Eysyslopterus | |
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Restoration of the carapace of E. patteni based on Tetlie and Poschmann's of 2008 | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Order: | † Eurypterida |
Superfamily: | † Adelophthalmoidea |
Family: | † Adelophthalmidae |
Genus: | † Eysyslopterus Tetlie & Poschmann, 2008 |
Type species | |
†Eysyslopterus patteni Størmer, 1934 | |
Synonyms | |
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Eysyslopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Eysyslopterus is classified as part of the family Adelophthalmidae, the only clade within the derived ("advanced") Adelophthalmoidea superfamily of eurypterids. One fossil of the single and type species, E. patteni, has been discovered in deposits of the Late Silurian period (Ludlow epoch) in Saaremaa, Estonia. The genus is named after Eysysla, the Viking name for Saaremaa, and opterus, a traditional suffix for the eurypterid genera, meaning "wing". The species name honors William Patten, an American biologist and zoologist who discovered the only known fossil of Eysyslopterus.
Eysyslopterus is a little-known basal genus that was distinguished from the rest of adelophthalmids by the position near the head margin of the eyes, different from the rest of its relatives. Its carapace was parabolic (approximately U-shaped) and with transverse deep furrows forming the ornamentation. With an estimated length of 8 cm (3.1 in), Eysyslopterus was a small eurypterid. It lived in a nearshore lagoonal quiet community along other eurypterid species.
Like the other adelophthalmid eurypterids, Eysyslopterus was a small eurypterid. The total size of the only known specimen is estimated at only 8 centimetres (3.1 inches), [1] far from the largest adelophthalmids like Adelophthalmus khakassicus of 32 cm (12.6 in) in length. [2]
Eysyslopterus is a little known eurypterid, with only one specimen collected that only preserves the prosoma ("head") with no appendages (limbs). The prosoma was rather broad, with a parabolic (approximately U-shaped) outline. It was 19.5 millimetres (0.77 in) long and 22 mm (0.87 in) wide. Although the anterior part of the prosoma is not preserved, the fossil mold indicates a pointed portion. The posterior border was slightly convex. A narrow rim surrounded the lateral margins of the carapace (dorsal shield of the prosoma). The prosoma surface was gently arched, with the central portion rather level. The eyes were reniform (bean-shaped) and with a thin and light-colored test (external "shield") on its surface. They measured 4.5 mm (0.18 in) in length. The ocelli (simple eyes) were located in the posterior half of the prosoma and consisted of two circular spots. The posterior part was smooth, but the previous part had a distinctive ornamentation. This ornamentation extended until the end of the eyes and was composed of transverse lines developed as deep furrows. [3]
Eysyslopterus was a basal ("primitive") genus with respect to the rest of adelophthalmids, with the eyes closer to the margin than to the ocelli, suggesting that the eyes migrated towards a central position from the basal genera to Adelophthalmus. [4]
Eysyslopterus is only known from one single specimen (and is therefore the holotype, AMNH 32720, housed at the American Museum of Natural History) which preserves the carapace. [4] This fossil was collected by the American biologist and zoologist William Patten during his extensive explorations at the Rootsikula Formation in Saaremaa, Estonia (then part of the Soviet Union). During a meeting between Patten and the Norwegian paleontologist and geologist Leif Størmer in 1932, Størmer observed Patten's private collection of eurypterids, among which he recognized a carapace of a new unknown species. He described it in 1934 as belonging to the genus Hughmilleria patteni, named after Patten, who died months after the meeting. Størmer noticed several differences between his new species and other Hughmilleria species, essentially the shape of the prosoma and the form or position of the eyes. He also suggested a close relationship between H. patteni and other species, especially the Scottish species H. lanceolata. [3]
However, in 1961, the American paleontologist Erik Norman Kjellesvig-Waering split Hughmilleria in two subgenera, H. (Hughmilleria) for species with large ovoid marginal eyes and H. ( Nanahughmilleria ) for small reniform intramarginal (within the margin) eyes. Since H. patteni fulfilled this condition, it was referred to H. (Nanahughmilleria). [5] It was not until 2008 when during a phylogenetic review of the entire Adelophthalmidae family by the Norwegian Odd Erik Tetlie and German Markus Poschmann paleontologists, N. patteni was recognized as sufficiently different to represent its own genus due to the anterior position of its eyes, much more exaggerated than in the other adelophthalmid genera. The first part of the generic name is composed of the word Eysysla, the Viking name for Saaremaa, and the second part by opterus, a suffix commonly used in eurypterids which means "wing" (and therefore, translated as "wing of Eysysla"). [4]
