Lake Malawi

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Lake Malawi (Lake Nyasa)
Lake Malawi seen from orbit.jpg
View from orbit. North is in upper right corner
Malawi relief location map.jpg
Red pog.svg
Lake Malawi (Lake Nyasa)
Africa relief location map.jpg
Red pog.svg
Lake Malawi (Lake Nyasa)
Coordinates 12°00′S34°30′E / 12.000°S 34.500°E / -12.000; 34.500 Coordinates: 12°00′S34°30′E / 12.000°S 34.500°E / -12.000; 34.500
Lake type Ancient lake, Rift lake
Primary inflows Ruhuhu River [1]
Primary outflows Shire River [1]
Basin  countries Malawi, Mozambique, and Tanzania
Max. length560 km (350 mi) [1] to 580 [2]
Max. width75 km (47 mi) [1]
Surface area 29,600 km2 (11,400 sq mi) [1]
Average depth292 m (958 ft) [3]
Max. depth706 m (2,316 ft) [3]
Water volume 8,400 km3 (2,000 cu mi) [3]
Surface elevation468 metres (1,535 ft) above sea level [4]
Islands Likoma and Chizumulu islets, Mumbo Island
References [1] [3]
Official nameLake Niassa and its Coastal Zone
Designated26 April 2011
Reference no.1964 [5]

Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, is an African Great Lake and the southernmost lake in the East African Rift system, located between Malawi, Mozambique and Tanzania.

Contents

It is the fifth largest fresh water lake in the world by volume, the ninth largest lake in the world by area—and the third largest and second deepest lake in Africa. Lake Malawi is home to more species of fish than any other lake in the world, [6] including at least 700 species of cichlids. [7] The Mozambique portion of the lake was officially declared a reserve by the Government of Mozambique on June 10, 2011, [8] while in Malawi a portion of the lake is included in Lake Malawi National Park. [6]

Lake Malawi is a meromictic lake, meaning that its water layers do not mix. The permanent stratification of Lake Malawi's water and the oxic-anoxic boundary (relating to oxygen in the water) are maintained by moderately small chemical and thermal gradients. [9]

Geography

Lake Malawi is between 560 kilometres (350 mi) [1] and 580 kilometres (360 mi) long, [2] and about 75 kilometres (47 mi) wide at its widest point. The lake has a total surface area of about 29,600 square kilometres (11,400 sq mi). [1] The lake is 706 m (2,316 ft) at its deepest point, located in a major depression in the north-central part. [10] Another smaller depression in the far north reaches a depth of 528 m (1,732 ft). [10] The southern half of the lake is shallower; less than 400 m (1,300 ft) in the south-central part and less than 200 m (660 ft) in the far south. [10] The lake has shorelines on western Mozambique, eastern Malawi, and southern Tanzania. The largest river flowing into it is the Ruhuhu River, and there is an outlet at its southern end, the Shire River, a tributary that flows into the very large Zambezi River in Mozambique. [2] Evaporation accounts for more than 80% of the water loss from the lake, considerably more than the outflowing Shire River. [11] The outflows from Lake Malawi into the Shire River are vital for the economy as the water resources support hydropower, irrigation and downstream biodiversity. [12] Concerns have been raised over the future climate change impacts of Lake Malawi due to the recent decline in lake levels and the overall drying trend. [13] The climate in the lake region is already experiencing changes, with the temperatures predicted to increase throughout the country. [14]

The lake is about 350 kilometres (220 mi) southeast of Lake Tanganyika, another of the great lakes of the East African Rift. [ citation needed ]

The Lake Malawi National Park is located at the southern end of the lake. [15]

Geological history

Topographic profile of Lake Malawi's rift shoulder FlankMalawi.png
Topographic profile of Lake Malawi's rift shoulder
The East African Rift (red) with the Rift Valley lakes, Malawi being in the south Map of Great Rift Valley.svg
The East African Rift (red) with the Rift Valley lakes, Malawi being in the south

Malawi is one of the major Rift Valley lakes and an ancient lake. The lake lies in a valley formed by the opening of the East African Rift, where the African tectonic plate is being split into two pieces. This is called a divergent plate tectonics boundary. Malawi has typically been estimated to be 1–2 million years old (mya), [16] [17] but more recent evidence points to a considerably older lake with a basin that started to form about 8.6 mya and deep-water condition first appeared 4.5 mya. [18] [19]

