Ensifera | |
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A bush cricket or katydid | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Orthoptera |
Suborder: | Ensifera |
Superfamilies and families | |
See text |
Ensifera is a suborder of insects that includes the various types of crickets and their allies including: true crickets, camel crickets, bush crickets or katydids, grigs, weta and Cooloola monsters. This and the suborder Caelifera (grasshoppers and their allies) make up the order Orthoptera. Ensifera is believed to be a more ancient group than Caelifera, with its origins in the Carboniferous period, [2] the split having occurred at the end of the Permian period. [3] Unlike the Caelifera, the Ensifera contain numerous members that are partially carnivorous, feeding on other insects, as well as plants.
Ensifer is Latin for "sword bearer", and refers to the typically elongated and blade-like ovipositor of the females. [4]
Characteristics shared by the two orthopteran suborders, Caelifera and Ensifera, are the mouthparts adapted for biting and chewing, the modified prothorax, the hind legs modified for jumping, the wing shape and venation, and the sound-producing stridulatory organs. [2]
Ensiferans are distinguished from Caeliferans by their elongated, threadlike antennae, which are often longer than the length of their bodies and have over 30 segments (except in the subterranean Cooloolidae family). For this reason, they are sometimes referred to as "long-horned orthopterans". In the families in which the males sing, the fore wings have modifications that include toothed veins and scrapers for making the noise, and the surrounding membranous areas amplify the sound. In these groups, the sound-detecting tympanal organs are located on the tibiae of the front legs. [5] The tarsi have three segments and the ovipositor is blade-like or needle-like. The male attaches the spermatophore externally to the female's gonopore. The spermatophore is often surrounded by a proteinaceous spermatophylax, the function of which is to provide a nutritional nuptial gift to the female. [5] [6]
The Orthoptera Species File database lists the following superfamilies and families. [7]
The phylogenetic relationships of the Ensifera, summarized by Darryl Gwynne in 1995 from his own work and that of earlier authors, [lower-alpha 1] are shown in the following cladogram, with the Orthoptera divided into two main groups, Ensifera and Caelifera (grasshoppers). Fossil Ensifera are found from the late Carboniferous period onwards. [5] [10]
The oldest known fossil in the Archaeorthoptera, the crown group of the Orthoptera, and also the oldest member of the Pterygota (winged insects), is from the Namurian (324 mya) Lower Carboniferous beds in the Upper Silesian Basin of the Czech Republic. [11]
Orthoptera |
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Insects in the family Tettigoniidae are commonly called katydids, or bush crickets. They have previously been known as "long-horned grasshoppers". More than 6,400 species are known. Part of the suborder Ensifera, the Tettigoniidae are the only extant (living) family in the superfamily Tettigonioidea.
Orthoptera is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grasshoppers, locusts, and close relatives; and Ensifera – crickets and close relatives.
Anostostomatidae is a family of insects in the order Orthoptera, widely distributed in the southern hemisphere. It is named Mimnermidae or Henicidae in some taxonomies, and common names include king crickets in South Africa and wētā in New Zealand. Prominent members include the Parktown prawn of South Africa, and the giant wētā of New Zealand. The distribution of this family reflects a common ancestry before the fragmenting of Gondwana.
The orthopteran family Rhaphidophoridae of the suborder Ensifera has a worldwide distribution. Common names for these insects include the cave wētā, cave crickets, camelback crickets, camel crickets, spider crickets and sand treaders. Those occurring in New Zealand, Australia, and Tasmania are typically referred to as jumping or cave wētā. Most are found in forest environments or within caves, animal burrows, cellars, under stones, or in wood or similar environments. All species are flightless and nocturnal, usually with long antennae and legs. More than 1100 species of Rhaphidophoridae are described.
The family Gryllidae contains the subfamilies and genera which entomologists now term true crickets. They belong to the Orthopteran subfamily Ensifera, having long, whip-like antennae and has been reduced in terms of the older literature, with taxa such as the spider-crickets and allies, sword-tail crickets, wood or ground crickets and scaly crickets elevated to family level. The type genus is Gryllus and the first use of the family name "Gryllidae" was by Walker.
