Irbzer Formation

Last updated
Irbzer Formation
Stratigraphic range: Maastrichtian-Paleocene
~70–60  Ma
Type Geological formation
AreaFes-Boulmane
Lithology
PrimaryGray mudstone
Location
Coordinates 33°1′N4°8′W / 33.017°N 4.133°W / 33.017; -4.133
Approximate paleocoordinates 24°5′N5°2′W / 24.083°N 5.033°W / 24.083; -5.033
Region Fes-Boulmane
CountryMorocco
Morocco relief location map.jpg
Green-orange pog.svg
Irbzer Formation (Morocco)

The Irbzer Formation is a geological formation located in Morocco in the area of Fes-Boulmane of latest Cretaceous (Maastrichtian) to Paleocene age. The lower section of this formation consists of yellowish, cross-bedded calcareous sandstone containing shark teeth, while the layer above comprises phosphatic marls with vertebrate bone beds, including dinosaur remains. [1]

Contents

Paleoenvironment

Reconstructions of the "Phosphate sea" of Morocco. The Irbzer Formation area is seen at the NE Palaeogeographic models Phosphate basins Morocco - Herbig 1991.png
Reconstructions of the "Phosphate sea" of Morocco. The Irbzer Formation area is seen at the NE

During the Maastrichtian the inner area was alocated within a Anoxic marine setting, as revelated by bituminous, very oily and dolomitic black shells. [2] Palynology of the underliying and coeval El Koubbat Formation has revelated an microbiota composed of Dinoflagellates, Pterospermopsis , Scolecodonts and Tasmanaceae. [3]

The formation reflects a shallow marine to lagoonal setting, transitioning from fine sands to marls and limestones. The presence of marine organisms, including bivalves and gastropods, along with phosphatic layers containing fish remains, indicates periodic marine incursions, with intermittent lagoonal or nearshore conditions. [4] Oxidation and sedimentary discontinuities suggest episodes of exposure or subaerial erosion. The overall environment was likely a shallow, low-energy, marginal marine setting with occasional deeper marine influences. [4] No planktonic foraminifera are present, and the few benthic foraminifera offer limited ecological insights. However, ostracodes, though poorly preserved, are more abundant and provide significant information about the environment. The ostracode assemblage including indicates a shallow (0–20 m), oligohaline, moderately alkaline, and relatively warm limnic environment, likely permanent and near the shoreline, as suggested by the presence of isolated dinosaur teeth and eggshell fragments in these layers. [4] [5]

Marine Amniote fauna is know from coeval layers of the El Koubbat Formation, mostly Mosasaurs (Halisaurinae, Gavialimimus , Globidens , Mosasaurus , Prognathodontini) and Elasmosauridae plesiosaurs. [6]


Biota

Color key
Taxon Reclassified taxonTaxon falsely reported as presentDubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Charophytes

GenusSpeciesLocationMaterialNotesImages
Feistiella [7]
  • F. sp.
Oudiksou synclineGyrogonitesA charophyte of the family Feistiellaceae
Lamprothamnium [7]
  • L. sp.
Oudiksou synclineGyrogonitesA charophyte of the family Characeae
Sphaerochara [7]
  • S. cf. edda
Oudiksou synclineGyrogonitesA charophyte of the family Characeae
Stephanochara [7]
  • S. sp.
Oudiksou synclineGyrogonitesA charophyte of the family Characeae

Ostracods

GenusSpeciesLocationMaterialNotesImages
Brachycythere [4]
  • B. cf. gr. sapucariensis
El KoubbatValvesA marine Ostracod of the family Brachycytheridae
Clithrocytheridea [4]
  • C. senegali
El KoubbatValvesA marine Ostracod of the family Cytherideidae
Costa [4]
  • C. sp
El KoubbatValvesA marine Ostracod of the family Trachyleberididae
Cytheretta [4]
  • C.? koubbatensis
El KoubbatValvesA marine Ostracod of the family Cytherettidae
Cytherella [4]
  • C. sp
El KoubbatValvesA marine Ostracod of the family Cytherellidae
Gomphocythere [5]
  • G. achloujensis
AchloujValvesA freshwater Ostracod of the family Limnocytheridae
Oertliella [4]
  • O. sp.
El KoubbatValvesA marine Ostracod of the family Trachyleberididae
Rehacythereis [4]
  • R. sp
El KoubbatValvesA marine Ostracod of the family Trachyleberididae
Paracandona [5]
  • P. occitanica
AchloujValvesA freshwater Ostracod of the family Candonidae
Paracypris [4]
  • P. cf. mdaouerensis
  • P. sp.
  • P.? sp.
El KoubbatValvesA marine Ostracod of the family Paracyprididae