The water levels have varied dramatically over time, ranging from almost 600 m (2,000 ft) below current level [20] to 10–20 m (33–66 ft) above. [18] During periods the lake dried out almost completely, leaving only one or two relatively small, highly alkaline and saline lakes in what currently are Malawi's deepest parts. [18] [20] A water chemistry resembling the current conditions only appeared about 60,000 years ago. [20] Major low-water periods are estimated to have occurred about 1.6 to 1.0–0.57 million years ago (where it might have dried out completely), 420,000 to 250,000–110,000 years ago, [18] about 25,000 years ago and 18,000–10,700 years ago. [19] During the peak of the low-water period between 1390 and 1860 AD, it may have been 120–150 m (390–490 ft) below current water levels. [17]

Water characteristics

The lake's water is alkaline (pH 7.7–8.6) and warm with a typical surface temperature between 24 and 29 °C (75–84 °F), while deep sections typically are about 22 °C (72 °F). [21] The thermocline is located at a depth of 40–100 m (130–330 ft). [11] The oxygen limit is at a depth of approximately 250 m (820 ft), effectively restricting fish and other aerobic organisms to the upper part. [22] The water is very clear for a lake and the visibility can be up to 20 m (66 ft), but slightly less than half this figure is more common and it is below 3 m (10 ft) in muddy bays. [10] However, during the rainy season months of January to March, the waters are more muddy due to muddy river inflows. [23]

European discovery and colonisation

The Portuguese trader Candido José da Costa Cardoso was the first European to visit the lake in 1846. [24] David Livingstone reached the lake in 1859, and named it Lake Nyasa. [2] He also referred to it by a pair of nicknames: Lake of Stars and Lake of Storms. [25] The Lake of Stars nickname came after Livingstone observed lights from the lanterns of the fishermen in Malawi on their boats, that resemble, from a distance, stars in the sky. [26] Later, after experiencing the unpredictable and extremely violent gales that sweep through the area, he also referred to it as the Lake of Storms. [26]

On 16 August 1914, Lake Malawi was the scene of a brief naval battle when the British gunboat SS Gwendolen, commanded by a Captain Rhoades, heard that World War I had broken out, and he received orders from the British Empire's high command to "sink, burn, or destroy" the German Empire's only gunboat on the lake, the Hermann von Wissmann , commanded by a Captain Berndt. Rhoades's crew found the Hermann von Wissmann in a bay near Sphinxhaven, in German East African territorial waters. Gwendolen disabled the German boat with a single cannon shot from a range of about 1,800 metres (2,000 yd). This very brief gunboat conflict was hailed by The Times in England as the British Empire's first naval victory of World War I. [27] [28]

Borders

Dashed line: current Malawi border
Dotted line: Tanzanian claim Malawi-Tanzania border dispute.png
Dashed line: current Malawi border
Dotted line: Tanzanian claim

Tanzania–Malawi dispute

The partition of the lake's surface area between Malawi and Tanzania is under dispute. Tanzania claims that the international border runs through the middle of the lake. [29] On the other hand, Malawi claims the whole of the surface of this lake that is not in Mozambique, including the waters that are next to the shoreline of Tanzania. [30] Both sides cite the Heligoland Treaty of 1890 between Great Britain and Germany concerning the border. The wrangle in this dispute occurred when the British colonial government, just after they had captured Tanganyika from Germany, placed all of the waters of the lake under a single jurisdiction, that of the territory of Nyasaland, without a separate administration for the Tanganyikan portion of the surface. Later in colonial times, two jurisdictions were established. [31]

The dispute came to a head in 1967 when Tanzania officially protested to Malawi; however nothing was settled. [32] Occasional flare-ups of conflict occurred during the 1990s and in the 21st century. [33] In 2012, Malawi's oil exploration initiative brought the issue to the fore, with Tanzania demanding that exploration cease until the dispute was settled. [34]

Malawi–Mozambique border

In 1954, an agreement was signed between the British and the Portuguese making the middle of the lake their boundary with the exception of Chizumulu Island and Likoma Island, which were kept by the British and are now part of Malawi. [31]

