Southern cassowary | |
---|---|
Wild individual seen in Etty Bay, Queensland, Australia | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Infraclass: | Palaeognathae |
Order: | Casuariiformes |
Family: | Casuariidae |
Genus: | Casuarius |
Species: | C. casuarius |
Binomial name | |
Casuarius casuarius | |
Native distribution of the southern cassowary |
The southern cassowary (Casuarius casuarius), also known as double-wattled cassowary, Australian cassowary, or two-wattled cassowary, is a large flightless black bird, found in Indonesia, Papua New Guinea, and northeastern Australia. It is one of the three living species of cassowary, alongside the dwarf cassowary and the northern cassowary. It is a ratite and therefore related to the emu, ostrich, rhea and kiwi.
The Australian population is listed as Endangered under federal and Queensland state legislation.
Presently, most authorities consider the southern cassowary monotypic, but several subspecies have been described. [3] It has proven very difficult to confirm the validity of these due to individual variations, age-related variations, the relatively few available specimens (and the bright skin of the head and neck – the basis upon which several subspecies have been described – fades in specimens), and that locals are known to have traded live cassowaries for hundreds, if not thousands of years, some of which are likely to have escaped/been deliberately introduced to regions away from their origin. [3]
Cassowaries are most closely related to the kiwis, both families diverging from a common ancestor approximately 40 million years ago. [3]
The binomial name Casuarius casuarius is derived from its Malay name kesuari. [4] The southern cassowary was first described by Carl Linnaeus, in his 18th-century work Systema Naturae , as Struthio casuarius, [5] from a specimen from Seram, in 1758. [6] It is now the type species of the genus Casuarius . [6] Common names for the species include southern cassowary, double-wattled cassowary, Australian cassowary, [2] and two-wattled cassowary. [6]
The southern cassowary has been described under a large number of scientific names, all of which are now considered taxonomic synonyms for the species. [7]
Synonyms |
---|
Struthio casuariusLinnaeus 1758 |
Casuarius casuarius altijugusSclater 1878 |
Casuarius altijugusSclater 1878 |
Casuarius casuarius aruensisSchlegel 1866 |
Casuarius aruensisSchlegel 1866 |
Casuarius australisWall 1854 |
Casuarius casuarius beccariiSclater 1875 |
Casuarius beccariiSclater 1875 |
Casuarius bicarunculatusSclater 1860 |
Casuarius casuarius bicarunculatusSclater 1860 |
Casuarius bistriatusvan Oort 1907 |
Casuarius casuarius bistriatusvan Oort 1907 |
Casuarius casuarius casuariusLinnaeus 1758 |
Casuarius casuarius chimaeraRothschild 1904 |
Cassowara eximiaPerry 1811 |
Casuarius casuarius grandisRothschild 1937 |
Casuarius galeatusBonnaterre 1790 |
Casuarius casuarius hamiltoniMathews 1915 |
Casuarius casuarius intensusRothschild 1898 |
Casuarius bicarunculatus intermediusRothschild 1928 |
Casuarius casuarius intermediusRothschild 1928 |
Casuarius casuarius johnsoniiMüller 1866 |
Casuarius johnsoniiMüller 1866 |
Casuarius casuarius lateralisRothschild 1925 |
Casuarius casuarius salvadoriiOustalet 1878 |
Casuarius salvadoriiOustalet 1878 |
Casuarius casuarius sclateriiSalvadori 1878 |
Casuarius sclateriiSalvadori 1878 |
Casuarius casuarius tricarunculatusBeccari 1876 |
Casuarius bicarunculatus tricarunculatusBeccari 1876 |
Casuarius tricarunculatusBeccari 1876 |
Casuarius casuarius violicollisRothschild 1899 |
Hippalectryo indicusGloger 1842 |
Casuarius hagenbeckiRothschild 1904 |
The southern cassowary has stiff, bristly black plumage, a blue face and a long neck, red on the cape and two red wattles measuring around 17.8 cm (7.0 in) in length hanging down around its throat. A horn-like brown casque, measuring 13 to 16.9 cm (5.1 to 6.7 in) high, sits atop the head. The bill can range from 9.8 to 19 cm (3.9 to 7.5 in). The plumage is sexually monomorphic, but the female is dominant and larger with a longer casque, larger bill and brighter-coloured bare parts. The juveniles have brown longitudinal striped plumage. [6]
The three-toed feet are thick and powerful, equipped with a lethal dagger-like claw up to 12 cm (4.7 in) on the inner toe. It is perhaps the largest member of the cassowary family and is tied as the third heaviest bird on earth (after the Somali ostrich and the common ostrich), at a maximum size estimated at 85 kg (187 lb) and 190 cm (6 ft 3 in) tall. Normally, this species ranges from 127 to 170 cm (4 ft 2 in to 5 ft 7 in) in length. [6] The height is normally 150 to 180 cm (4 ft 11 in to 5 ft 11 in) ; females average 58.5 kg (129 lb), while males average 29 to 34 kg (64–75 lb). The northern cassowary is about the same size on average and is perhaps very mildly less sexually dimorphic than the southern. [6] Most adult birds will weigh between 17 and 70 kg (37 and 154 lb). [8] It is technically the largest Asian bird (since the extinction of the Arabian ostrich) and the largest Australian bird (though the emu may be slightly taller).