Echinoderms

GenusSpeciesLocationMaterialNotesImages
Cretasterias [8]
  • C. reticulatus
BakritThousands of specimensA sea star of the family Asteriidae

Fish

GenusSpeciesLocationMaterialNotesImages
Cretalamna [1]
  • C. maroccana
Oukdiksou synclineTeethA Shark of the family Otodontidae
Cretalmana reconstruction Cretalamna reconstruction.png
Cretalmana reconstruction
Dalpiazia [1]
  • D. strollleri
Oukdiksou synclineTeethA Chondrichthian of the family Sclerorhynchidae
Ganopristis [1]
  • G. leptodoll
Oukdiksou synclineTeethA Chondrichthian of the family Sclerorhynchidae
Serratolamna [1]
  • S. serrata
Oukdiksou synclineTeethA Shark of the family Serratolamnidae
Squalicorax [1]
  • S. pristodontus
  • S. bassallii
Oukdiksou synclineTeethA Shark of the family Anacoracidae
Squalicorax reconstruction Squalicorax2DB.jpg
Squalicorax reconstruction

Crocodrylomorpha

GenusSpeciesLocationMaterialNotesImages
Krokolithidae [1]
  • Indeterminate
Oukdiksou synclineeggshellsIndeterminate Crocodrilian eggshells

Sauropoda

GenusSpeciesLocationMaterialNotesImages
Pseudomegaloolithus [1]
  • P. atlasi
Oukdiksou synclineeggshellsIndeterminate Sauropod eggshells
Megaloolithus [1]
  • M. maghrebiensis
  • M. sp.
Oukdiksou synclineeggshellsEggshells referred to Titanosauridae sauropods
Example of Megaloolithus Underside of a clutch of Megaloolithus siruguei at Pinyes locality - Tremp Formation.jpg
Example of Megaloolithus

Theropoda

GenusSpeciesLocationMaterialNotesImages
Pseudogeckoolithus [1]
  • P. tirboulensis
Oukdiksou synclineeggshellsTheropod eggs referred to Maniraptora
Rodolphoolithus [1]
  • R. arioul
Oukdiksou synclineeggshellsIndeterminate eggs likely laid by small theropods
" Paronychodon " [1]
  • "P. sp."
Oukdiksou synclineTeethA theropod, probably related with Troodontidae or Unenlagiinae
Tipoolithus [1]
  • T. achloujensis
Oukdiksou synclineeggshellsTheropod eggs, referred to an enantiornithine bird or small theropod

Related Research Articles

<span class="mw-page-title-main">Mosasaur</span> Extinct marine lizards of the Late Cretaceous

Mosasaurs are an extinct group of large aquatic reptiles within the family Mosasauridae that lived during the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes.

<i>Mosasaurus</i> Extinct genus of marine squamate reptile from the Late Cretaceous

Mosasaurus is the type genus of the mosasaurs, an extinct group of aquatic squamate reptiles. It lived from about 82 to 66 million years ago during the Campanian and Maastrichtian stages of the Late Cretaceous. The genus was one of the first Mesozoic marine reptiles known to science—the first fossils of Mosasaurus were found as skulls in a chalk quarry near the Dutch city of Maastricht in the late 18th century, and were initially thought to be crocodiles or whales. One skull discovered around 1780 was famously nicknamed the "great animal of Maastricht". In 1808, naturalist Georges Cuvier concluded that it belonged to a giant marine lizard with similarities to monitor lizards but otherwise unlike any known living animal. This concept was revolutionary at the time and helped support the then-developing ideas of extinction. Cuvier did not designate a scientific name for the animal; this was done by William Daniel Conybeare in 1822 when he named it Mosasaurus in reference to its origin in fossil deposits near the Meuse River. The exact affinities of Mosasaurus as a squamate remain controversial, and scientists continue to debate whether its closest living relatives are monitor lizards or snakes.

<span class="mw-page-title-main">Maastrichtian</span> Sixth and last age of the Late Cretaceous

The Maastrichtian is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from 72.1 to 66 million years ago. The Maastrichtian was preceded by the Campanian and succeeded by the Danian.