Transport

A jetty juts into the lake at Nkhata Bay Nkhata Bay, Malawi.jpg
A jetty juts into the lake at Nkhata Bay

MV Chauncy Maples began service on the lake in 1901 as the SS Chauncy Maples: a floating clinic and church for the Universities' Mission to Central Africa. She later served as a ferry and is currently being renovated into a mobile clinic at Monkey Bay. The renovation was expected to be complete during the first half of 2014, but was halted in 2017. [35] MV Mpasa entered service in 1935. [36] The ferry MV Ilala entered service in 1951. In recent years she has often been out of service, but when operational she runs between Monkey Bay at the southern end of the lake to Karonga on the northern end, and occasionally to the Iringa Region of Tanzania. The ferry MV Mtendere entered service in 1980. [36] By 1982 she was carrying 100,000 passengers each year., [36] but as of 2014 she was out of service. [37] She normally serves the southern part of the lake but if Ilala was out of service she operated the route to Karonga. The Tanzanian ferry MV Songea was built in 1988. [38] Her operator was the Tanzania Railway Corporation Marine Division until 1997, when it became the Marine Services Company Limited. [39] Songea plies weekly between Liuli and Nkhata Bay via Itungi and Mbamba Bay. [38] The worst Lake disaster was a ship accident by the MV Vipya in 1946 which resulted in 145 deaths. [40]

Wildlife

An African fish eagle catching a fish in Lake Malawi Haliaeetus vocifer -Cape Maclear, Malawi -fishing-8b.jpg
An African fish eagle catching a fish in Lake Malawi

Wildlife found in and around Lake Malawi or Nyasa includes Nile crocodiles, hippopotamus, monkeys, [41] and a significant population of African fish eagles that feed off fish from the lake. [42]

Fish

Fishing

Lake Malawi sardines (Engraulicypris sardella) spread out to dry on the shore of the lake Pesce ad essiccare sulla riva del lago malawi.JPG
Lake Malawi sardines (Engraulicypris sardella) spread out to dry on the shore of the lake

Lake Malawi has for millennia provided a major food source to the residents of its shores since its waters are rich in fish. Among the most popular are the four species of chambo, consisting of any one of four species in the subgenus Nyasalapia ( Oreochromis karongae , O. lidole , O. saka and O. squamipinnis ), as well as the closely related O. shiranus . [43] Other species that support important fisheries include the Lake Malawi sardine (Engraulicypris sardella) and the large kampango catfish (Bagrus meridionalis). [10] Most fishing provides food for the increasing human population near the lake, but some are exported from Malawi. The wild population of fish is increasingly threatened by overfishing and water pollution. [44] [45] A drop in the lake's water level represents another threat, and is believed to be driven by water extraction by the increasing human population, climate change and deforestation. [45] The chambo and kampango have been particularly overfished (the kampango declined by about 90% from 2006 to 2016, [46] O. karongae and O. squamipinnis by about 94%, and O. lidole might already be extinct [47] [48] ) and they are now seriously threatened. [49] The IUCN recognises 117 species of Malawi cichlids as threatened; some of these have tiny ranges and may be restricted to rocky coastlines only a few hundred metres long. [50]

Malawi cichlids
Diplotaxodon.jpg
Adult male livingstonii.png
Copadichromis azureus.jpg
Frankfurt Zoo - Aulonocara stuartgranti Usisya.jpg
Fossorochromis rostratus - male en aquarium 01.jpg
Pseudotropheus saulosi.jpg
Oreochromis squamipinnis.jpg
Top to bottom:

1. Diplotaxodon , one of the very few cichlid genera that occurs offshore in relatively deep water. [22]
2. Nimbochromis livingstonii is a piscivorous hap that is famous for playing dead to lure prey close. [10] [51] [52]
3. As typical of utaka, Copadichromis azureus has bright blue males (shown) and duller females that are silvery with dark spots. [53]
4. Aulonocara stuartgranti is part of a group of relatively peaceful species popularly known as peacock cichlids. [54]
5. Fossorochromis rostratus is an "aberrant" hap that often sifts mouthfuls of sand to extract small food organisms. [10] [55]
6. Like many mbuna, Pseudotropheus saulosi is a small cichlid where both male (blue and black) and female (yellow) are colorful. [56]

