Dicynodon is a genus of dicynodont therapsid that flourished during the Upper Permian period. Like all dicynodonts, it was an herbivorous animal. This synapsid was toothless, except for prominent tusks, hence the name. It probably cropped vegetation with a horny beak, much like a tortoise, while the tusks may have been used for digging up roots and tubers.
Emydops is an extinct genus of dicynodont therapsids from the Middle Permian to Late Permian of what is now South Africa. The genus is generally small and herbivorous, sharing the dicynodont synapomorphy of bearing two tusks. In the following years, the genus grew to include fourteen species. Many of these species were erected on the basis of differences in the teeth and the positioning of the frontal and parietal bones. A 2008 study narrowed Emydops down to two species, E. arctatus and the newly described E. oweni.
Dicynodontoides is a genus of small to medium-bodied, herbivorous, emydopoid dicynodonts from the Late Permian. The name Dicynodontoides references its “dicynodont-like” appearance due to the caniniform tusks featured by most members of this infraorder. Kingoria, a junior synonym, has been used more widely in the literature than the more obscure Dicynodontoides, which is similar-sounding to another distantly related genus of dicynodont, Dicynodon. Two species are recognized: D. recurvidens from South Africa, and D. nowacki from Tanzania.
Daptocephalus is an extinct genus of dicynodont synapsid, which was found in Late Permian strata, in a biozone known precisely for the presence of fossils of this dicynodont, the Daptocephalus Assemblage Zone, in the Karoo Basin in South Africa. An additional species, D. huenei, is known from the Usili Formation in Tanzania and was formerly assigned to the genus Dicynodon before a study in 2019 recognised that the type specimen belonged to Daptocephalus.
Vivaxosaurus is a genus of dicynodont from Late Permian (Changhsingian) of Russia. It has been found at Sokolki on the Northern Dvina River near Kotlas in Arkhangelsk Oblast, Russia. It lived during the latest Permian, and was a contemporary of Inostrancevia, Scutosaurus and Dvinia. Like all members of the genus, this animal was toothless, except for prominent tusks, and probably cropped vegetation with a horny beak, like a tortoise.
Lystrosauridae is a family of dicynodont therapsids from the Permian and Triassic time periods. It includes two genera, Lystrosaurus and Kwazulusaurus. Kwazulusaurus includes a single species, K. shakai, from the Late Permian of South Africa and Lystrosaurus includes many species from the Late Permian and Early Triassic of South Africa, India, and Antarctica.
Elph is an extinct genus of dicynodont therapsids from Russia. Four specimens have been found from the Sokolki Assemblage in European Russia, representing a fauna that dates back to the Late Permian. The holotype, PIN 2353/37, was originally considered a juvenile specimen due to its small size, before being recognized as a mature individual.
Peramodon is an extinct genus of dicynodont therapsid from the Late Permian Scutosaurus karpinskii Zone of the Salarevo Formation of Russia. The type species, P. amalitzkii, was first named in 1926 as Dicynodon amalitzkii.
Keyseria is an extinct genus of dicynodont therapsid. The type species K. benjamini was first named in 1948 as Dicynodon benjamini.
Syops is an extinct genus of dicynodont therapsid. The type species S. vanhoepeni was first named in 1938 as Dicynodon vanhoepeni. Fossils of the genus have been found in the Cistecephalus Assemblage Zone in the Usili Formation of the Ruhuhu Basin, Tanzania and the Upper Madumabisa Mudstone Formation of the Luangwa Basin, Zambia. Its phylogenetic placement is somewhat uncertain, with multiple different studies finding it as either a basal geikiid, rhachiocephalid a dicynodontoid more derived than the most basal genera but less derived than Lystrosauridae, or a lystrosaurid.
Basilodon is an extinct genus of dicynodont therapsid. The type species, Basilodon woodwardi was originally named in 1921 as Dicynodon woodwardi. Fossils have been found in the Cistecephalus Assemblage Zone and Dicynodon Assemblage Zone of the Balfour Formation of the Beaufort Group in South Africa.
Therochelonia is a group of dicynodont therapsids. The group was named by British paleontologist Harry Seeley in 1894 and fell into disuse in the following century. Therochelonia was redefined as a node-based clade in 2009. It is defined as the last common ancestor of Cistecephalus microrhinus and Dicynodon lacerticeps, and all of its descendants. Below is a simplified cladogram from Kammerer et al. (2011) showing the phylogenetic placement of Therochelonia:
Sintocephalus is an extinct genus of dicynodont therapsid from the Late Permian of South Africa. Fossils are known from the Cistecephalus Assemblage Zone of the Beaufort Group. The type species of Sintocephalus, S. alticeps, was first named in 1913 as a species of Dicynodon. The genus was erected in 1934, but in subsequent years its species were often regarded as members of other dicynodont genera.
