2013 in paleobotany

Last updated

List of years in paleobotany
In paleontology
2010
2011
2012
2013
2014
2015
2016
In arthropod paleontology
2010
2011
2012
2013
2014
2015
2016
In paleoentomology
2010
2011
2012
2013
2014
2015
2016
In paleomalacology
2010
2011
2012
2013
2014
2015
2016
In reptile paleontology
2010
2011
2012
2013
2014
2015
2016
In archosaur paleontology
2010
2011
2012
2013
2014
2015
2016
In mammal paleontology
2010
2011
2012
2013
2014
2015
2016
In paleoichthyology
2010
2011
2012
2013
2014
2015
2016

This list of 2013 in paleobotany records new fossil plant taxa that were described during 2013, as well as other significant discoveries and events related to paleobotany that occurred in the year.

Contents

Chlorophyta

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Cymatiosphaera scitula [1]

Sp. nov.

Valid

Wicander & Playford

Devonian (Famennian)

Saverton Shale

Flag of the United States.svg  United States

A member of Chlorophyta belonging to the group Prasinophyceae.

Bryophyta

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Brachytheciites sachalinensis [2]

Sp. nov

Valid

Ignatov & Perkovsky

Middle Eocene

Sakhalinian amber

Flag of Russia.svg  Russia

A moss belonging to the family Brachytheciaceae.

Bryiidites [3]

Gen. et sp. nov.

Valid

Barclay et al.

Late Cretaceous (late Cenomanian)

Middle Dakota Formation

Flag of the United States.svg  United States

A member of Bryidae of uncertain phylogenetic placement. The type specimen is Bryiidites utahensis.

Bryokhutuliinia crassimarginata [4]

Sp. nov

Valid

Ignatov et al.

Late Jurassic

Flag of Russia.svg  Russia

A moss.

Calymperites [5]

Gen. et sp. nov

Valid

Ignatov & Perkovsky

Late Eocene

Rovno amber

Flag of Ukraine.svg  Ukraine

A moss belonging to the group Dicranales (sensu lato). Genus includes new species C. ucrainicus.

Pseudoleskeellites [2]

Gen. et sp. nov

Valid

Ignatov & Perkovsky

Middle Eocene

Sakhalinian amber

Flag of Russia.svg  Russia

A moss belonging to the order Hypnales. Genus includes new species P. obscurus.

Ferns and fern allies

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Ashicaulis beipiaoensis [6]

Sp. nov

Valid

Tian et al.

Middle Jurassic

Tiaojishan Formation

Flag of the People's Republic of China.svg  China

A member of Osmundaceae. Originally described as a species of Ashicaulis ; transferred to the genus Millerocaulis by Bomfleur, Grimm & McLoughlin (2017). [7]

Mygdykitia [8]

Gen. et sp. nov

Valid

Golovneva & Shczepetov

Late Cretaceous (Campanian)

Ola Formation

Flag of Russia.svg  Russia

A fern belonging to the family Dipteridaceae. Genus includes new species M. ochotica.

Palaeosorum [9]

Gen. et sp. nov

Valid

Jacques & Zhou in Jacques, Su & Zhou

Miocene

Dajie Formation

Flag of the People's Republic of China.svg  China

A member of Polypodiaceae. The type specimen is Palaeosorum ellipticum.

Todea amissa [10]

Sp. nov.

Valid

Carvalho in Carvalho et al.

Early Eocene

Flag of Argentina.svg  Argentina

A species of Todea .

Yavanna [11]

Gen. et sp. nov

Valid

Vera

Early Cretaceous (Aptian)

Cerro Negro Formation

Blank flag.svg  Antarctica
(Livingston Island)

A cyathealean of uncertain placement. Type species is Y. chimaerica.

Ginkgophytes

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Allicospermum angrenicum [12]

Sp. nov.

