Elphidae

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Elphidae
Temporal range: Late Permian
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Suborder: Anomodontia
Clade: Dicynodontia
Clade: Bidentalia
Family: Elphidae
Kurkin, 2010
Genera

Elphidae is a clade of bidentalian dicynodonts containing Elph , Katumbia , and Interpresosaurus . [1] [2] It is exclusively known from the Late Permian of Russia and Tanzania. Elphidae is variously recovered as either at the base of a paraphyletic Cryptodontia, [2] or as basal dicynodontoids. [3]

The clade Elphinae was erected by Kurkin (2010) to pertain to a clade containing Elph and Interpresosaurus, two dicynodonts from Late Permian Russia. This clade was diagnosed by a sharply shortened preorbital skull, relatively small tusks located below the anterior orbit, and maxillae with depressions located below the naris. [4] Elphinae is often recovered forming a clade with the Tanzanian Katumbia , [1] to which the name Elphidae can be applied under the rules of the ICZN. [2]

Related Research Articles

<span class="mw-page-title-main">Dicynodontia</span> Extinct clade of therapsids

Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers.

<span class="mw-page-title-main">Anomodontia</span> Suborder of stem-mammals

Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores. Anomodonts were very diverse during the Middle Permian, including primitive forms like Anomocephalus and Patranomodon and groups like Venyukovioidea and Dromasauria. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage (Kannemeyeriiformes) evolved into large, stocky forms that became dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline, finally going extinct during the Triassic–Jurassic extinction event.

<i>Kannemeyeria</i> Extinct genus of dicynodonts

Kannemeyeria is a genus of dicynodont that lived during the Anisian age of Middle Triassic period in what is now Africa and South America. The generic name is given in honor of Daniel Rossouw Kannemeyer, the South African fossil collector who discovered the original specimen. It is one of the first representatives of the family, and hence one of the first large herbivores of the Triassic.

<span class="mw-page-title-main">Dromasauria</span>

"Dromasaurs" are an artificial grouping of small anomodont therapsids from the Middle and Late Permian of South Africa. They represent either a paraphyletic grade or a polyphyletic grouping of small non-dicynodont basal anomodonts rather than a clade, and as such are considered an invalid group today. "Dromasaurs" were historically united by their superficially similar appearances that were unlike other known anomodonts. They are all small in size with slender limbs and long tails, and have short skulls with very large eye sockets. "Dromasauria" traditionally includes three genera, all from the Karoo Supergroup of South Africa: Galepus, Galechirus, and Galeops. These genera have sometimes been divided into two subgroups, the monotypic family Galeopidae and the Galechiridae for Galechiris and Galepus.

<i>Dicynodon</i> Extinct genus of dicynodonts

Dicynodon is a genus of dicynodont therapsid that flourished during the Upper Permian period. Like all dicynodonts, it was an herbivorous animal. This synapsid was toothless, except for prominent tusks, hence the name. It probably cropped vegetation with a horny beak, much like a tortoise, while the tusks may have been used for digging up roots and tubers.

<span class="mw-page-title-main">Stahleckeriidae</span> Extinct family of dicynodonts

Stahleckeriidae is a family of dicynodont therapsids whose fossils are known from the Triassic of North America, South America, Asia and Africa.

Uralokannemeyeria is an extinct genus of kannemeyeriiform dicynodont known from the Middle Triassic Donguz Formation of Bashkortostan, Russia.

Interpresosaurus is a genus of dicynodont from the Late Permian (Changhsingian) Vyazniki Assemblage of Russia. The type and only species, Interpresosaurus blomi, is known from a single fragmentary skull, PIN 3584/1.

Katumbia is a genus of dicynodont from Late Permian (Changhsingian) Kawinga Formation of the Ruhuhu Basin, Tanzania. and possibly the Upper Madumabisa Mudstone Formation of the Luangwa Basin, Zambia. The type species, K. parringtoni, was originally referred to the genus Cryptocynodon, which is now recognized as a junior synonym of Endothiodon.

<i>Vivaxosaurus</i> Extinct genus of dicynodonts

Vivaxosaurus is a genus of dicynodont from Late Permian (Changhsingian) of Russia. It has been found at Sokolki on the Northern Dvina River near Kotlas in Arkhangelsk Oblast, Russia. It lived during the latest Permian, and was a contemporary of Inostrancevia, Scutosaurus and Dvinia. Like all members of the genus, this animal was toothless, except for prominent tusks, and probably cropped vegetation with a horny beak, like a tortoise.

<span class="mw-page-title-main">Lystrosauridae</span> Extinct family of dicynodonts

Lystrosauridae is a family of dicynodont therapsids from the Permian and Triassic time periods. It includes two genera, Lystrosaurus and Kwazulusaurus. Kwazulusaurus includes a single species, K. shakai, from the Late Permian of South Africa and Lystrosaurus includes many species from the Late Permian and Early Triassic of South Africa, India, and Antarctica.

<span class="mw-page-title-main">Venyukovioidea</span> Extinct infraorder of therapsids

Venyukovioidea is an infraorder of anomodont therapsids related to dicynodonts from the Permian of Russia. They have also been known as 'Venjukovioidea', as well as by the similar names 'Venyukoviamorpha' or 'Venjukoviamorpha' in literature. This in part owes to a misspelling by Russian palaeontologist Ivan Efremov in 1940 when he mistakenly spelt Venyukovia, the namesake of the group, with a 'j' instead of a 'y', which permeated through subsequent therapsid literature before the mistake was caught and corrected. The order Ulemicia has also been coined for a similar taxonomic concept in Russian scientific literature, which notably excludes Suminia and Parasuminia.

