Tyto Temporal range: | |
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African grass owl, Tyto capensis The "grass owls" are two rather long-legged species of Tyto. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Strigiformes |
Family: | Tytonidae |
Subfamily: | Tytoninae |
Genus: | Tyto Billberg, 1828 |
Type species | |
Tyto alba (formerly Strix flammea) [1] Linnaeus, 1766 | |
Species | |
see text | |
Synonyms | |
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Tyto is a genus of birds consisting of true barn owls, grass owls and masked owls that collectively make up all the species within the subfamily Tytoninae of the barn owl family, Tytonidae.
The genus Tyto was introduced in 1828 by the Swedish naturalist Gustaf Johan Billberg with the western barn owl as the type species. [2] [3] The name is from the Ancient Greek tutō meaning "owl". [4]
The barn owl (Tyto alba) was formerly considered to have a global distribution with around 28 subspecies. [5] In the list of birds maintained by Frank Gill, Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC) the barn owl is now split into four species: the western barn owl (Tyto alba) (10 subspecies), the American barn owl (Tyto furcata) (12 subspecies), the eastern barn owl (Tyto javanica) (7 subspecies) and the Andaman masked owl (Tyto deroepstorffi). [6] This arrangement is followed here. Some support for this split was provided by a molecular phylogenetic study by Vera Uva and collaborators published in 2018 that compared the DNA sequences of three mitochondrial and one nuclear loci. [7] This split has not been adopted by other taxonomic authorities such as the Clements Checklist of Birds of the World maintained by members of Cornell University or by the list maintained by BirdLife International that is used by the International Union for Conservation of Nature. [8] [9]
The cladogram below is based on the 2018 phylogenetic study. The Andaman masked owl (Tyto deroepstorffi) and Itombwe owl (Tyto prigoginei) were not sampled. The Manus masked owl (Tyto manusi) was embedded in a clade with subspecies of the Australian masked owl. [7]
Tyto |
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Throughout their evolutionary history, Tyto owls have shown a better capability to colonize islands than other owls. Several such island forms have become extinct, some long ago, but some in comparatively recent times. A number of insular barn owls from the Mediterranean and the Caribbean were very large or truly gigantic species.
Seventeen species are recognized: [6]
Image | Common name | Scientific name | Distribution |
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Greater sooty owl | Tyto tenebricosa | Australia | |
Lesser sooty owl | Tyto multipunctata | Australia | |
Minahasa masked owl | Tyto inexspectata | Sulawesi, Indonesia | |
Taliabu masked owl | Tyto nigrobrunnea | Sula Islands, Maluku, Indonesia | |
Moluccan masked owl | Tyto sororcula | south Moluccas of Indonesia | |
Manus masked owl | Tyto manusi | Manus Island in the Admiralty Islands | |
Golden masked owl | Tyto aurantia | the island of New Britain, Papua New Guinea | |
Australian masked owl | Tyto novaehollandiae | Southern New Guinea and the non-desert areas of Australia. | |
Sulawesi masked owl | Tyto rosenbergii | the Indonesian islands of Sulawesi, Sangihe and Peleng | |
Red owl | Tyto soumagnei | Madagascar | |
Western barn owl | Tyto alba | Eurasia and Africa. | |
American barn owl | Tyto furcata | the Americas | |
Eastern barn owl | Tyto javanica | southeast Asia and Australasia. | |
Andaman masked owl | Tyto deroepstorffi | southern Andaman Islands | |
Ashy-faced owl | Tyto glaucops | Hispaniola (Haiti and the Dominican Republic). | |
African grass owl | Tyto capensis | southern Congo and northern Angola to the central coast of Mozambique and the other centred on South Africa from the Western Cape north to the southern extremities of Zimbabwe, Botswana and Mozambique up to Kenya and Ethiopia. | |
Eastern grass owl | Tyto longimembris | eastern, southern and southeast Asia, parts of New Guinea, Australia (mainly in Queensland) and the western Pacific | |
Itombwe owl | Tyto prigoginei | Itombwe Mountains in the eastern Democratic Republic of Congo | |
A number of owl fossils were at one time assigned to the present genus, but are nowadays placed elsewhere. While there are clear differences in osteology between typical owls and barn owls, there has been parallel evolution to some degree and thus isolated fossil bones cannot necessarily be assigned to either family without thorough study. Notably, the genus Strix has been misapplied by many early scientists as a "wastebasket taxon" for many owls, including Tyto. [12]
They are darker on the back than the front, usually an orange-brown colour, the front being a paler version of the back or mottled, although there is considerable variation even amongst species. Tyto owls have a divided, heart-shaped facial disc, and lack the ear-like tufts of feathers found in many other owls. Tyto owls tend to be larger than bay owls. The name tyto (τυτώ) is onomatopeic Greek for owl.
Systematics and distribution of the living and fossil small barn owls of the West Indies (Aves: Strigiformes: Tytonidae)
". Zootaxa. 4830 (3): 544–564. doi:10.11646/zootaxa.4830.3.4. ISSN 1175-5334. PMID 33056145. S2CID 222819958.Owls are birds from the order Strigiformes, which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers adapted for silent flight. Exceptions include the diurnal northern hawk-owl and the gregarious burrowing owl.
