Chibanian

Last updated
Chibanian
0.774 – 0.129 Ma
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Chronology
Etymology
Name formalityFormal
Name ratifiedJanuary 2020
Synonym(s)Middle Pleistocene
Ionian
Usage information
Celestial body Earth
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unit Age
Stratigraphic unit Stage
Time span formalityFormal
Lower boundary definition1.1 m below the directional midpoint of the Brunhes-Matuyama magnetic reversal
Lower boundary GSSP Chiba, Japan
35°17′39″N140°08′47″E / 35.2943°N 140.1465°E / 35.2943; 140.1465
Lower GSSP ratifiedJanuary 2020 [3]
Upper boundary definitionNot formally defined
Upper boundary definition candidatesMarine Isotope Substage 5e
Upper boundary GSSP candidate section(s)None

The Chibanian, widely known as the Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. [4] The Chibanian name was officially ratified in January 2020. It is currently estimated to span the time between 0.770 Ma (770,000 years ago) and 0.126 Ma (126,000 years ago), also expressed as 770–126 ka. It includes the transition in palaeoanthropology from the Lower to the Middle Paleolithic over 300 ka.

Contents

The Chibanian is preceded by the Calabrian and succeeded by the proposed Tarantian. [5] The beginning of the Chibanian is the Brunhes–Matuyama reversal, when the Earth's magnetic field last underwent reversal. [6] Its end roughly coincides with the termination of the Penultimate Glacial Period and the onset of the Last Interglacial period (corresponding to the beginning of Marine Isotope Stage 5). [7]

The term Middle Pleistocene was in use as a provisional or "quasi-formal" designation by the International Union of Geological Sciences (IUGS). While the three lowest ages of the Pleistocene, the Gelasian, Calabrian and Chibanian have been officially defined, the Late Pleistocene has yet to be formally defined. [8]

Definition process

The International Union of Geological Sciences (IUGS) had previously proposed replacement of the Middle Pleistocene by an Ionian Age based on strata found in Italy. In November 2017, however, the Chibanian (based on strata at a site in Chiba Prefecture, Japan) replaced the Ionian as the Subcommission on Quaternary Stratigraphy's preferred GSSP proposal for the age that should replace the Middle Pleistocene sub-epoch. [9] The "Chibanian" name was ratified by the IUGS in January 2020. [4]

Climate

By early Middle Pleistocene, the Mid-Pleistocene Transition had changed the glacial cycles from an average 41,000 year periodicity present during most of the Early Pleistocene to a 100,000 year periodicity, [10] with the glacial cycles becoming asymmetric, having long glacial periods punctuated by short warm interglacial periods. [11] Millennial-scale climatic variability continued to be highly sensitive to precession and obliquity cycles. [12]

Events

The Early-Middle Pleistocene boundary saw the migration of true horses out of North America and into Eurasia. [13] Also around this time, the European mammoth species Mammuthus meridionalis became extinct and was replaced by the Asian species Mammuthus trogontherii (the steppe mammoth). This was coincident with the migration of the elephant genus Palaeoloxodon out of Africa and into Eurasia, including the first appearance of species like the European straight-tusked elephant (Palaeoloxodon antiquus). [14] With the extinction of Sinomastodon in East Asia at the Early-Middle Pleistocene boundary, gomphotheres became completely extinct in Afro-Eurasia, [15] [16] but continued to persist in the Americas into the Late Pleistocene. [16] There was a major extinction of carnivorous mammals in Europe around the Early-Middle Pleistocene transition, including the giant hyena Pachycrocuta. [17] The mid-late Middle Pleistocene saw the emergence of the woolly mammoth (Mammuthus primigenius), and its replacement of Mammuthus trogontherii, with the replacement of M. trogontherii in Europe by woolly mammoths being complete by around 200,000 years ago. [14] [18] The last member of the notoungulate family Mesotheriidae, Mesotherium , has its last records around 220,000 years ago, leaving Toxodontidae as the sole family of notoungulates to persist into the Late Pleistocene. [19] During the late Middle Pleistocene, around 195,000-135,000 years ago, the steppe bison (the ancestor of the modern American bison) migrated across the Bering land bridge into North America, marking the beginning of the Rancholabrean faunal stage. [20] Around 500,000 years ago, the last members of the largely European aquatic frog genus Palaeobatrachus and by extension the family Palaeobatrachidae became extinct. [21]