The southern cassowary is distributed in Indonesia, Papua New Guinea, and northeastern Australia. [9] It mainly inhabits tropical rainforests but may make use of nearby savannah forests or mangroves stands. [1] The species prefers elevations below 1,100 m (3,600 ft) in Australia, [6] and 500 m (1,600 ft) on New Guinea. [1]
As of 2018 [update] , according to Birdlife International, the breeding populations were as follows: [1]
Location | Population | Trend |
---|---|---|
Southern New Guinea | Unknown | Declining |
Seram, Indonesia | Unknown | Unknown |
Aru Islands, Indonesia | Unknown | Unknown |
Northeastern Australia | 1,500 to 2,500 | Declining |
Paluma Range, Qld, Australia | Unknown | Declining |
McIlwraith Range, Qld, Australia | 1,000+ | Declining |
Apudthama National Park, Qld, Australia | Unknown | Unknown |
Total | 4,000+ | Declining |
Southern cassowaries forage on the forest floor for fallen fruit and seeds and are capable of safely digesting some fruits and seeds toxic to other animals. They also eat fungi, and some insects, other invertebrates (such as snails), small vertebrates (such as fish, frogs, small birds and their eggs and nestlings, and small mammals), and carrion. When food is scarce, both wild and captive cassowaries have been seen to ingest earth, perhaps seeking supplementary minerals. [10] Inspection of the faeces reveals that commonly ingested fruits are Davidsonia pruriens , Syzygium divaricatum , and members of the palm (Arecaceae), quandong (Elaeocarpaceae), laurel (Lauraceae), and myrtle (Myrtaceae) families. [11] [10]
The southern cassowary is a solitary bird, which pairs only in breeding season, in late winter or spring. The male builds a nest on the ground, [6] a mattress of herbaceous plant material 5 to 10 centimetres (2–4 in) thick and up to 100 centimetres (39 in) wide. This is thick enough to let moisture drain away from the eggs. The male also incubates the eggs and raises the chicks alone. A clutch of three or four eggs are laid measuring 138 by 95 millimetres (5.4 in × 3.7 in). They have a granulated surface and are initially bright pea-green in colour although they fade with age. [6] [12] Southern cassowaries make a thunderous call during mating season, and hissing and rumblings otherwise. Chicks will make frequent high-pitched contact whistles and chirps to call the male. [1]
Southern cassowaries have a reputation for being dangerous to humans and other animals, and are often regarded as aggressive. The birds can jump quite high and kick powerfully with their blade-like claws. However, deadly encounters with southern cassowaries are rare. Only two human deaths have been reported since 1900. A 2003 historical study of 221 southern cassowary attacks showed that 150 had been against humans: 75% of these had been from southern cassowaries that had been fed by people, 71% of the time the bird had chased or charged the victim, 15% of the time they kicked. Of the attacks, 73% involved the birds expecting or snatching food, 5% involved defending their natural food sources, 15% involved defending themselves from attack, and 7% involved defending their chicks or eggs. Only one human death was reported among those 150 attacks. [13]
The first documented human death caused by a southern cassowary was on 6 April 1926. In Australia, 16-year-old Phillip McClean and his brother, age 13, came across a southern cassowary on their property and decided to try and kill it by striking it with clubs. The bird kicked the younger boy, who fell and ran away as his older brother struck the bird. The older McClean then tripped and fell to the ground. While he was on the ground, the cassowary kicked him in the neck, opening a 1.25 cm (0.5 in) wound that may have severed his jugular vein. The boy died of his injuries shortly thereafter. [14]
Another human death due to a southern cassowary was recorded in Florida, United States on 12 April 2019. The bird's owner, a 75-year-old man who had raised the animal, was apparently clawed to death after he fell to the ground. [15]
Being fed by people tempts southern cassowaries into closer associations with human-inhabited areas, increasing the already high risk of vehicle strikes – a major cause of southern cassowary mortality – and increasing the likelihood of encounters with humans. [16]
In a 2017 Australian Birdlife article, Karl Brandt suggested Aboriginal encounters with the southern cassowary may have inspired the myth of the bunyip. [17]