The true owls or typical owls are one of the two generally accepted families of owls, the other being the barn owls and bay owls (Tytonidae). This large family comprises 230 living or recently extinct species in 24 genera. The Strigidae owls have a cosmopolitan distribution and are found on every continent except Antarctica.
The bird family Tytonidae, which includes the barn owlsTyto and the bay owlsPhodilus, is one of the two families of owls, the other being the true owls or typical owls, Strigidae. They are medium to large owls with large heads and characteristic heart-shaped faces. They have long, strong legs with powerful talons. They also differ from the Strigidae in structural details relating in particular to the sternum and feet.
The American horned owls and the Old World eagle-owls make up the genus Bubo, at least as traditionally described. The genus name Bubo is Latin for owl.
The barn owls are the most widely distributed group of owls in the world. They are medium-sized owls with large heads and characteristic heart-shaped faces. They have long, strong legs with powerful talons. The term may be used to describe:
Tringa is a genus of waders, containing the shanks and tattlers. The genus name Tringa is the Neo-Latin name given to the green sandpiper by the Italian naturalist Ulisse Aldrovandi in 1599. They are mainly freshwater birds, often with brightly coloured legs as reflected in the English names of six species, as well as the specific names of two of these and the green sandpiper. They are typically associated with northern hemisphere temperate regions for breeding. Some of this group—notably the green sandpiper—nest in trees, using the old nests of other birds, usually thrushes.
Gargano is a historical and geographical sub-region in the province of Foggia, Apulia, southeast Italy, consisting of a wide isolated mountain massif made of highland and several peaks and forming the backbone of the Gargano Promontory projecting into the Adriatic Sea, the "spur" on the Italian "boot".
Asio is a genus of typical owls, or true owls, in the family Strigidae. This group has representatives over most of the planet, and the short-eared owl is one of the most widespread of all bird species, breeding in Europe, Asia, North and South America, the Caribbean, Hawaii and the Galápagos Islands. Its geographic range extends to all continents except Antarctica and Australia.
Strix is a genus of owls in the typical owl family (Strigidae), one of the two generally accepted living families of owls, with the other being Tytonidae. Common names are earless owls or wood owls, though they are not the only owls without ear tufts, and "wood owl" is also used as a more generic name for forest-dwelling owls.
Ninox is a genus of true owls comprising 36 species found in Asia and Australasia. Many species are known as hawk-owls or boobooks, but the northern hawk-owl is not a member of this genus.
Lybiidae is a family of birds also known as the African barbets. There are 42 species ranging from the type genus Lybius of forest interior to the tinkerbirds (Pogoniulus) of forest and scrubland. They are found throughout sub-Saharan Africa, with the exception of the far south-west of South Africa.
Athene is a genus of owls, containing nine living species, depending on classification. These birds are small, with brown and white speckles, yellow eyes, and white eyebrows. This genus is found on all continents except for Australia, Antarctica, and Sub-Saharan Africa. An evolutionary radiation of 4 species is also present in the Solomon Islands.
The Sulawesi masked owl is a species of owl in the family Tytonidae. It is endemic to the Indonesian islands of Sulawesi, Sangihe and Peleng. It is listed by the IUCN as being of least concern.
The African terrestrial barbets are the bird genus Trachyphonus in the African barbet family (Lybiidae), which was formerly included in the Capitonidae and sometimes in the Ramphastidae. These birds are more terrestrial than the other African barbets and differ in some other respects too; they are thus separated in a monotypic subfamily Trachyphoninae.
Tyto robusta was a prehistoric barn-owl. It lived at what is now Monte Gargano in Italy, and was an island throughout much of the Neogene when sea levels were higher. The owl's remains date back to the Miocene-Pliocene boundary 5.5 to 5 million years ago. The fossil bones are about 60% as long again as a modern barn owl, giving a total length of about 50–65 cm for T. robusta. This owl provides an interesting case study of evolution and insular gigantism.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1969.
The Andaman masked owl is a barn owl endemic to the southern Andaman Islands archipelago of India, in the Bay of Bengal, Indian Ocean. Regarded by some authors as a subspecies of the common barn owl, it is recognized by others as a species in its own right.
The eastern barn owl is usually considered a subspecies group and together with the American barn owl group, the western barn owl group, and sometimes the Andaman masked owl make up the barn owl. The cosmopolitan barn owl is recognized by most taxonomic authorities. A few separate them into distinct species, as is done here. The eastern barn owl is native to southeastern Asia and Australasia.
The northern masked owl is a large forest owl in the family Tytonidae. The northern kimberli subspecies was identified as a novel race of the Australian masked owl by the Australian ornithologist Gregory Macalister Mathews in his 1912 reference list of Australian birds. The northern masked owl occurs in forest and woodland habitats in northern Australia, ranging from the northern Kimberley region to the northern mainland area of the Northern Territory and the western Gulf of Carpentaria. While the Australian masked owl is recognized as the largest species in the family Tytonidae, the northern masked owl is one of the smallest of the Australian masked owl subspecies.