Palaeoanthropology

The Chibanian includes the transition in palaeoanthropology from the Lower to the Middle Paleolithic: i.e., the emergence of Homo sapiens sapiens between 300 ka and 400 ka. [22] The oldest known human DNA dates to the Middle Pleistocene, around 430,000 years ago. This is the oldest found, as of 2016. [23]

After analyzing 2,496 remains of Castor fiber (Eurasian beaver) and Trogontherium cuvieri found at Bilzingsleben in Germany, a team of scientists concluded that, around 400 ka, hominids in the area hunted and exploited beavers. They may have been targeted for their meat (based on cut marks on the bones) and skin. [24]

Chronology

Agepaleoclimateglaciationpalaeoanthropology
790–761 kaMIS 19 Günz (Elbe) glaciation Peking Man ( Homo erectus )
761–712 kaMIS 18
712–676 kaMIS 17
676–621 kaMIS 16
621–563 kaMIS 15 Gunz-Haslach interglacial Heidelberg Man ( Homo heidelbergensis ), Bodo cranium
563–524 kaMIS 14
524–474 ka MIS 13 end of Cromerian (Günz-Mindel) interglacial Boxgrove Man ( Homo heidelbergensis )
474–424 kaMIS 12 Anglian Stage in Britain; Haslach glaciation Tautavel Man ( Homo erectus )
424–374 ka MIS 11 Hoxnian (Britain), Yarmouthian (North America) Swanscombe Man ( Homo heidelbergensis )
374–337 kaMIS 10 Mindel glaciation, Elster glaciation, Riss glaciation
337–300 ka MIS 9 Purfleet Interglacial in Britain Mousterian
300–243 kaMIS 8 Irhoud 1 ( Homo sapiens ); Middle Paleolithic; Haplogroup A (Y-DNA)
243–191 kaMIS 7Aveley Interglacial in Britain Galilee Man; Haua Fteah
191–130 kaMIS 6 Illinoian Stage Herto Man ( Homo sapiens ); Macro-haplogroup L (mtDNA); Mousterian
130–123 ka MIS 5e peak of Eemian interglacial sub-stage, or Ipswichian in Britain Klasies River Caves; Sangoan

See also

Related Research Articles

<span class="mw-page-title-main">Mammoth</span> Extinct genus of mammals

A mammoth is any species of the extinct elephantid genus Mammuthus. The various species of mammoth were commonly equipped with long, curved tusks. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are more closely related to living Asian elephants than African elephants.

<span class="mw-page-title-main">Neogene</span> Second geologic period in the Cenozoic Era 23–2.6 million years ago

The Neogene is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period 23.03 million years ago (Mya) to the beginning of the present Quaternary Period 2.58 million years ago. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). The earlier term Tertiary Period was used to define the span of time now covered by Paleogene and Neogene and, despite no longer being recognized as a formal stratigraphic term, "Tertiary" still sometimes remains in informal use.

<span class="mw-page-title-main">Pleistocene</span> First epoch of the Quaternary Period

The Pleistocene is the geological epoch that lasted from c. 2.58 million to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek πλεῖστος (pleîstos), meaning "most", and καινός, meaning "new".

<span class="mw-page-title-main">Quaternary</span> Third and current period of the Cenozoic Era, from 2.58 million years ago to the present

The Quaternary is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS). It follows the Neogene Period and spans from 2.58 million years ago to the present. As of 2023, the Quaternary Period is divided into two epochs: the Pleistocene and the Holocene ; a third epoch, the Anthropocene, has recently been proposed, but it is not officially recognised by the ICS.

<span class="mw-page-title-main">Elephantidae</span> Family of mammals

Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.