Eysyslopterus is classified as part of the family Adelophthalmidae, the only clade ("group") within the superfamily Adelophthalmoidea. [6] E. patteni was originally described as a species of the genus Hughmilleria, [3] but it was considered different enough to represent a new separate genus in 2008. [4]
Eysyslopterus is considered to be the most basal adelophthalmid genus due to the position of its eyes. These are placed closer to the margin than to the ocelli, in contrast to any other adelopthalmid. In fact, the carapace of Eysyslopterus and other basal genera belonging to superfamilies, Orcanopterus and Herefordopterus , were almost identical. In the waeringopteroid Orcanopterus, the eyes were separated from the margin by the marginal rim, whereas in Herefordopterus they were completely marginal, a feature present in all pterygotioid genera. [4] In addition, the triangular anterior carapace margin present in Eysyslopterus was shared with Hughmilleria, indicating that it might be a plesiomorphic trait (a trait present in a common ancestor). [7] These characteristics have led some authors to question whether Eysyslopterus really represents an adelophthalmid or the sister taxon of a group formed by Pterygotioidea and Adelophthalmoidea, but until more fossil material is found this can not be proven. [4]
The cladogram below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the Diploperculata infraorder of eurypterids (Adelophthalmoidea, Pterygotioidea and the waeringopteroids), as inferred by Tetlie and Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis. [4]
Diploperculata |
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The single known specimen of Eysyslopterus has been recovered from Silurian deposits of the Ludlow epoch of the Rootsikula Formation of Saaremaa, Estonia. [4] This formation was the habitat of a fauna of eurypterids, among them Erettopterus laticauda, E. osiliensis, Eurypterus tetragonophthalmus , Mixopterus simonsi and Strobilopterus laticeps . Fossil remains of indeterminate osteostracid and thelodontid fishes have also been found. The habitat in which Eysyslopterus lived is considered to be a nearshore lagoonal quiet community. The lithology (the physical characteristics of the rocks) of the zone was composed of mudcrack and burrows of dolomite and limestone. [8]
Eurypterus is an extinct genus of eurypterid, a group of organisms commonly called "sea scorpions". The genus lived during the Silurian period, from around 432 to 418 million years ago. Eurypterus is by far the most well-studied and well-known eurypterid and its fossil specimens probably represent more than 90% of all known eurypterid specimens.
Stylonurina is one of two suborders of eurypterids, a group of extinct arthropods commonly known as "sea scorpions". Members of the suborder are collectively and informally known as "stylonurine eurypterids" or "stylonurines". They are known from deposits primarily in Europe and North America, but also in Siberia.
Slimonia is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Slimonia have been discovered in deposits of Silurian age in South America and Europe. Classified as part of the family Slimonidae alongside the related Salteropterus, the genus contains three valid species, S. acuminata from Lesmahagow, Scotland, S. boliviana from Cochabamba, Bolivia and S. dubia from the Pentland Hills of Scotland and one dubious species, S. stylops, from Herefordshire, England. The generic name is derived from and honors Robert Slimon, a fossil collector and surgeon from Lesmahagow.
Carcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Carcinosoma are restricted to deposits of late Silurian age. Classified as part of the family Carcinosomatidae, which the genus lends its name to, Carcinosoma contains seven species from North America and Great Britain.
Hughmilleria is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Hughmilleria have been discovered in deposits of the Silurian age in China and the United States. Classified as part of the basal family Hughmilleriidae, the genus contains three species, H. shawangunk from the eastern United States, H. socialis from Pittsford, New York, and H. wangi from Hunan, China. The genus is named in honor of the Scottish geologist Hugh Miller.
Alkenopterus is a genus of prehistoric eurypterid classified as part of the family Onychopterellidae. The genus contains two species, A. brevitelson and A. burglahrensis, both from the Devonian of Germany.
Bassipterus is a genus of eurypterid, an extinct group of aquatic arthropods. Bassipterus is classified as part of the family Adelophthalmidae, the only clade within the derived ("advanced") Adelophthalmoidea superfamily of eurypterids. Fossils of the single and type species, B. virgnicus, have been discovered in deposits of the Late Silurian age in West Virginia and Maryland, United States. The genus is named after Bass, where most of the fossils have been recovered.