7. Oreochromis squamipinnis is one of only six native tilapia species in the lake, but these are important to fisheries. Notice the white genital tassels, longer when fully extended and unique to male chambo. [10] [43]

Cichlids

Lake Malawi is noted for being the site of evolutionary radiations among several groups of animals, most notably cichlid fish. [57] There are at least 700 cichlid species in Lake Malawi, [7] with some estimating that the actual figure is as high as 1,000 species. [8] [58] The actual number is labelled with some uncertainty because of the many undescribed species and the extreme variation among some species, making the task of delimiting them very complex. [7] [10] Except for four species ( Astatotilapia calliptera , Coptodon rendalli , Oreochromis shiranus and Serranochromis robustus), all cichlids in the lake are endemic to the Malawi system, which also includes nearby smaller Lake Malombe and the upper Shire River. [10] [59] [60] Many of these have become popular among aquarium owners due to their bright colors. Recreating a Lake Malawi biotope to host cichlids became quite popular in the aquarium hobby. [61] Most Malawi cichlids are found in relatively shallow coastal waters, [10] but Diplotaxodon has been recorded down to depths of 200–220 m (660–720 ft) and several (especially Diplotaxodon, Rhamphochromis and Copadichromis quadrimaculatus ) are known from pelagic waters. [22]

The cichlids of the lake are divided into two groups and the vast majority of the species are haplochromines. The sister species to the Malawi haplochromines is Astatotilapia sp. Ruaha (a currently undescribed species from Great Ruaha River), and these two separated between 2.13 and 6.76 million years ago (mya). [62] The earliest divergence within the Malawi haplochromines occurred between 1.20 and 4.06 mya, [62] but most radiations in this group are far younger; in extreme cases species may have diverged only a few hundred years ago. [17] The Malawi haplochromines are mouthbrooders, but otherwise vary extensively in general behaviour and ecology. [10] Within the Malawi haplochromines there are two main groups, the haps and the mbuna. The haps (they were formerly included in Haplochromis ) can be further subdivided into three subgroups: The relatively large, often more than 20 cm (8 in) long, and aggressive piscivores that roam various habitats in pursuit of prey, the open-water (although often not far from sand or rocks) utaka that feed in schools on zooplankton and typically are of medium size, and finally a subgroup of "aberrant" species that essentially are defined by them not fitting clearly into the other subgroups. [10] [52] [63] Adult male haps generally display bright colors, while juveniles of both sexes and adult females typically show a silvery or grey coloration with sometimes irregular black bars or other markings. [10] [52] The second main haplochromine group are the mbuna, a name used both locally and popularly, which means "rockfish" in Tonga. [64] They are found at rocky outcrops, territorially aggressive (although commonly found in high densities) and often specialised aufwuchs feeders. [10] [52] The mbuna species tend to be relatively small, mostly less than 13 cm (5 in) long, and often both sexes are brightly colored with males having egg-shaped yellow spots on their anal fin (a feature particularly prevalent in the mbuna, but not exclusive to this group). [10] [52]

The second group, the tilapia, comprises only six species in two genera in Lake Malawi: The redbreast tilapia (Coptodon rendalli), a widespread African species, is the only substrate-spawning cichlid in the lake. [10] [65] This large cichlid mainly feeds on macrophytes. [10] [66] The remaining are five mouthbrooding species of Oreochromis ; four chambo in the subgenus Nyasalapia (O. karongae, O. lidole, O. saka and O. squamipinnis) that are endemic to the Lake Malawi system, as well as the closely related O. shiranus, which also is found in Lake Chilwa. [10] [43] [59] The Malawi Oreochromis mainly feed on phytoplankton, reach lengths up to 26–42 cm (10–17 in) depending on the exact species, and are mostly black or silvery-gray with relatively indistinct dark bars. [10] [59] [67] Male chambo have unique genital tassels when breeding, which aid in egg fertilisation in a manner comparable to the egg-spots on the anal fin of haplochromines. [10] [43]

Non-cichlids

The kampango (Bagrus meridionalis), one of the largest catfish, reaching up to 1.5 m (4.9 ft) in length Catfish in Lake Malawi.jpg
The kampango (Bagrus meridionalis), one of the largest catfish, reaching up to 1.5 m (4.9 ft) in length