Valid

Nosova

Middle Jurassic

Angren Formation

Flag of Uzbekistan.svg  Uzbekistan

A Ginkgo -like plant described on the basis of seeds.

Nagrenia [12]

Gen. et sp. nov.

Valid

Nosova

Middle Jurassic

Angren Formation

Flag of Uzbekistan.svg  Uzbekistan

A member of Ginkgoales described on the basis of peduncles furcating into several pedicels. Genus includes new species N. samylinae.

Gnetophyta

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Chengia [13]

Gen. et sp. nov.

Valid

Yang, Lin & Wang

Early Cretaceous (Aptian)

Yixian Formation

Flag of the People's Republic of China.svg  China

A member of Ephedraceae. The type specimen is Chengia laxispicata.

Friedsellowia gracilifolia [14]

Gen. nov. et sp. nov.

Valid

Löwe et al.

Early Cretaceous

Crato Formation

Flag of Brazil.svg  Brazil

A gnetophyte. The type of a new genus.

Itajuba [ citation needed ]

Gen. et sp. nov

Valid

Ricardi-Branco et al.

Early Cretaceous (Aptian or Albian)

Crato Member of the Santana Formation

Flag of Brazil.svg  Brazil

A member of Gnetales. The type specimen is Itajuba yansanae.

Pachytestopsida

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Stephanospermum braidwoodensis [15]

Sp. nov.

Valid

Spencer, Hilton & Sutton

Carboniferous

Mazon Creek fossil beds

Flag of the United States.svg  United States

A member of Medullosales, a species of Stephanospermum.

Stephanospermum shuichengensis [16]

Sp. nov.

Valid

Spencer et al.

Permian (Lopingian)

Wangjiazhai Formation

Flag of the People's Republic of China.svg  China

A member of Medullosales, a species of Stephanospermum.

Pinophyta

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Bancroftiastrobus [17]

Gen. et sp. nov.

Valid

Rothwell et al.

Late Jurassic (early Kimmeridgian)

Kimmeridge Clay Formation

Flag of the United Kingdom.svg  United Kingdom

A pinophyte of uncertain phylogenetic placement.
Type species is B. digitata.

Brachyoxylon currumilii [18]

Sp. nov.

Valid

Bodnar et al.

Early–Middle Jurassic

Cañadón Asfalto Formation

Flag of Argentina.svg  Argentina

A member of Pinophyta described on the basis of fossil wood.

Cunninghamia taylorii [19]

Sp. nov.

Valid

Serbet, Bomfleur & Rothwell

Late Cretaceous (Campanian)

Horseshoe Canyon Formation

Flag of Canada (Pantone).svg  Canada

A species of Cunninghamia .

Elatocladus anorovii [8]

Sp. nov

Valid

Golovneva & Shczepetov

Late Cretaceous (Campanian)

Ola Formation

Flag of Russia.svg  Russia

A conifer of uncertain phylogenetic placement.

Pagiophyllum heshanense [20]

Sp. nov.

Valid

Liu & Yao

Permian

Flag of the People's Republic of China.svg  China

A species of Pagiophyllum.

Pararaucaria delfueyoi [21]

Sp. nov.

Valid

Escapa et al.

Late Jurassic

Cañadón Calcáreo Formation

Flag of Argentina.svg  Argentina

A member of Pinophyta belonging to the family Cheirolepidiaceae, a species of Pararaucaria.

Picea yanensis [8]

Sp. nov

Valid

Golovneva & Shczepetov

Late Cretaceous (Campanian)

Ola Formation

Flag of Russia.svg  Russia

A spruce.

Prumnopitys anglica [22]

Sp. nov.

Valid

Greenwood, Hill & Conran

Eocene

Branksome Formation

Flag of the United Kingdom.svg  United Kingdom

A species of Prumnopitys .

Schizolepidopsis canicularis [23]

Sp. nov.

Valid

Leslie et al.