Gordonia is an extinct genus of dicynodont therapsid from the Late Permian of Scotland. Fossils have been found from the Elgin sandstone of Cutties Hillock Sandstone in Elgin, Moray. These are among the many amniote fossils referred to as the Elgin Reptiles. Gordonia was named in 1893 with four species: G. traquairi, G. duffiana, G. huxleyana, and G. juddiana. Currently, the only recognized species is the type G. traquairi. All other species are considered synonyms of the type.
Daqingshanodon is an extinct genus of dicynodont therapsid from the Late Permian of Inner Mongolia, China. The type species D. limbus was described in 1989 from a single skull found in the Naobaogou Formation. Daqingshanodon belongs to a group of dicynodonts called cryptodonts. It is the smallest known cryptodont, and the only one known from China. Like other cryptodonts, it has a pair of rounded nasal bosses above its nostrils and a ridge of bone on the upper jaw called the postcaniniform process. Daqingshanodon has a pair of elongated, recurved tusks extending from its beak-like snout. It is distinguished from other dicynodonts by the presence of a distinct ridge running along the side of the skull from below the eye socket to the area around the tusks. The skull of Daqingshanodon is less than 10 centimetres (3.9 in) long, yet this specimen is thought to have been an adult on the basis of its well-developed nasal bosses.
Jimusaria is an extinct genus of dicynodont therapsid from the Late Permian (Changhsingian) of China. The type species J. sinkianensis from the Guodikeng Formation in Xinjiang, was originally named as a species of Dicynodon, the first from Asia, but was given its own genus in 1963 before being sunk back into Dicynodon in 1988. The genus was resurrected in 2011 by palaeontologist Christian Kammerer in a taxonomic revision of the genus Dicynodon. Jimusaria was a mid-sized dicynodont, and was similar in appearance to the South African Dicynodon, but differed from it in features such as its narrower snout. A second species, Jimusaria monanensis was described from the Naobaogou Formation of northern China in 2023.
Turfanodon is an extinct genus of dicynodont therapsid from the Late Permian Sunan, Guodikeng, and Naobaogou Formations of China. The holotype of T. bogdaensis was discovered between 1963-1964 and was originally named in 1973 by A. Sun with the type species Turfanodon bogdaensis, Turfanodon was reclassified as a junior synonym of the related Dicynodon in 1988 by G. M. King. T. bogdaensis remained a species of Dicynodon for over two decades before the genus was reinstated in 2011 in a revision of the taxonomy of Dicynodon by palaeontologist Christian Kammerer. A second species from Inner Mongolia, T. jiufengensis, was named in 2021 by palaeontologist Jun Liu from a nearly complete skeleton and other referred bones. Turfanodon was a relatively large dicynodont, and similar in appearance to the related Daptocephalus from South Africa.
Kunpania is an extinct genus of dicynodont therapsid from the Quanzijie Formation of Xinjiang, China. The type and only species is K. scopulusa, and it is known only by a single incomplete specimen including parts of the skull and forelimb. Since its initial description in 1978 by palaeontologist Ailing Sun, it has sometimes been considered to be another species of Dicynodon by other researchers, or potentially undiagnostic. However, a redescription in 2021 reaffirmed its distinctiveness, including a uniquely well developed muscle attachment on the humerus. Kunpania is perhaps the oldest known member of the derived dicynodont group Dicynodontoidea, potentially dating to the Middle Permian period during the Capitanian, and so may fill a knowledge gap in the history of dicynodont evolution.
Counillonia is an extinct genus of dicynodont therapsid from the area of Luang Prabang in Laos, Southeast Asia that lived at around the time of the Permian-Triassic boundary and possibly dates to the earliest Early Triassic. Its type and only known species is C. superoculis. Counillonia was related to the Triassic dicynodonts such as Lystrosaurus and the Kannemeyeriiformes that survived the Permian mass extinction, but it was more closely related to the Permian genus Dicynodon than to either of these lineages. Counillonia may then possibly represent another line of dicynodonts that survived the Permian mass extinction into the Triassic period, depending on its age. The discovery of Counillonia in Laos and its unexpected evolutionary relationships hint at the less well understood geographies of dicynodont diversity across the Permo-Triassic boundary outside of well explored regions like the Karoo Basin in South Africa.
Taoheodon is an extinct genus of dicynodont therapsid from the Sunjiagou Formation in the Shanxi province of China, dated to the Wuchiapingian age of the Late Permian. Its type and only known species is T. baizhijuni. Taoheodon was a close relative of the well known Dicynodon, and may represent a biogeographical link between the South African Dicynodon and similar dicynodonts found in Laos.