The vast majority of the fish species in the lake are cichlids. Among the non-cichlid native fish are several species of cyprinids (in genera Barbus , Labeo and Opsaridium , and the Lake Malawi sardine Engraulicypris sardella), airbreathing catfish ( Bathyclarias and Clarias , and the kampango Bagrus meridionalis), mochokid catfish ( Chiloglanis and Malawi squeaker Synodontis njassae), Mastacembelus spiny eel, mormyrids ( Marcusenius , Mormyrops and Petrocephalus ), the African tetra Brycinus imberi , the poeciliid Aplocheilichthys johnstoni , the spotted killifish (Nothobranchius orthonotus), and the mottled eel (Anguilla nebulosa). [10]

At a genus level, most of these are widespread in Africa, but Bathyclarias is entirely restricted to the lake. [69]

Invertebrates

Molluscs

Lake Malawi is home to 28 species of freshwater snails (including 16 endemics) and 9 bivalves (2 endemics, Aspatharia subreniformis and the unionid Nyassunio nyassaensis ). [70] [71] The endemic freshwater snails are all members of the genera Bellamya , Bulinus , Gabbiella , Lanistes and Melanoides . [72]

Lake Malawi is home to a total of four snail species in the genus Bulinus, which is a known intermediate host of bilharzia. A survey in Monkey Bay in 1964 found two endemic species of snails of the genus ( B. nyassanus and B. succinoides ) in the lake, and two non-endemic species ( B. globosus and B. forskalli ) in lagoons separated from it. The latter species are known intermediate hosts of bilharzia, and larvae of the parasite were detected in water containing these, but in experiments C. Wright of the British Museum of Natural History was unable to infect the two species endemic to the lake with the parasites. The field workers, who spent many hours on and in the lake, did not find either B. globosus or B. forskalli in the lake itself. [73] More recently, the disease has become a problem in the lake itself as the endemic B. nyassanus has become an intermediate host. This change, first noticed in the mid-1980s, is possibly related to a decline in snail-eating cichlids (for example, Trematocranus placodon ) due to overfishing and/or a new strain of the bilharzia parasite. [21]

Crustaceans

Unlike Lake Tanganyika with its many endemic freshwater crabs and shrimp, there are few such species in Lake Malawi. The Malawi blue crab, Potamonautes lirrangensis (syn. P. orbitospinus), is the only crab in the lake and it is not endemic. [74] [75] The atyid shrimp Caridina malawensis is endemic to the lake, but it is poorly known and has historically been confused with C. nilotica , which is not found in the lake. [76] Pelagic zooplanktonic species include two cladocerans ( Diaphanosoma excisum and Bosmina longirostris ), three copepods ( Tropodiaptomus cunningtoni, Thermocyclops neglectus and Mesocyclops aequatorialis), [77] and several ostracods (including both described and undescribed species). [78]

Lake flies

Huge swarms of lake flies (Chaoborus edulis), resembling distant plumes of smoke over the water Lake flies, Lake Malawi (2499202894).jpg
Huge swarms of lake flies ( Chaoborus edulis), resembling distant plumes of smoke over the water

Lake Malawi is famous for the huge swarms of tiny, harmless lake flies, Chaoborus edulis. [79] These swarms, typically appearing far out over water, can be mistaken for plumes of smoke and were also noticed by David Livingstone when he visited the lake. [79] [80] [81] The aquatic larvae feed on zooplankton, spending the day at the bottom and the night in the upper water levels. [79] When they pupate they float to the surface and transform into adult flies. [81] The adults are very short-lived and the swarms, which can be several hundred metres tall and often have a spiraling shape, are part of their mating behaviour. [79] [82] They lay their eggs at the water's surface and the adults die. [82] The larvae are an important food source for fish, [77] [79] [83] and the adult flies are important both to birds and local people, who collect them to make kungu cakes/burgers, a local delicacy with a very high protein content. [80] [81]

2015 mine leak

In January 2015, a sediment control tank collapsed at the Paladin Energy-owned uranium mine in Northern Malawi after a high intensity rain storm hit the area. It was revealed that approximately 50 litres of non radioactive material leaked into a local creek. Despite reports in local media of radioactive contamination the government conducted independent scientific tests on the local river system and found that there was no effect on the environment. [84] [85]