<span class="mw-page-title-main">Timeline of glaciation</span> Chronology of the major ice ages of the Earth

There have been five or six major ice ages in the history of Earth over the past 3 billion years. The Late Cenozoic Ice Age began 34 million years ago, its latest phase being the Quaternary glaciation, in progress since 2.58 million years ago.

<span class="mw-page-title-main">Gomphothere</span> Extinct family of proboscidean mammals

Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to western South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.

The Cromerian Stage or Cromerian Complex, also called the Cromerian, is a stage in the Pleistocene glacial history of north-western Europe, mostly occurring more than half a million years ago. It is named after the East Anglian town of Cromer in Great Britain where interglacial deposits that accumulated during part of this stage were first discovered. The stratotype for this interglacial is the Cromer Forest Bed situated at the bottom of the coastal cliff near West Runton. The Cromerian stage preceded the Anglian and Elsterian glacials and show an absence of glacial deposits in western Europe, which led to the historical terms Cromerian interglacial and the Cromerian warm period. It is now known that the Cromerian consisted of multiple glacial and interglacial periods.

The Hoxnian Stage was a middle Pleistocene stage of the geological history of the British Isles. It was an interglacial which preceded the Wolstonian Stage and followed the Anglian Stage. It is equivalent to Marine Isotope Stage 11. Marine Isotope Stage 11 started 424,000 years ago and ended 374,000 years ago. The Hoxnian is divided into sub-stages Ho I to Ho IV.

The Wolstonian Stage is a middle Pleistocene stage of the geological history of Earth from approximately 374,000 until 130,000 years ago. It precedes the Eemian Stage in Europe and follows the Hoxnian Stage in the British Isles.

The Illinoian Stage is the name used by Quaternary geologists in North America to designate the period c.191,000 to c.130,000 years ago, during the Chibanian stage of the Pleistocene, when sediments comprising the Illinoian Glacial Lobe were deposited. It precedes the Sangamonian Stage and follows the Pre-Illinoian Stage in North America. The Illinoian Stage is defined as the period of geologic time during which the glacial tills and outwash, which comprise the bulk of the Glasford Formation, accumulated to create the Illinoian Glacial Lobe. It occurs at about the same time as the penultimate glacial period.

<span class="mw-page-title-main">Mammoth steppe</span> Prehistoric biome

During the Last Glacial Maximum, the mammoth steppe, also known as steppe-tundra, was once the Earth's most extensive biome. It stretched east-to-west, from the Iberian Peninsula in the west of Europe, across Eurasia to North America, through Beringia and Canada; from north-to-south, the steppe reached from the arctic islands southward to China. The mammoth steppe was cold and dry, and relatively featureless, though topography and geography varied considerably throughout. Some areas featured rivers which, through erosion, naturally created gorges, gulleys, or small glens. The continual glacial recession and advancement over millennia contributed more to the formation of larger valleys and different geographical features. Overall, however, the steppe is known to be flat and expansive grassland. The vegetation was dominated by palatable, high-productivity grasses, herbs and willow shrubs.

<span class="mw-page-title-main">Late Pleistocene</span> Third division (unofficial) of the Pleistocene Epoch

The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as the upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently defined as the time between c. 129,000 and c. 11,700 years ago. The late Pleistocene equates to the proposed Tarantian Age of the geologic time scale, preceded by the officially ratified Chibanian. The beginning of the Late Pleistocene is the transition between the end of the Penultimate Glacial Period and the beginning of the Last Interglacial around 130,000 years ago. The Late Pleistocene ends with the termination of the Younger Dryas, some 11,700 years ago when the Holocene Epoch began.

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<span class="mw-page-title-main">Straight-tusked elephant</span> Extinct species of elephant

The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. It was larger than any living elephant, with some adult males suggested to reach 3.81–4.2 metres (12.5–13.8 ft) in shoulder height, and 11.3–15 tonnes in weight. Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.