<span class="mw-page-title-main">Mosasaurinae</span> Subfamily of reptiles

The Mosasaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "mosasaurines" and their fossils have been recovered from every continent except for South America.

<i>Globidens</i> Extinct genus of lizards

Globidens is an extinct genus of mosasaurid oceanic lizard classified as part of the Globidensini tribe in the Mosasaurinae subfamily. Globidens belongs to the family Mosasauridae, which consists of several genera of predatory marine lizards of various sizes that were prevalent during the Late Cretaceous. Specimens of Globidens have been discovered in Angola, Brazil, Morocco, Syria and the United States. Among mosasaurs, Globidens is probably most well known for the highly rounded, globe-like teeth that give it its name.

The Coon Creek Formation or Coon Creek Tongue is a geologic unit and Konservat-Lagerstätte located in western Tennessee and extreme northeast Mississippi. It is a sedimentary sandy marl deposit, Late Cretaceous (Maastrichtian) in age, about 70 million years old. The formation is renowned for its pristine fossils of Late Cretaceous marine invertebrates, including gastropods, bivalves, decapod crustaceans, and ammonites, particularly at Coon Creek in McNairy County, Tennessee, which the formation is named for. It is also known for producing fosslis of marine vertebrates, such as mosasaurs and plesiosaurs. Notable fossils from this formation is the gastropod Turritella, the bivalve Pterotrigonia thoracica, as well as other fossils such as crabs.

<i>Pluridens</i> Extinct genus of lizards

Pluridens is an extinct genus of marine lizard belonging to the Mosasauridae. Pluridens is placed in the subfamily Halisaurinae with the genera Phosphorosaurus, Eonatator and Halisaurus. Compared to related halisaurines, Pluridens had longer jaws with more teeth, and smaller eyes. It also grew large size, measuring 5–6 m (16–20 ft) long and perhaps over 9 m (30 ft) in some individuals. The jaws in some specimens are robust, and sometimes show injuries suggestive of combat. The jaws may have been used for fighting over mates or territories.

<span class="mw-page-title-main">Aganane Formation</span> Geologic formation in Azilal Province, central Morocco

The Aganane Formation is a Pliensbachian geologic formation in the Azilal, Béni-Mellal, Ouarzazate, Tinerhir and Errachidia provinces, central Morocco, being the remnant of a local massive Carbonate platform, and known mostly for its rich tracksites including footprints of thyreophoran, sauropod and theropod dinosaurs. It may also include the fossiliferous levels of the Calcaires du Bou Dahar, if true, it would be one of the richest Early Jurassic formations in the entire tethys area.

<span class="mw-page-title-main">Ouled Abdoun Basin</span> Phosphate basin in Morocco

The Oulad Abdoun Basin is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years.

<i>Kaikaifilu</i> Extinct genus of marine squamate reptiles

Kaikaifilu is an extinct genus of large mosasaurs that lived during the Maastrichtian stage of the Late Cretaceous, in what is now northern Antarctica. The only species known, K. hervei, was described in 2017 from an incomplete specimen discovered in the López de Bertodano Formation, in Seymour Island, Antarctic Peninsula. The taxon is named in reference to Coi Coi-Vilu, a reptilian ocean deity of the Mapuche cosmology. Early observations of the holotype classify it as a member of the subfamily Tylosaurinae. However, later observations note that several characteristics show that this attribution is problematic.

<i>Chenanisaurus</i> Extinct genus of dinosaurs

Chenanisaurus is a genus of predatory abelisaurid dinosaur, with a single known species C. barbaricus. It comes from the upper Maastrichtian phosphates of the Ouled Abdoun Basin in Morocco, North Africa. The animal is known from a holotype, consisting of a partial jaw bone, and several isolated teeth found in the same beds. Chenanisaurus is one of the largest members of the Abelisauridae, and one of the last, being a contemporary of the North American Tyrannosaurus. It would have been among the dinosaur species wiped out by the Chicxulub asteroid impact and the Cretaceous-Paleogene mass extinction that followed.

<i>Alcione elainus</i> Genus of pterosaur from the Late Cretaceous

Alcione is a genus of pterosaur from the Ouled Abdoun Basin of Morocco, dating back to the Maastrichtian stage of the Late Cretaceous period. Only one species, A. elainus, is known. This pterosaur lived in a marine environment alongside several other pterosaurs, including Simurghia and Barbaridactylus.