Swimming

The 25 km solo swim across Lake Malawi between Cape Ngomba and Senga Bay has been accomplished on 5 occasions by 16 swimmers

1992: Lewis Pugh 9hrs 52 minutes (UK/SA) [86] and Otto Thanning (SA) 10hrs 5 minutes

2010: Abigail Brown (UK) 9hrs 45 minutes [87]

2013: Milko van Gool (Netherlands) 8hrs 46 minutes [88] and Kaitlin Harthoorn (US) 9hrs 17 minutes

2016: (current record) Jean Craven (SA), Robert Dunford (Kenya), Michiel Le Roux (SA), Samantha Whelpton (SA), Greig Bannatyne (SA), Haydn Von Maltitz (SA), Douglas Livingstone-Blevins (SA) 7hrs 53 mins [89]

2019: Chris Stapley (Eswatini) and Jay Azran (SA) 8hrs 40 minutes, Andrew Stevens (Australia) 10hrs 50 minutes, and Ruth Azran (SA) 11hrs 8 minutes. [90] [91]

In 2019, Martin Hobbs (SA), became the first person to swim the full length of Lake Malawi (54 days), as well as setting the world record for longest solo swim in a lake [92]

See also

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<i>Labidochromis caeruleus</i> Species of fish

Labidochromis caeruleus is a species of cichlid endemic to the central western coastal region of Lake Malawi in East Africa. It is also known as lemon yellow lab, the blue streak hap, the electric yellow or yellow prince, depending on the colour morph. A naturally occurring yellow-coloured variant from Lion's Cove is one of the most popular cichlids amongst aquarium hobbyists.

Mbuna Common name for several species of fish

Mbuna is the common name for a large group of African cichlids from Lake Malawi, and are members of the haplochromine family. The name mbuna means "rockfish" in the language of the Tonga people of Malawi. As the name implies, most mbuna are cichlids that live among the piles of rocks and along the rocky shores of Lake Malawi, as opposed to the utaka, cichlids that live in the open water or on sandy shores or soft substrates. Some species of mbuna are highly sexually dimorphic, although many are not. Among biologists, almost all of the cichlid species of Lake Malawi, including mbuna and non mbuna such as the utaka, are believed to have descended from one or a very few species that became isolated in the lake. With rising water levels, new habitats could be colonized and the many isolated rocky outcrops allowed new mbuna species to form. Their striking colors, intriguing behavioral characteristics, and relative hardiness make them very popular despite their unique demands for the home aquarist.

<i>Maylandia estherae</i> Species of fish

Maylandia estherae is a haplochromine cichlid. It is a rock dwelling fish or mbuna from Lake Malawi. This fish, like most cichlids from Lake Malawi, is a mouthbrooder - females hold their fertilized eggs then fry in their mouths until they are released after about 21 days.

<i>Maylandia</i> Genus of fishes

Maylandia or Metriaclima is a genus of haplochromine cichlids endemic to Lake Malawi in East Africa. They belong to the mbuna (rock-dwelling) haplochromines.

Haplochromine Tribe of fishes

The haplochromine cichlids are a tribe of cichlids in subfamily Pseudocrenilabrinae called Haplochromini. This group includes the type genus (Haplochromis) plus a number of closely related genera such as Aulonocara, Astatotilapia, and Chilotilapia. They are endemic to eastern, southern and northern Africa, except for Astatotilapia flaviijosephi in the Middle East. A common name in a scientific context is East African cichlids – while they are not restricted to that region, they are the dominant Cichlidae there. This tribe was extensively studied by Ethelwynn Trewavas, who made major reviews in 1935 and 1989, at the beginning and at the end of her career in ichthyology. Even today, numerous new species are being described each year.

<i>Diplotaxodon</i> Genus of fishes

Diplotaxodon is a small genus of seven formally described, as well as a number of undescribed, deep-water species of cichlid fish endemic to Lake Malawi in east Africa. These fishes represent a remarkable adaptive radiation of offshore and deep-water adapted fish descended from ancestral shallow water forms. They include the dominant zooplankton-feeding fish of the offshore and deep-water regions of the lake, as well as a number of larger species that appear to feed on small pelagic fishes. Adult sizes range from 10 to 30 cm in total length, depending on species.