<span class="mw-page-title-main">Chainosauria</span> Extinct clade of therapsids

Chainosauria is a large and speciose clade of anomodont therapsid that includes the highly diverse dicynodonts and a small number of closely related basal genera —although the total composition and taxonomic scope of Chainosauria is in flux. Chainosauria was named in 1923 to group together the dicynodonts and their close relatives, namely three small anomodont genera from South Africa that made up the now defunct group 'Dromasauria'. The name soon fell into disuse, however, as it was functionally replaced by Anomodontia. Chainosauria was later revived cladistically in 2009, preserving the association of dicynodonts and the 'dromasaurs' and has since served in effect as both a cladistic and a biogeographic counterpart to the Laurasian venyukovioids, with early chainosaurs appearing to have been a Gondwanan radiation.

Elph is an extinct genus of dicynodont therapsids from Russia. Four specimens have been found from the Sokolki Assemblage in European Russia, representing a fauna that dates back to the Late Permian. The holotype, PIN 2353/37, was originally considered a juvenile specimen due to its small size, before being recognized as a mature individual.

<span class="mw-page-title-main">Pylaecephalidae</span> Extinct family of dicynodonts

Pylaecephalidae is a family of dicynodont therapsids that includes Diictodon, Robertia, and Prosictodon from the Permian of South Africa. Pylaecephalids were small burrowing dicynodonts with long tusks. The family was first named in 1934 and was redefined in 2009. Diictodontidae and Robertiidae are considered junior synonyms of Pylaecephalidae; although Pylaecephalus itself is considered a junior synonym of Diictodon, the name Pylaecephalidae predates these names and therefore takes priority.

<span class="mw-page-title-main">Kannemeyeriiformes</span> Extinct clade of dicynodonts

Kannemeyeriiformes is a group of large-bodied Triassic dicynodonts. As a clade, Kannemeyeriiformes has been defined to include the species Kannemeyeria simocephalus and all dicynodonts more closely related to it than to the species Lystrosaurus murrayi.

Syops is an extinct genus of dicynodont therapsid. The type species S. vanhoepeni was first named in 1938 as Dicynodon vanhoepeni. Fossils of the genus have been found in the Cistecephalus Assemblage Zone in the Usili Formation of the Ruhuhu Basin, Tanzania and the Upper Madumabisa Mudstone Formation of the Luangwa Basin, Zambia. Its phylogenetic placement is somewhat uncertain, with multiple different studies finding it as either a basal geikiid, rhachiocephalid a dicynodontoid more derived than the most basal genera but less derived than Lystrosauridae, or a lystrosaurid.

<span class="mw-page-title-main">Bidentalia</span> Extinct clade of dicynodonts

Bidentalia is a group of dicynodont therapsids. Bidentalia was one of the first names used to describe dicynodonts; the group was established in 1876, while the name "bidentals" dates back as far as 1845. With the increasing prominence of phylogenetics, the group was redefined as a clade in 2009. Bidentalia is now considered a stem-based taxon that includes all taxa more closely related to Aulacephalodon bainii and Dicynodon lacerticeps than Emydops arctatus.

<span class="mw-page-title-main">Anomocephaloidea</span> Extinct clade of therapsids

Anomocephaloidea is a clade of basal anomodont therapsids related to the dicynodonts known from what is now South Africa and Brazil during the Middle Permian. It includes only two species, Anomocephalus africanus from the Karoo Basin of South Africa and Tiarajudens eccentricus from the Paraná Basin of Brazil. Anomocephaloidea was named in 2011 with the discovery of Tiarajudens, although Anomocephalus itself has been known since 1999.

<i>Rastodon</i> Extinct genus of dicynodonts

Rastodon is an extinct genus of dicynodont. Uniquely among dicynodonts, its tusks curve forward. The type and only species is R. procurvidens.

References

  1. 1 2 Boos, Alessandra D. S.; Kammerer, Christian F.; Schultz, Cesar L.; Soares, Marina B.; Ilha, Ana L. R. (2016). "A new dicynodont (Therapsida: Anomodontia) from the Permian of southern Brazil and its implications for bidentalian origins". PLOS ONE. 11 (5): e0155000. Bibcode:2016PLoSO..1155000B. doi: 10.1371/journal.pone.0155000 . PMC   4880204 . PMID   27224287.
  2. 1 2 3 Angielczyk, Kenneth D.; Kammerer, Christian F. (2017). "The Cranial Morphology, Phylogenetic Position and Biogeography of the Upper Permian Dicynodont Compsodon helmoedi Van Hoepen (Therapsida, Anomodontia)". Papers in Palaeontology. 3 (4): 513–545. doi:10.1002/spp2.1087. S2CID   134092461.
  3. Kammerer, C. F.; Fröbisch, J. R.; Angielczyk, K. D. (2013). Farke, Andrew A (ed.). "On the Validity and Phylogenetic Position of Eubrachiosaurus browni, a Kannemeyeriiform Dicynodont (Anomodontia) from Triassic North America". PLOS ONE. 8 (5): e64203. Bibcode:2013PLoSO...864203K. doi: 10.1371/journal.pone.0064203 . PMC   3669350 . PMID   23741307.
  4. Kurkin, A.A. (2010). "Late Permian Dicynodonts of Eastern Europe". Paleontological Journal. 44 (6): 672–681. doi:10.1134/S0031030110060092. S2CID   131459807.