Although subject to ongoing habitat loss (some due to logging), limited range, and overhunting in some areas, the southern cassowary was evaluated as Least concern in 2018 by the International Union for Conservation of Nature (IUCN). Other threats include feral animals eating their eggs and roadkill. [1] [6] Road-building, feral animals and hunting are the worst of these threats. It has an occurrence range of 396,000 km2 (153,000 sq mi), and between 10,000 and 20,000 birds were estimated in a 2002 study, with between 1,500 and 2,500 in Australia. [1]
The Australian population is listed as Endangered under federal (EPBC Act). Under the Nature Conservation (Animals) Regulation 2020 (Queensland), November 2022 list, the northern population is considered vulnerable, while the southern population remains as endangered. [2] A draft recovery plan to save the species was published by the federal government in June 2023, at which time there were estimated to be around 5,000 individuals in Australia. [18] A study published in Biological Conservation in March 2023 listed 23 species which the authors considered to no longer meet the criteria as threatened species under the EPBC Act, including the Southern cassowary. A 2021 study had shown that extensive reservation had led to recovery of the species, and legislation had prevented the previously rapid loss of habitat. The reason for their assessment was given as "Populations now stable or declining at a rate less than threshold". [19] The team, led by John Woinarski of Charles Darwin University looked at all species listed as threatened under the act in 2000 and 2022. [20]
Southern cassowaries have been bred in zoos around the world, including White Oak Conservation in Yulee, Florida, United States. [21]
The common ostrich, or simply ostrich, is a species of flightless bird native to certain large areas of Africa. It is one of two extant species of ostriches, the only living members of the genus Struthio in the ratite order of birds. The other is the Somali ostrich, which was recognized as a distinct species by BirdLife International in 2014 having been previously considered a distinctive subspecies of ostrich.
Cassowaries are flightless birds of the genus Casuarius in the order Casuariiformes. They are classified as ratites: flightless birds without a keel on their sternum bones. Cassowaries are native to the tropical forests of New Guinea, The Moluccas, and northeastern Australia.
The emu is a species of flightless bird endemic to Australia, where it is the tallest native bird. It is the only extant member of the genus Dromaius and the third-tallest living bird after its African ratite relatives, the common ostrich and Somali ostrich. The emu's native ranges cover most of the Australian mainland. The Tasmanian, Kangaroo Island and King Island subspecies became extinct after the European settlement of Australia in 1788.
Ratites are a polyphyletic group consisting of all birds within the infraclass Palaeognathae that lack keels and cannot fly. They are mostly large, long-necked, and long-legged, the exception being the kiwi, which is also the only nocturnal extant ratite.
Tinamous are members of the order Tinamiformes, and family Tinamidae, divided into two distinct subfamilies, containing 46 species found in Mexico, Central America, and South America. The word "tinamou" comes from the Galibi term for these birds, tinamu. Tinamous are the only living group of palaeognaths able to fly, and were traditionally regarded as the sister group of the flightless ratites, but recent work places them well within the ratite radiation as most closely related to the extinct moa of New Zealand, implying flightlessness emerged among ratites multiple times. Tinamous first appear in the fossil record in the Miocene epoch. They are generally sedentary, ground-dwelling and, though not flightless, when possible avoid flight in favour of hiding or running away from danger. They are found in a variety of habitats, ranging from semi-arid alpine grasslands to tropical rainforests. The two subfamilies are broadly divided by habitat, with the Nothurinae referred to as steppe or open country tinamous, and the Tinaminae known as forest tinamous.
The great white pelican also known as the eastern white pelican, rosy pelican or simply white pelican is a bird in the pelican family. It breeds from southeastern Europe through Asia and Africa, in swamps and shallow lakes.
The wattled crane is a large, threatened species of crane found in wetlands and grasslands of eastern and southern Africa, ranging from Ethiopia to South Africa. Some authorities consider it the sole member of the genus Bugeranus.