Early Cretaceous (probably Aptian-Albian)

Tevshiin Govi Formation

Flag of Mongolia.svg  Mongolia

A member of Pinophyta, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis.

Xuanweioxylon [24]

Gen. et sp. nov.

Valid

He et al.

Permian (late Wuchiapingian-Changhsingian)

Xuanwei Formation

Flag of the People's Republic of China.svg  China

A member of Pinophyta. Genus includes new species X. scalariforme.

Angiosperms

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Anonaspermum warmanensis [25]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A member of Annonaceae.

Atli [26]

Gen. et sp. nov.

Valid

Smith et al.

Late Cretaceous (Campanian)

Flag of Canada (Pantone).svg  Canada

A ranunculalean known from a stem with lianoid anatomy. The type specimen is Atli morinii.

Bambusiculmus [27]

Gen. et 2 sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

Flag of the People's Republic of China.svg  China

A bamboo. The type specimen is Bambusiculmus angustus; genus also includes Bambusiculmus latus.

Bambusium angustifolia [27]

Sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

Flag of the People's Republic of China.svg  China

A bamboo, a species of Bambusium .

Bambusium latifolia [27]

Sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

Flag of the People's Republic of China.svg  China

A bamboo, a species of Bambusium .

Bombacacioxylon [28]

Gen. et sp. nov.

Valid

Mehrotra et al.

Late Miocene to early Pliocene

Tipam Group

Flag of India.svg  India

A possible relative of Bombax . The type specimen is Bombacacioxylon tertiarum.

Carpolithus warmanensis [25]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A fossil fruit.

Cornerocarpon [25]

Gen. et sp. nov.

Valid

Grote in Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A member of Rosales belonging or related to the family Moraceae. Genus includes new species C. copiosum.

Corylopsis grotei [25]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A species of Corylopsis .

Diospyros neyveliensis [29]

Sp. nov.

Valid

Mukherjee & Prasad

Miocene

Neyveli Formation

Flag of India.svg  India

A member of the family Ebenaceae described on the basis of fossil wood. Originally described as a species of Diospyros ; Srivastava et al. (2018) transferred this species to the genus Ebenoxylon. [30]

Firmiana oligocenica [31]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

Flag of India.svg  India

A species of Firmiana .

Grotea [25]

Gen. et sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A member of the family Bignoniaceae. Genus includes new species G. warmanensis.

Guadua morronei [32]

Sp. nov.

Valid

Brea, Zucol & Franco

Pliocene–Pleistocene

Ituzaingó Formation

Flag of Argentina.svg  Argentina

A bamboo, a species of Guadua .

Hamamelis warmanensis [25]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A witch-hazel.

Indovitis [33]

Gen. et sp. nov.

Valid

Manchester, Kapgate & Wen

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

Flag of India.svg  India

A member of the family Vitaceae. Genus includes new species I. chitaleyae.

Iterophyllum [34]

Gen. et sp. nov.

Valid

Barral et al.

Early Cretaceous (late Barremian)

Flag of Spain.svg  Spain

A basal eudicot. The type specimen is Iterophyllum lobatum.

Jaguariba wiersemana [35]

Gen. nov. et sp. nov.

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous

Crato Formation

Flag of Brazil.svg  Brazil

Fossil Nymphaeales.

Lagerstroemia corvinusii [36]

Sp. nov.

Valid

Prasad

Mio-Pliocene

Churia Formation

Flag of Nepal.svg    Nepal

A species of Lagerstroemia .

Laurelia otagoensis [37]

Sp. nov

Valid

Conran, Bannister & Lee

Early Miocene

Flag of New Zealand.svg  New Zealand

A species of Laurelia .

Lauroflorum [25]

Gen. et sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

Flag of the United States.svg  United States

A member of Lauraceae. Genus includes new species L. warmanensis.

Lindera acuminatissima [38]

Sp. nov

Valid

Dao & Sun in Dao et al.