Nanahughmilleria is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Nanahughmilleria have been discovered in deposits of Devonian and Silurian age in the United States, Norway, Russia, England and Scotland, and have been referred to several different species.
Pittsfordipterus is a genus of eurypterid, an extinct group of aquatic arthropods. Pittsfordipterus is classified as part of the family Adelophthalmidae, the only clade in the derived ("advanced") Adelophthalmoidea superfamily of eurypterids. Fossils of the single and type species, P. phelpsae, have been discovered in deposits of Silurian age in Pittsford, New York state. The genus is named after Pittsford, where the two only known specimens have been found.
Salteropterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Salteropterus have been discovered in deposits of Late Silurian age in Britain. Classified as part of the family Slimonidae, the genus contains one known valid species, S. abbreviatus, which is known from fossils discovered in Herefordshire, England, and a dubious species, S. longilabium, with fossils discovered in Leintwardine, also in Herefordshire. The generic name honours John William Salter, who originally described S. abbreviatus as a species of Eurypterus in 1859.
Erettopterus is a genus of large predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Erettopterus have been discovered in deposits ranging from Early Silurian to the Early Devonian, and have been referred to several different species. Fossils have been recovered from two continents; Europe and North America. The genus name is composed by the Ancient Greek words ἐρέττω (eréttō), which means "rower", and πτερόν (pterón), which means "wing", and therefore, "rower wing".
Parahughmilleria is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Parahughmilleria have been discovered in deposits of the Devonian and Silurian age in the United States, Canada, Russia, Germany, Luxembourg and Great Britain, and have been referred to several different species. The first fossils of Parahughmilleria, discovered in the Shawangunk Mountains in 1907, were initially assigned to Eurypterus. It would not be until 54 years later when Parahughmilleria would be described.
Adelophthalmus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Adelophthalmus have been discovered in deposits ranging in age from the Early Devonian to the Early Permian, which makes it the longest lived of all known eurypterid genera, with a total temporal range of over 120 million years. Adelopthtalmus was the final genus of the Eurypterina suborder of eurypterids and consisted the only known genus of swimming eurypterids from the Middle Devonian until its extinction during the Permian, after which the few surviving eurypterids were all walking forms of the suborder Stylonurina.
Unionopterus is a genus of eurypterid, an extinct group of aquatic arthropods commonly known as "sea scorpions". Fossils have been registered from the Early Carboniferous period. The genus contains only one species, U. anastasiae, recovered from deposits of Tournaisian to Viséan stages in Kazakhstan. Known from one single specimen which was described in a publication of Russian language with poor illustrations, Unionopterus' affinities are extremely poorly known.
Stylonuridae is a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is one of two families contained in the superfamily Stylonuroidea, which in turn is one of four superfamilies classified as part of the suborder Stylonurina.
Pterygotioidea is a superfamily of eurypterids, an extinct group of aquatic arthropods. Pterygotioids were the most derived members of the infraorder Diploperculata and the sister group of the adelophthalmoid eurypterids. The group includes the basal and small hughmilleriids, the larger and specialized slimonids and the famous pterygotids which were equipped with robust and powerful cheliceral claws.
Adelophthalmidae is a family of eurypterids, an extinct group of aquatic arthropods. Adelophthalmidae is the only family classified as part of the superfamily Adelophthalmoidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina.
Hughmilleriidae is a family of eurypterids, an extinct group of aquatic arthropods. The hughmilleriids were the most basal members of the superfamily Pterygotioidea, in contrast with the more derived families Pterygotidae and Slimonidae. Despite their classification as pterygotioids, the hughmilleriids possessed several characteristics shared with other eurypterid groups, such as the lanceolate telson.
Borchgrevinkium is an extinct genus of chelicerate arthropod. A fossil of the single and type species, B. taimyrensis, has been discovered in deposits of the Early Devonian period in the Krasnoyarsk Krai, Siberia, Russia. The name of the genus honors Carsten Borchgrevink, an Anglo-Norwegian explorer who participated in many expeditions to Antarctica. Borchgrevinkium represents a poorly known genus whose affinities are uncertain.
This timeline of eurypterid research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of eurypterids, a group of extinct aquatic arthropods closely related to modern arachnids and horseshoe crabs that lived during the Paleozoic Era.