The El Mers Group is a geological group in the Middle Atlas of Morocco. It is subdivided into three formations named the El Mers I, II, and III Formations, respectively. It is a marine deposit primarily consisting of marl, with gypsum present in the upper part of unit 3. It is the lateral equivalent of the terrestrial Guettioua Sandstone. Dinosaur remains are among the fossils that have been recovered from the group, most notably those of sauropods and the unusual thyreophorans Adratiklit, Thyreosaurus, and Spicomellus.

<span class="mw-page-title-main">Azilal Formation</span> Geological unit in Morocco

The Azilal Formation, also known as "Aguerd-nˈTazoult" Formation,Toundoute Continental Series and "Wazzant" Formation, is a geological unit in the Azilal, Béni-Mellal, Imilchil, Zaouiat Ahansal, Ouarzazate, Tinerhir and Errachidia areas of the High Atlas of Morocco, part of the Tafraout Group that covers the Early Toarcian to Middle Aalenian stages of the Jurassic Period. While there have been atributions of it´s lowermost layers to the Latest Pliensbachian, the current oldest properly measured are part of the Earliest Toarcian regresion ("MRST10"), part of the Lower-Middle Palymorphum biozone. The Azilal Formation consists mainly of claystones rich in continental plant debris and laminated microbial facies. It is a continental deposit which overlies marine dolomites of equivalent age to the Rotzo Formation of Italy, mostly part of the Aganane Formation. The formation is the continental to marginal marine part of a massive Siliciclastic-Carbonate platform, best assigned to an alluvial environment occasionally interrupted by shallow marine incursions and marks a dramatic decrease of the carbonate productivity under increasing terrigenous sedimentation. Dinosaur remains, such the sauropod Tazoudasaurus and the basal ceratosaur Berberosaurus are known from the unit, along with several undescribed genera. The toarcian High Atlas is divided in 5 units: the continental layers with paralic deposits belong to the Azilal, along the shoreface layers of the Tagoudite Formation & Tafraout Formation, connected with the offshore Ait Athmane Formation and the deeper shelf deposits of the Agoudim 1 Formation.

<span class="mw-page-title-main">Kristianstad Basin</span> Cretaceous geological formation in Skåne, Sweden

The Kristianstad Basin is a Cretaceous-age structural basin and geological formation in northeastern Skåne, the southernmost province of Sweden. The basin extends from Hanöbukten, a bay in the Baltic Sea, in the east to the town of Hässleholm in the west and ends with the two horsts Linderödsåsen and Nävlingeåsen in the south. The basin's northern boundary is more diffuse and there are several outlying portions of Cretaceous-age sediments. During the Cretaceous, the region was a shallow subtropical to temperate inland sea and archipelago.

<i>Xenodens</i> Extinct genus of lizards

Xenodens is an extinct genus of marine lizard belonging to the mosasaur family. It currently contains a single species, X. calminechari, which is known from Late Maastrichtian phosphate deposits in the Ouled Abdoun Basin, Morocco. Its closest known relative is believed to be the durophagous Carinodens.

<i>Spicomellus</i> Extinct genus of dinosaurs

Spicomellus is an extinct genus of early ankylosaurian dinosaur from the El Mers III Formation (Bathonian-Callovian) of Morocco. The genus contains a single species, S. afer, known from a single rib with fused osteoderms. Spicomellus represents the oldest named ankylosaur.

<i>Thalassotitan</i> Large bodied African mosasaur

Thalassotitan is an extinct genus of large mosasaurs that lived during the late Maastrichtian of the Cretaceous period in what is now Morocco, around 67 to 66 million years ago. The only known species is T. atrox, described in 2022 from fossils discovered in the Ouled Abdoun Basin, initially identified as coming from other genera such as Mosasaurus or Prognathodon. Hypothetical Thalassotitan specimens may have been found in other corners of the world, although researchers also note the possibility that they come from distinct, related taxa. It is considered to be close to the genera Prognathodon and Gnathomortis, together forming the tribe Prognathodontini. The prognathodontines are separated from other mosasaurs based on their massive jaws and robust teeth.