<span class="mw-page-title-main">Steppe mammoth</span> Extinct species of mammal

Mammuthus trogontherii, sometimes called the steppe mammoth, is an extinct species of mammoth that ranged over most of northern Eurasia during the Early and Middle Pleistocene, approximately 1.7 million-200,000 years ago. One of the largest mammoth species, it evolved in East Asia during the Early Pleistocene, around 1.8 million years ago, before migrating into North America around 1.5 million years ago, and into Europe during the Early/Middle Pleistocene transition, around 1 to 0.7 million years ago. It was the ancestor of the woolly mammoth and Columbian mammoth of the later Pleistocene.

<span class="mw-page-title-main">Woolly mammoth</span> Extinct species of mammoth from the Quaternary period

The woolly mammoth is an extinct species of mammoth that lived from the Middle Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with the African Mammuthus subplanifrons in the early Pliocene. The woolly mammoth began to diverge from the steppe mammoth about 800,000 years ago in Siberia. Its closest extant relative is the Asian elephant. The Columbian mammoth lived alongside the woolly mammoth in North America, and DNA studies show that the two hybridised with each other.

The Pre-Illinoian Stage is used by Quaternary geologists for the early and middle Pleistocene glacial and interglacial periods of geologic time in North America from ~2.5–0.2 Ma.

The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time between 2.580 ± 0.005 Ma and 0.773 ± 0.005 Ma. The term Early Pleistocene applies to both the Gelasian Age and the Calabrian Age.

<span class="mw-page-title-main">Mid-Pleistocene Transition</span> Change in glacial cycles c. 1m years ago

The Mid-Pleistocene Transition (MPT), also known as the Mid-Pleistocene Revolution (MPR), is a fundamental change in the behaviour of glacial cycles during the Quaternary glaciations. The transition occurred gradually, taking place approximately 1.25–0.7 million years ago, in the Pleistocene epoch. Before the MPT, the glacial cycles were dominated by a 41,000-year periodicity with low-amplitude, thin ice sheets, and a linear relationship to the Milankovitch forcing from axial tilt. Because of this, sheets were more dynamic during the Early Pleistocene. After the MPT there have been strongly asymmetric cycles with long-duration cooling of the climate and build-up of thick ice sheets, followed by a fast change from extreme glacial conditions to a warm interglacial. This led to less dynamic ice sheets. Interglacials before the MPT had lower levels of atmospheric carbon dioxide compared to interglacials after the MPT. One of the MPT's effects was causing ice sheets to become higher in altitude and less slippery compared to before. The cycle lengths have varied, with an average length of approximately 100,000 years.