Flightless birds have, through evolution, lost the ability to fly. There are over 60 extant species, including the well-known ratites and penguins. The smallest flightless bird is the Inaccessible Island rail. The largest flightless bird, which is also the largest living bird in general, is the common ostrich.
The African oystercatcher or African black oystercatcher is a large charismatic wader resident to the mainland coasts and offshore islands of southern Africa. This near-threatened oystercatcher has a population of over 6,000 adults, which breed between November and April. The scientific name moquini commemorates the French naturalist Alfred Moquin-Tandon who discovered and named this species before Bonaparte.
The great spotted kiwi, great grey kiwi or roroa is a species of kiwi endemic to the South Island of New Zealand. The great spotted kiwi, as a member of the ratites, is flightless. It is the largest of the kiwi. The rugged topography and harsh climate of the high altitude alpine part of its habitat render it inhospitable to a number of introduced mammalian predators, which include dogs, ferrets, cats, and stoats. Because of this, populations of this species have been less seriously affected by the predations of these invasive species compared to other kiwi. Nonetheless, there has been a 43% decline in population in the past 45 years, due to these predators and habitat destruction. This has led it to be classified as vulnerable. There are less than 16,000 great spotted kiwis in total, almost all in the more mountainous parts of northwest Nelson, the northwest coast, and the Southern Alps. A minority live on island reserves.
The Australian brushturkey, Australian brush-turkey, or gweela, also frequently called the scrub turkey or bush turkey, is a common, widespread species of mound-building bird from the family Megapodiidae found in eastern Australia from Far North Queensland to Eurobodalla on the South Coast of New South Wales. The Australian brushturkey has also been introduced to Kangaroo Island in South Australia. It is the largest extant representative of the family Megapodiidae, and is one of three species to inhabit Australia.
The Casuariiformes is an order of large flightless birds that has four surviving members: the three species of cassowary, and the only remaining species of emu. They are divided into either a single family, Casuariidae, or more typically two, with the emu splitting off into its own family, Dromaiidae.
Darwin's rhea or the lesser rhea is a large flightless bird, the smaller of the two extant species of rheas. It is found in the Altiplano and Patagonia in South America.
The greater rhea is a species of flightless bird native to eastern South America. Other names for the greater rhea include the grey, common, or American rhea; ema (Portuguese); or ñandú. One of two species in the genus Rhea, in the family Rheidae, it inhabits a variety of open areas, such as grasslands, savanna or grassy wetlands. Weighing 20–27 kilograms (44–60 lb), the greater rhea is the largest native bird in the Americas. In the wild, the greater rhea has a life expectancy of 10.5 years. It is also notable for its reproductive habits, and for the fact that a population has established itself in Northern Germany in recent years. The species is listed as Near Threatened by the IUCN.
The dwarf cassowary, also known as Bennett's cassowary, little cassowary, mountain cassowary or muruk, is the smallest of the three extant species of cassowaries.
The northern cassowary, also known as the one-wattled cassowary, single-wattled cassowary, or golden-necked cassowary, is a large, stocky flightless bird of northern New Guinea. It is one of the three living species of cassowary, alongside the dwarf cassowary and the southern cassowary. It is a member of the superorder Paleognathae.
Palaeognathae is an infraclass of birds, called paleognaths or palaeognaths, within the class Aves of the clade Archosauria. It is one of the two extant infraclasses of birds, the other being Neognathae, both of which form Neornithes. Palaeognathae contains five extant branches of flightless lineages, termed ratites, and one flying lineage, the Neotropic tinamous. There are 47 species of tinamous, five of kiwis (Apteryx), three of cassowaries (Casuarius), one of emus (Dromaius), two of rheas (Rhea) and two of ostriches (Struthio). Recent research has indicated that paleognaths are monophyletic but the traditional taxonomic split between flightless and flighted forms is incorrect; tinamous are within the ratite radiation, meaning flightlessness arose independently multiple times via parallel evolution.
The blue petrel is a small seabird in the shearwater and petrel family, Procellariidae. This small petrel is the only member of the genus Halobaena, but is closely allied to the prions. It is distributed across the Southern Ocean but breeds at a few island sites, all close to the Antarctic Convergence zone.
The Chilean tinamou is a type of tinamou commonly found in high elevation shrubland in subtropical regions of central Chile.
The South Island giant moa is an extinct species of moa in the genus Dinornis, known in Māori by the name moa nunui. It was one of the tallest-known bird species to walk the Earth, exceeded in weight only by the heavier but shorter elephant bird of Madagascar.
S. pedibus tridactylís, vertice palearibusque nudis