<span class="mw-page-title-main">Tafraout Group</span> Geological formations in Morocco

The Tafraout Group is a geological group of formations of Toarcian-Aalenian age in the Azilal, Béni-Mellal, Imilchil, Zaouiat Ahansal, Ouarzazate, Tinerhir and Errachidia areas of the High Atlas of Morocco. The Group represents the remnants of a local massive Siliciclastic-Carbonate platform, best assigned to succession W-E of alluvial environment occasionally interrupted by shallow marine incursions and inner platform to open marine settings, and marks a dramatic decrease of the carbonate productivity under increasing terrigenous sedimentation. Fossils include large reef biotas with richness in "lithiotid" bivalves and coral mounts, but also by remains of vertebrates such as the sauropod Tazoudasaurus and the basal ceratosaur Berberosaurus, along with several undescribed genera. While there have been attributions of its lowermost layers to the Latest Pliensbachian, the current oldest properly measured are part of the Earliest Toarcian regression ("MRST10"), part of the Lower-Middle Palymorphum biozone. This group is composed of the following units, which extend from west to east: the Azilal Formation ; the "Amezraï" Formation ; the Tafraout Formation & the Tagoudite Formation. They are connected with the offshore Ait Athmane Formation and the deeper shelf deposits of the Agoudim 1 Formation. Overall, this group represents a mixed carbonate-siliciclastic system of several hundred meters thick, dominated by deposits of shallow marine platforms linked to a nearby hinterland dominated by conglomerates. The strata of the group extend towards the central High Atlas, covering different anticlines and topographic features along the mountain range.

References

  1. 1 2 3 4 5 6 7 8 9 10 11 12 13 Garcia, G.; Tabuce, R.; Cappetta, H.; Marandat, B.; Bentaleb, I.; Benabdallah, A.; Vianey-Liaud, M. (2003). "First record of dinosaur eggshells and teeth from the North-West African Maastrichtian (Morocco)". Palaeovertebrata. 32 (2–4): 59–69.
  2. Jacobshagen, Volker H. (1988). "The Atlas System of Morocco Volume 15 || Synsedimentary tectonics in the Northern Middle Atlas (Morocco) during the late Cretaceous and Tertiary". Studies on Its Geodynamic Evolution. 18: 321–337. doi:10.1007/BFb0011599.
  3. Rauscher, R.; Schuler, M.; Benalioulhaj, N. (1986). "Les Conditions de Depôt des Schistes Bitumineux et des Phosphates Maastrichtiens au Maroc: une Approche Palynologique". Documents du Bureau de Recherches Géologiques et Minières. 110 (1): 127–139.
  4. 1 2 3 4 5 6 7 8 9 10 11 Andreu, Bernard; Tronchetti, Guy (1996). "Ostracodes et foraminifèresdu Crétacé supérieur du synclinal d'El Koubbat, Moyen Atlas, Maroc". Geobios. 29 (1): 45–71. Bibcode:1996Geobi..29...45A. doi:10.1016/s0016-6995(96)80071-4. ISSN   0016-6995.
  5. 1 2 3 Colin, Jean-Paul; Tabuce, Rodolphe (2004). "Ostracodes limniques de la Formation d'Irbzer, Crétacé terminal du Moyen-Atlas, Maroc: taxonomie, biostratigraphie, paléoécologie, paléobiogéographie". Revue de Micropaléontologie. 47 (2): 103–109. Bibcode:2004RvMic..47..103C. doi:10.1016/j.revmic.2004.04.001. ISSN   0035-1598.
  6. Vinkeles Melchers, Alexander P. M.; Martens, Brennan P.; Rempert, Trevor H. "The marine reptile fauna of the Bekrit syncline (El Koubbat Formation, lower Maastrichtian), Middle Atlas Mountains, Morocco". 7th Triennial Mosasaur Meeting – A Global Perspective on Mesozoic Marine AmniotesAt: Maastricht, the Netherlands.
  7. 1 2 3 4 Schudack, Michael; Herbig, H.G. (1995). "Charophytes from the Cretaceous -Tertiary Boundary Beds of the Middle Atlas Mountains, Morocco". Géologie Méditerranéenne. 22 (3): 125–139. doi:10.3406/geolm.1995.1574. ISSN   0397-2844.
  8. GALE, ANDY; VILLIER, LOÏC (2013). "Mass mortality of an asteriid starfish (Forcipulatida, Asteroidea, Echinodermata) from the late Maastrichtian (Late Cretaceous) of Morocco". Palaeontology. 56 (3): 577–588. Bibcode:2013Palgy..56..577G. doi:10.1111/pala.12002. ISSN   0031-0239.