered in phylogenetic studies. [1] [3] In 2014, a new family, Trogloraptoridae, was created for a recently discovered species Trogloraptor marchingtoni. It was suggested that Trogloraptoridae may be the most basal member of the Dysderoidea clade. [1] [4] However, a later study found that Trogloraptoridae was placed outside the Dysderoidea and concluded that it was not part of this clade. [5]
Dysderoidea are members of the Haplogynae clade: spiders with simpler copulatory organs (palpal bulbs and epigynes) than other araneomorphs. One hypothesis for relationships within the Haplogynae is shown below. [6] The status of the Trogloraptoridae is unclear. The family was not included in one study which otherwise found the same topography, [7] but it was placed outside even the Filistatidae in a 2014 study based on ribosomal DNA. [5]
Haplogynae |
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Other studies have suggested that Caponiidae rather than Tetrablemmidae are the sister of Dysderoidea. [8]
Crevice weaver spiders (Filistatidae) comprise cribellate spiders with features that have been regarded as "primitive" for araneomorph spiders. They are weavers of funnel or tube webs. The family contains 18 genera and more than 120 described species worldwide.
Spider taxonomy is that part of taxonomy that is concerned with the science of naming, defining and classifying all spiders, members of the Araneae order of the arthropod class Arachnida with more than 48,500 described species. However, there are likely many species that have escaped the human eye to this day, and many specimens stored in collections waiting to be described and classified. It is estimated that only one third to one half of the total number of existing species have been described.
Norman Ira Platnick was an American biological systematist and arachnologist. At the time of his death, he was a professor emeritus of the Richard Gilder Graduate School and Peter J. Solomon Family Curator Emeritus of the invertebrate zoology department of the American Museum of Natural History. A 1973 Ph.D. recipient at Harvard University, Platnick described over 1,800 species of spiders from around the world, making him the second most prolific spider taxonomist in history, behind only Eugène Simon. Until 2014 he was also the maintainer of the World Spider Catalog, a website formerly hosted by the AMNH which tracks the arachnology literature, and attempts to maintain a comprehensive list, sorted taxonomically, of every species of spider which has been formally described. In 2007 he received the International Society of Arachnology's Bonnet award, named for Pierre Bonnet, in recognition of his work on the catalog.
The Micropholcommatinae are a subfamily of araneomorph spiders in the family Anapidae. They were previously treated as the family Micropholcommatidae. Micropholcommatins are extremely small, with body lengths typically between 0.5 and 2 mm. They are usually found among leaf litter or moss.
The Dionycha are a clade of spiders (Araneomorphae:Entelegynae), characterized by the possession of two tarsal claws with tufts of hairs (setae) beside them, which produce strong adhesion, enabling some species to climb glass. The circumscription of the group has varied widely; a 2021 analysis resulted in about 20 families, including Salticidae, Gnaphosidae, and Clubionidae.
Lycosoidea is a clade or superfamily of araneomorph spiders. The traditional circumscription was based on a feature of the eyes. The tapetum is a reflective layer at the back of the eye, thought to increase sensitivity in low light levels. Lycosoids were then defined by having a "grate-shaped" tapetum. Research from the late 1990s onwards suggests that this feature has evolved more than once, possibly as many as five times, so that the original Lycosoidea is paraphyletic. Studies published in 2014 and 2015 suggest that a smaller group of families does form a clade.
Scytodoidea is a taxon of araneomorph spiders, at the rank of superfamily. It contains four families:
The Leptonetoidea are a superfamily of haplogyne araneomorph spiders with three families. Phylogenetic studies have provided weak support for the relationship among the families. The placement of one of the families within the Haplogynae has been questioned.
The Pholcoidea or pholcoids are a superfamily of araneomorph spiders. The group has been circumscribed to contain the following three families:
The Eresoidea or eresoids are a group of araneomorph spiders that have been treated as a superfamily. As usually circumscribed, the group contains three families: Eresidae, Hersiliidae and Oecobiidae. Studies and reviews based on morphology suggested the monophyly of the group; more recent gene-based studies have found the Eresidae and Oecobiidae to fall into different clades, placing doubt on the acceptability of the taxon. Some researchers have grouped Hersiliidae and Oecobiidae into the separate superfamily Oecobioidea, a conclusion supported in a 2017 study, which does not support Eresoidea.
The Agelenoidea or agelenoids are a superfamily or informal group of entelegyne araneomorph spiders. Phylogenetic studies since 2000 have not consistently recovered such a group, with more recent studies rejecting it.
The Dictynoidea or dictynoids are a group of araneomorph spiders that have been treated as a superfamily. The composition of the group has varied. Phylogenetic studies in the 21st century have failed to confirm the monophyly of the dictynoids as originally defined.
The Austrochiloidea or austrochiloids are a group of araneomorph spiders, treated as a superfamily. The taxon contains two families of eight-eyed spiders:
The Haplogynae or haplogynes are one of the two main groups into which araneomorph spiders have traditionally been divided, the other being the Entelegynae. Morphological phylogenetic studies suggested that the Haplogynae formed a clade; more recent molecular phylogenetic studies refute this, although many of the ecribellate haplogynes do appear to form a clade, Synspermiata.
The Entelegynae or entelegynes are a subgroup of araneomorph spiders, the largest of the two main groups into which the araneomorphs were traditionally divided. Females have a genital plate (epigynum) and a "flow through" fertilization system; males have complex palpal bulbs. Molecular phylogenetic studies have supported the monophyly of Entelegynae.
Chileolobus is a monotypic genus of Chilean araneomorph spiders in the family Orsolobidae containing the single species, Chileolobus eden. It was first described by Raymond Robert Forster & Norman I. Platnick in 1985, and is only found in Chile.
Dugdalea is a monotypic genus of Polynesian araneomorph spiders in the family Orsolobidae containing the single species, Dugdalea oculata. It was first described by Raymond Robert Forster & Norman I. Platnick in 1985, and is only found in New Zealand.
Tautukua is a monotypic genus of Polynesian araneomorph spiders in the family Orsolobidae containing the single species, Tautukua isolata. It was first described by Raymond Robert Forster & Norman I. Platnick in 1985, and is only in New Zealand.
Synspermiata is a clade of araneomorph spiders, comprising most of the former "haplogynes". They are united by having simpler genitalia than other araneomorph spiders, lacking a cribellum, and sharing an evolutionary history of synspermia – a particular way in which spermatozoa are grouped together when transferred to the female.
Avicularioidea is a clade of mygalomorph spiders, one of the two main clades into which mygalomorphs are divided. It has been treated at the rank of superfamily.