Late Pliocene

Mangbang Formation

Flag of the People's Republic of China.svg  China

A species of Lindera .

"Magnolia" szakmanycsabae [39]

Sp. nov.

Valid

Hably

Late Miocene

Flag of Hungary.svg  Hungary

A plant of uncertain phylogenetic placement, possibly a relative of MagnoliaceaeLauraceae.

Micropetasos [40]

Gen. et sp. nov

Valid

Poinar, Chambers & Wunderlich

Late Cretaceous (Cenomanian)

Burmese amber

Flag of Myanmar.svg  Myanmar

An early flowering plant of uncertain phylogenetic placement. Genus includes new species M. burmensis.

Myrtineoxylon [41]

Gen. et sp. nov.

Valid

Oskolski, Feng & Jin

Late Eocene

Youganwo Formation

Flag of the People's Republic of China.svg  China

A member of Myrtaceae described on the basis of fossil wood. Genus includes new species M. maomingensis.

Palmoxylon vastanensis [42]

Sp. nov.

Valid

Prasad et al.

Eocene

Cambay Shale Formation

Flag of India.svg  India

A palm tree, a species of Palmoxylon .

Paranephelium makumensis [31]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

Flag of India.svg  India

A species of Paranephelium .

Poeciloneuron preindicum [43]

Sp. nov.

Valid

Srivastava & Mehrotra

Late Oligocene

Flag of India.svg  India

A species of Poeciloneuron .

Pongamia arjunkholaensis [36]

Sp. nov.

Valid

Prasad

Mio-Pliocene

Churia Formation

Flag of Nepal.svg    Nepal

A species of Pongamia .

Potomacapnos [44]

Gen. et sp. nov.

Valid

Jud & Hickey

Early Cretaceous (Aptian)

Potomac Group

Flag of the United States.svg  United States

A close relative of ranunculalean eudicots. The type specimen is Potomacapnos apeleutheron.

Pterygota palaeoalata [31]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

Flag of India.svg  India

A species of Pterygota .

Sapindus palaeoemarginatus [31]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

Flag of India.svg  India

A species of Sapindus .

Spixiarum kipea [45]

Gen. nov. et sp. nov.

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous

Crato Formation

Flag of Brazil.svg  Brazil

Fossil Aroid.

Trapa pannonica [39]

Sp. nov.

Valid

Hably

Late Miocene

Flag of Hungary.svg  Hungary

A water caltrop.

Trapa rozsaszentmartoni [39]

Sp. nov.

Valid

Hably

Late Miocene

Flag of Hungary.svg  Hungary

A water caltrop.

Virola dominicana [46]

Sp. nov.

Valid

Poinar & Steeves

Cenozoic (Eocene or Miocene)

Flag of the Dominican Republic.svg  Dominican Republic

A member of Myristicaceae found in Dominican amber, a species of Virola .

Vitis szakmanygyorgyi [39]

Sp. nov.

Valid

Hably

Late Miocene

Flag of Hungary.svg  Hungary

A species of Vitis .

Other seed plants

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Donponoxylon [47]

Gen. et 2 sp. nov.

Valid

Tidwell, Britt & Wright

Middle to Late Jurassic

Flag of Australia (converted).svg  Australia
Flag of New Zealand.svg  New Zealand

A seed plant of uncertain phylogenetic placement. The type specimen is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii.

Dordrechtites arcanus [48]

Sp. nov.

Valid

Bergene, Taylor & Taylor

Middle Triassic (Anisian)

Fremouw Formation

Blank flag.svg  Antarctica

A gymnosperm of uncertain phylogenetic placement.

Other plants

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Acinosporites tristratus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jubah Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Alatisporites? trisacculus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Biornatispora elegantula [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Belgium (civil).svg  Belgium
Flag of Germany.svg  Germany
Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Biornatispora microclavata [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Brochotriletes crameri [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Jubah Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Brochotriletes tenellus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Brochotriletes tripapillatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin II Formation

Flag of Libya.svg  Libya

A trilete spore.