References

  1. ↑ Cohen, K. M.; Finney, S. C.; Gibbard, P. L.; Fan, J.-X. (January 2020). "International Chronostratigraphic Chart" (PDF). International Commission on Stratigraphy. Retrieved 23 February 2020.
  2. ↑ Mike Walker; et al. (December 2018). "Formal ratification of the subdivision of the Holocene Series/Epoch (Quaternary System/Period)" (PDF). Episodes. Subcommission on Quaternary Stratigraphy (SQS). 41 (4): 213–223. doi:10.18814/epiiugs/2018/018016 . Retrieved 11 November 2019.
  3. ↑ "Global Boundary Stratotype Section and Point". International Commission of Stratigraphy. Retrieved 26 December 2020.
  4. 1 2 Hornyak, Tim (30 January 2020). "Japan Puts Its Mark on Geologic Time with the Chibanian Age". Eos – Earth & Space Science News. American Geophysical Union . Retrieved 31 January 2020.
  5. ↑ Cohen, K. M.; Finney, S. C.; Gibbard, P. L.; Fan, J.-X. (January 2020). "International Chronostratigraphic Chart" (PDF). International Commission on Stratigraphy. Retrieved 23 February 2020.
  6. ↑ Gradstein, Felix M.; Ogg, James G.; Smith, Alan G., eds. (2004). A Geological Time Scale 2004 (3rd ed.). Cambridge: Cambridge University Press. p.  28. ISBN   9780521786737.
  7. ↑ D. Dahl-Jensen & others (2013). "Eemian interglacial reconstructed from a Greenland folded ice core" (PDF). Nature. 493 (7433): 489–494. Bibcode:2013Natur.493..489N. doi:10.1038/nature11789. PMID   23344358. S2CID   4420908.
  8. ↑ P. L. Gibbard (17 April 2015). "The Quaternary System/Period and its major sub-divisions". Russian Geology and Geophysics. Special Issue: Topical Problems of Stratigraphy and Evolution of the Biosphere. Elsevier BV. 56 (4): 686–688. Bibcode:2015RuGG...56..686G. doi:10.1016/j.rgg.2015.03.015 . Retrieved 13 November 2019.
  9. ↑ "Japan-based name 'Chibanian' set to represent geologic age of last magnetic shift". The Japan Times. 14 November 2017. Retrieved 13 November 2019.
  10. ↑ Berends, C. J.; Köhler, P.; Lourens, L. J.; van de Wal, R. S. W. (June 2021). "On the Cause of the Mid-Pleistocene Transition". Reviews of Geophysics. 59 (2). Bibcode:2021RvGeo..5900727B. doi:10.1029/2020RG000727. hdl: 1874/412413 . ISSN   8755-1209. S2CID   236386405.
  11. ↑ Chalk, Thomas B.; Hain, Mathis P.; Foster, Gavin L.; Rohling, Eelco J.; Sexton, Philip F.; Badger, Marcus P. S.; Cherry, Soraya G.; Hasenfratz, Adam P.; Haug, Gerald H.; Jaccard, Samuel L.; Martínez-García, Alfredo; Pälike, Heiko; Pancost, Richard D.; Wilson, Paul A. (2017-12-12). "Causes of ice age intensification across the Mid-Pleistocene Transition". Proceedings of the National Academy of Sciences. 114 (50): 13114–13119. Bibcode:2017PNAS..11413114C. doi: 10.1073/pnas.1702143114 . ISSN   0027-8424. PMC   5740680 . PMID   29180424.
  12. ↑ Sun, Youbin; McManus, Jerry F.; Clemens, Steven C.; Zhang, Xu; Vogel, Hendrik; Hodell, David A.; Guo, Fei; Wang, Ting; Liu, Xingxing; An, Zhisheng (1 November 2021). "Persistent orbital influence on millennial climate variability through the Pleistocene". Nature Geoscience . 14 (11): 812–818. Bibcode:2021NatGe..14..812S. doi:10.1038/s41561-021-00794-1. ISSN   1752-0908. S2CID   240358493 . Retrieved 26 February 2024.
  13. ↑ Vershinina, Alisa O.; Heintzman, Peter D.; Froese, Duane G.; Zazula, Grant; Cassatt-Johnstone, Molly; Dalén, Love; Der Sarkissian, Clio; Dunn, Shelby G.; Ermini, Luca; Gamba, Cristina; Groves, Pamela; Kapp, Joshua D.; Mann, Daniel H.; Seguin-Orlando, Andaine; Southon, John (December 2021). "Ancient horse genomes reveal the timing and extent of dispersals across the Bering Land Bridge". Molecular Ecology. 30 (23): 6144–6161. Bibcode:2021MolEc..30.6144V. doi:10.1111/mec.15977. hdl: 10995/118212 . ISSN   0962-1083. PMID   33971056.