<i>Nimbochromis polystigma</i> Species of fish

Nimbochromis polystigma is a species of cichlid endemic to Lake Malawi, in the Great Rift Valley system of Africa. This species prefers areas with rock/sand substrate or areas with plentiful vegetation. It can grow to a length of 23 centimetres (9.1 in) TL.

<i>Copadichromis borleyi</i> Species of fish

Copadichromis borleyi is a species of haplochromine cichlid fish endemic to Lake Malawi in East Africa. The species is popular in the fishkeeping hobby where it is frequently kept in aquariums. The species has numerous common names, including redfin hap and goldfin hap.

<i>Pseudotropheus johannii</i> Species of fish

Pseudotropheus johannii or the bluegray mbuna is an African freshwater fish from the family Cichlidae.

<i>Buccochromis atritaeniatus</i> Species of fish

Buccochromis atritaeniatus is a species of haplochromine cichlid. It is found in Lake Malawi in Malawi, Mozambique, and Tanzania. Its natural habitat is freshwater lakes and intermediate and sandy areas and has been caught between 10 and 40 meters; it feeds on Haplochromines and hunts in pairs or large groups. Breeding males are found in colonies at depths about 30 meters and, while breeding, build sandcastle nests on steeply sloping sandy shores. The only known threat is fisheries and is known as Haplochromis mbowe in the fishing trade. Although FishBase and the Catalog of Fishes recognise this species, the IUCN treats it as a junior synonym of Buccochromis nototaenia.

Buccochromis oculatus is a species of haplochromine cichlid. It is endemic to Lake Malawi where it is found at 18 meters depth in Malawi, Mozambique, and Tanzania. Its natural habitat is freshwater lakes. The only threat may be potential overfishing. This taxon is regarded a junior synonym of Buccochromis nototaenia by the IUCN.

Verduya's hap is a species of haplochromine cichlid which is endemic to Lake Malawi where it has a lake wide range and so is found in Malawi, Mozambique, and Tanzania. This species occurs in open water, sometimes forming large shoals.

<i>Corematodus shiranus</i> Species of fish

Corematodus shiranus is a species of haplochromine cichlid fish native to Lake Malawi, Lake Malombe and the upper reaches of the Shire River in East Africa. It is an aggressive mimic of the chambo cichlids in both color pattern and swimming mode. It is therefore able to approach unsuspecting schools of these species and rapidly take a mouthful of scales or fin. Due to overfishing, the chambo cichlid populations have drastically declined, adversely affecting C. shiranus, which was last reported in 1997 and might now be extinct.

<i>Oreochromis karongae</i> Species of fish

Oreochromis karongae is a critically endangered species of cichlid that is endemic to Lake Malawi, Lake Malombe, and upper and middle Shire River in Malawi, Mozambique, and Tanzania. This species is important to local commercial fisheries, but has declined drastically due to overfishing.

<i>Oreochromis lidole</i> Species of fish

Oreochromis lidole is a species of freshwater fish in the family Cichlidae. This tilapia is native to Malawi, Mozambique and Tanzania, where it is found in Lake Malawi, Lake Malombe, the Shire River and perhaps some crater lakes further north. It is important in fisheries, but has drastically declined; it may already be extinct. This oreochromine cichlid is locally called chambo, a name also used for two other closely related species found in the same region, O. karongae and O. squamipinnis.

<i>Oreochromis squamipinnis</i> Species of fish

Oreochromis squamipinnis is a critically endangered species of cichlid fish that is endemic to Lake Malawi, Lake Malombe and Shire River in East Africa, where found in a wide range of habitats, but especially in shallow water. This species is important to local commercial fisheries and can also be found in the aquarium trade, but it has declined drastically due to overfishing.

<i>Oreochromis variabilis</i> Species of fish

Oreochromis variabilis, the Victoria tilapia, is a species of African cichlid native to Lake Victoria and its tributaries, Lake Kyoga, Lake Kwania, and Lake Bisina (Salisbury), as well as being found in the Victoria Nile above Murchison Falls. This species can reach a standard length of 30 cm (12 in). This species is important to local commercial fisheries and is potentially important in aquaculture. It is also found in the aquarium trade.

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Further reading