Camarozonotriletes asperulus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Awaynat Wanin III Formation

Flag of Brazil.svg  Brazil
Flag of Tunisia.svg  Tunisia

A trilete spore.

Chelinospora carnosa [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Chelinospora condensata [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Chelinospora densa [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Chelinospora laxa [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Chelinospora vulgata [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Coronaspora inornata [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Cristatisporites streelii [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin II Formation

Flag of Brazil.svg  Brazil
Flag of Tunisia.svg  Tunisia

A trilete spore.

Cymbosporites variabilis [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Cymbosporites variegatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin II Formation

Flag of Brazil.svg  Brazil
Flag of Libya.svg  Libya
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia
Flag of the United Kingdom.svg  United Kingdom

A trilete spore.

Cymbosporites wellmanii [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Cyrtospora tumida [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Awaynat Wanin III Formation

Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

Diaphanospora milleri [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Ouan Kasa Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Dibolisporites tuberculatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Jauf Formation
Jubah Formation

Flag of Brazil.svg  Brazil
Flag of Libya.svg  Libya
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Spain.svg  Spain [50]
Flag of Tunisia.svg  Tunisia
Flag of the United States.svg  United States

A trilete spore.

Dibolisporites verecundus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Dictyotriletes hemeri [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jubah Formation

Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Spain.svg  Spain

A trilete spore.

Dictyotriletes marshallii [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Ouan Kasa Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Elenisporis gondwanensis [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Jubah Formation

Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

Emphanisporites laticostatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin III Formation
Jubah Formation

Flag of Brazil.svg  Brazil
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

Emphanisporites plicatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Jubah Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Franhueberia [51]

Gen. et sp. nov.

Valid

Hoffman & Tomescu

Devonian (Emsian)

Battery Point Formation

Flag of Canada (Pantone).svg  Canada

A basal member of Euphyllophytina. The type specimen is F. gerriennei.

Geminospora convoluta [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin II Formation
Jubah Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Grandispora maura [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Awaynat Wanin III Formation

Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

Granulatisporites concavus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Jauf Formation
Jubah Formation

Flag of Libya.svg  Libya
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Spain.svg  Spain [50]

A trilete spore.

Hystricosporites brevispinus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin II Formation

Flag of Tunisia.svg  Tunisia

A trilete spore.

Raistrickia commutata [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin II Formation
Awaynat Wanin III Formation

Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

Raistrickia jaufensis [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Retusotriletes atratus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Jubah Formation

Flag of Australia (converted).svg  Australia
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Spain.svg  Spain [50]

A trilete spore.

Retusotriletes celatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Samarisporites tunisiensis [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Ouan Kasa Formation

Flag of Tunisia.svg  Tunisia

A trilete spore.

Verrucosisporites nafudensis [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Verrucosisporites onustus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Verrucosisporites stictus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Ouan Kasa Formation

Flag of Saudi Arabia.svg  Saudi Arabia

A trilete spore.

Zonotriletes brevivelatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Jauf Formation
Jubah Formation
Ouan Kasa Formation

Flag of Algeria.svg  Algeria
Flag of Libya.svg  Libya
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

Zonotriletes rotundus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Awaynat Wanin I Formation
Awaynat Wanin II Formation
Ouan Kasa Formation

Flag of Brazil.svg  Brazil
Flag of Tunisia.svg  Tunisia

A trilete spore.

Zonotriletes venatus [49]

Sp. nov

Valid

Breuer & Steemans

Devonian

Jauf Formation
Ouan Kasa Formation

Flag of Algeria.svg  Algeria
Flag of Brazil.svg  Brazil
Flag of Saudi Arabia.svg  Saudi Arabia
Flag of Tunisia.svg  Tunisia

A trilete spore.

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