  14. 1 2 Lister, Adrian M. (2004), "Ecological Interactions of Elephantids in Pleistocene Eurasia", Human Paleoecology in the Levantine Corridor, Oxbow Books, pp. 53–60, ISBN   978-1-78570-965-4 , retrieved 2020-04-14
  15. ↑ Wang, Yuan; Jin, Chang-zhu; Mead, Jim I. (August 2014). "New remains of Sinomastodon yangziensis (Proboscidea, Gomphotheriidae) from Sanhe karst Cave, with discussion on the evolution of Pleistocene Sinomastodon in South China". Quaternary International. 339–340: 90–96. Bibcode:2014QuInt.339...90W. doi:10.1016/j.quaint.2013.03.006.
  16. 1 2 Cantalapiedra, Juan L.; Sanisdro, Oscar L.; Zhang, Hanwen; Alberdi, Mª Teresa; Prado, Jose Luis; Blanco, Fernando; Saarinen, Juha (1 July 2021). "The rise and fall of proboscidean ecological diversity". Nature Ecology & Evolution. 355 (9): 1266–1272. Bibcode:2021NatEE...5.1266C. doi:10.1038/s41559-021-01498-w. PMID   34211141. S2CID   235712060 . Retrieved 21 August 2021– via Escience.magazine.org.
  17. ↑ Palombo, Maria Rita; Sardella, Raffaele; Novelli, Micaela (March 2008). "Carnivora dispersal in Western Mediterranean during the last 2.6Ma". Quaternary International. 179 (1): 176–189. Bibcode:2008QuInt.179..176P. doi:10.1016/j.quaint.2007.08.029.
  18. ↑ Lister, Adrian M. (October 2022). "Mammoth evolution in the late Middle Pleistocene: The Mammuthus trogontherii-primigenius transition in Europe". Quaternary Science Reviews. 294: 107693. Bibcode:2022QSRv..29407693L. doi:10.1016/j.quascirev.2022.107693. S2CID   252264887.
  19. ↑ Fernández-Monescillo, Marcos; Martínez, Gastón; García López, Daniel; Frechen, Manfred; Romero-Lebrón, Eugenia; Krapovickas, Jerónimo M.; Haro, J. Augusto; Rodríguez, Pablo E.; Rouzaut, Sabrina; Tauber, Adan A. (February 2023). "The last record of the last typotherid (Notoungulata, Mesotheriidae, Mesotherium cristatum) for the middle Pleistocene of the western Pampean region, Córdoba Province, Argentina, and its biostratigraphic implications". Quaternary Science Reviews. 301: 107925. Bibcode:2023QSRv..30107925F. doi:10.1016/j.quascirev.2022.107925. S2CID   254913691.
  20. ↑ Froese, Duane; Stiller, Mathias; Heintzman, Peter D.; Reyes, Alberto V.; Zazula, Grant D.; Soares, André E. R.; Meyer, Matthias; Hall, Elizabeth; Jensen, Britta J. L.; Arnold, Lee J.; MacPhee, Ross D. E. (2017-03-28). "Fossil and genomic evidence constrains the timing of bison arrival in North America". Proceedings of the National Academy of Sciences. 114 (13): 3457–3462. Bibcode:2017PNAS..114.3457F. doi: 10.1073/pnas.1620754114 . ISSN   0027-8424. PMC   5380047 . PMID   28289222.
  21. ↑ Wuttke, Michael; Přikryl, Tomáš; Ratnikov, Viacheslav Yu.; Dvořák, Zdeněk; Roček, Zbyněk (September 2012). "Generic diversity and distributional dynamics of the Palaeobatrachidae (Amphibia: Anura)". Palaeobiodiversity and Palaeoenvironments. 92 (3): 367–395. Bibcode:2012PdPe...92..367W. doi:10.1007/s12549-012-0071-y. ISSN   1867-1594. S2CID   130080167.
  22. ↑ D. Richter & others (8 June 2017). "The Age of Hominin Fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age". Nature. 546 (7657): 293–296. Bibcode:2017Natur.546..293R. doi:10.1038/nature22335. PMID   28593967. S2CID   205255853..
  23. ↑ Crew, Bec (15 March 2016). "The Oldest Human Genome Ever Has Been Sequenced, And It Could Rewrite Our History". ScienceAlert. Retrieved 5 June 2019.
  24. ↑ Gaudzinski-Windheuser, Sabine; Kindler, Lutz; Roebroeks, Wil (2023-11-13). "Beaver exploitation, 400,000 years ago, testifies to prey choice diversity of Middle Pleistocene hominins". Scientific Reports. 13 (1): 19766. Bibcode:2023NatSR..1319766G. doi: 10.1038/s41598-023-46956-6 . hdl: 1887/3674398 . ISSN   2045-2322. PMID   37957223.
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