The family Prophalangopsidae are insects belonging to the order Orthoptera. They are the only extant members of the supefamily Hagloidea. There is only one extant genus in North America, where they are known as grigs, four genera in Asia, and many extinct genera.
The Caelifera are a suborder of orthopteran insects. They include the grasshoppers and grasshopper-like insects, as well as other superfamilies classified with them: the ground-hoppers (Tetrigoidea) and pygmy mole crickets (Tridactyloidea). The latter should not be confused with the mole crickets (Gryllotalpidae), which belong to the other Orthopteran sub-order Ensifera.
Gryllacrididae are a family of non-jumping insects in the suborder Ensifera occurring worldwide, known commonly as leaf-rolling crickets or raspy crickets. The family historically has been broadly defined to include what are presently several other families, such as Stenopelmatidae and Rhaphidophoridae, now considered separate. As presently defined, the family contains two subfamilies: Gryllacridinae and Hyperbaeninae. They are commonly wingless and nocturnal. In the daytime, most species rest in shelters made from folded leaves sewn with silk. Some species use silk to burrow in sand, earth or wood. Raspy crickets evolved the ability to produce silk independently from other insects, but their silk has many convergent features to silkworm silk, being made of long, repetitive proteins with an extended beta-sheet structure.
Crickets are Orthopteran insects which are related to bush crickets, and, more distantly, to grasshoppers. In older literature, such as Imms, "crickets" were placed at the family level, but contemporary authorities including Otte now place them in the superfamily Grylloidea. The word has been used in combination to describe more distantly related taxa in the suborder Ensifera, such as king crickets and mole crickets.
The Tridactylidae are a family in the insect order Orthoptera. They are small, mole-cricket-like insects, almost always less than 20 mm (0.79 in) long when mature. Generally they are shiny, dark or black, sometimes variegated or sandy-coloured. They commonly live in short tunnels and are commonly known as pygmy mole crickets, though they are not closely related to the true "mole crickets" (Ensifera), as they are included in the Caelifera suborder.
Grylloidea is the superfamily of insects, in the order Orthoptera, known as crickets. It includes the "true crickets", scaly crickets, wood crickets and other families, some only known from fossils.
Acrididea including the Acridomorpha is an infraorder of insects that describe the grasshoppers and ground-hoppers. It contains a large majority of species in the suborder Caelifera and the taxon Acridomorpha may also be used, which excludes the Tetrigoidea. Both names are derived from older texts, such as Imms, which placed the "short-horned grasshoppers" and locusts at the family level (Acrididae). The study of grasshopper species is called acridology.
The cohort Polyneoptera is a proposed taxonomic ranking for the Orthoptera and all other Neopteran insects believed to be more closely related to Orthoptera than to any other insect orders. These winged insects, now in the Paraneoptera, were formerly grouped as the Hemimetabola or Exopterygota on the grounds that they have no metamorphosis, the wings gradually developing externally throughout the nymphal stages.
The superfamily Hagloidea are insects belonging to the order Orthoptera: Ensifera; they are now represented by the extant Prophalangopsidae, with many extinct genera and families.
Tettigoniidea is an infraorder of the order Orthoptera, with six extant families.
Gryllidea is an infraorder that includes crickets and similar insects in the order Orthoptera. There are two superfamilies, and more than 6,000 described species in Gryllidea.
The infraorder Tridactylidea has a single extant superfamily which includes pygmy mole crickets; they are thought to represent living representatives of the most basal Caelifera: the Orthopteran suborder that includes grasshoppers.
The Gryllotalpoidea are a superfamily of insects that includes the mole crickets and the ant crickets. The type genus is Gryllotalpa.
The Phalangopsidae are a recently reconstituted family of crickets, based on the type genus PhalangopsisServille, 1831 from South America. Priority for family-group names based on this genus dates from Blanchard's "Phalangopsites".
Elcanidae are an extinct family of Mesozoic and early Cenozoic ensiferans. Members of the family are distinguished by the presence of spurs on the distal part of the metatibia, unique among orthopterans, these have been suggested to have been used for controlling gliding, swimming aids, or for jumping on water. They are known from the Late Triassic to Paleocene of Eurasia, North and South America.