2012 in arthropod paleontology

List of years in arthropod paleontology
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This list of fossil arthropods described in 2012 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2012. The list only includes taxa at the level of genus or species.

Arachnids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Ampezzoa [1]	Gen. et sp. nov.	Valid	Lindquist & Grimaldi in Schmidt et al.	Carnian		Italy	A mite. The type species is Ampezzoa triassica.
Archaelagonops [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Lagonomegopidae. Type species is A. salticoides.
Baltleucauge propinqua [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A long-jawed orb weaver
Craspedisia yapchoontecki [4]	Sp. nov.	Valid	Penney et al.	Miocene	Dominican amber	Dominican Republic	A theridiid
Eodoter lonimammillae [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A sac spider
Eohalinobius calefactus [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A succinomid
Eohalinobius hiddenseeensis [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A succinomid
Eohalinobius patina [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A succinomid
?Eophantes seorsum [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A linyphiid
Ephalmator tredecim [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	An ephalmatorid
Ero (Succinero) clunis [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A pirate spider
Ero (Succinero) gracilitibialis [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A pirate spider
Ero (Succinero) veta [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A pirate spider
Geratonephila [5]	Gen. et sp. nov.	jr synonym	Poinar in Poinar & Buckley	Early Cretaceous	Burmese amber	Myanmar	A nephiline. The type species is G. burmanica. Moved to Nephila in 2015.	Nephila burmanica
Graea magnocoli [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	An orb-weaver spider
Hypotheridiosoma [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	Originally classified as a ray spider, but subsequently transferred to the araneoid family Zarqaraneidae. [6] The type species is H. paracymbium.
Korearachne [7]	gen et sp nov	Valid	Selden et al..	Early Cretaceous	Jinju Formation	South Korea	A member of Araneomorphae, possibly a lycosoid. The type species is K. jinju.
Lagonoburmops [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Lagonomegopidae. The type species is L. plumosus.
Leclercera longissipes [2]	Sp. nov	Jr synonym	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Psilodercidae. Originally described as a species of Leclercera , but subsequently transferred to the genus Priscaleclercera. [8]
Leclercera spicula [2]	Sp. nov	Jr synonym	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Psilodercidae. Originally described as a species of Leclercera , but subsequently transferred to the genus Priscaleclercera. [8]
Leviunguis [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	Originally classified as a ray spider, subsequently transferred to the separate araneoid family Leviunguidae. [6] The type species is L. bruckschi.
Mesobunus dunlopi [9]	Sp. nov.	Valid	Giribet et al.	Middle Jurassic	Daohugou Beds	China	A sclerosomatid harvestman.
Myanlagonops [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Lagonomegopidae. The type species is M. gracilipes.
Ocululoborus [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Uloboridae. The type species is O. curvatus.
Orchestina (Baltorchestina) angulata [10]	Nom. nov	Valid	Wunderlich	Eocene	Bitterfeld amber	Germany	An oonopid; a replacement name for Orchestina (Baltorchestina) rectangulata Wunderlich (2011).
Orchestina gappi [11]	sp nov	Valid	Saupe et al..	Latest Albian or earliest Cenomanian	Chartense amber	France	An oonopid
Orchestina rabagensis [11]	Species	Valid	Saupe et al.	Early Albian		Spain	An oonopid, a species of Orchestina.
Palaeogrosphus jacquesi [12]	Sp. nov	Valid	Lourenço & Henderickx	Holocene	Malagasy copal	Madagascar	A buthid scorpion.
Palaeoleptoneta [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	Originally described as a member of Leptonetidae, subsequently considered to be a member of Araneomorphae of uncertain affinities by Magalhaes et al.. [13] The type species is P. calcar.
Palaeolychas weitschati [14]	Sp. nov.	Valid	Lourenço	Eocene		Russia (Kaliningrad Oblast)	A buthid scorpion, a species of Palaeolychas .
Palaeotrilineatus [15]	Gen. et sp. nov.	Valid	Lourenço	Early Cretaceous	Burmese amber	Myanmar	A scorpion. The type species is P. ellenbergeri.
?Pholcochyrocer baculum [2]	Sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Pholcochyroceridae, possibly a species of Pholcochyrocer .
Pholcochyrocer pecten [2]	Sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Pholcochyroceridae, a species of Pholcochyrocer .
Piankhi [16]	Gen. et sp. nov.	Valid	Dunlop, Bartel & Mitov	Eocene	Baltic amber	Europe	A harvestman found in Baltic amber. The type species is P. steineri.	Piankhi steineri
?Poliochera cretacea [17]	Sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Ricinulei, possibly a species of Poliochera .
Praoppiella [18]	Gen. et sp. nov.	Valid	Miko & Mourek in Miko et al.	Early Pleistocene		Slovenia	A mite. The type species is P. oanae.
Pseudogarypus synchrotron [19]	Sp. nov	Valid	Henderickx in Henderickx, Tafforeau & Soriano	Late Eocene	Baltic Amber	Europe	A pseudoscorpion	Pseudogarypus synchrotron
?Psiloderces filiformis [2]	Sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	First described as possibly species of Psiloderces ; transferred to the genus Eopsiloderces. [8]
Rhinoppioides [18]	Gen. et sp. nov.	Valid	Miko in Miko et al.	Early Pleistocene		Slovenia	A mite. The type species is R. quadrituberculatus.
Saetosoma [2]	Gen. et sp. nov	Valid	Wunderlich	Cretaceous	Burmese amber	Myanmar	A member of Tetrablemmidae. The type species is S. filiembolus.
Samlandicmeta [3]	Gen. et sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A long-jawed orb weaver. The type species is S. mutila.
?Scytodes hani [2]	Sp. nov	Valid	Wunderlich	Cretaceous	Lebanese amber	Jordan	A spider of uncertain affinities. Originally described as a spitting spider, possibly a species of Scytodes ; Magalhaes et al. subsequently argued that it cannot be reliably assigned to Scytodoidea. [13]
Succiniropsis runcinata [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A zoropsid.
?Succinomus gibbosus [3]	Sp. nov	Valid	Wunderlich	Eocene	Baltic amber	Europe	A succinomid, possibly a species of Succinomus .
Trhypochthonius lopezvallei [20]	Sp. nov.	Valid	Arillo, Subías & Shtanchaeva	Early Cretaceous (Albian)		Spain	A trhypochthoniid mite.
Triasacarus [1]	Gen. et sp. nov.	Valid	Lindquist & Grimaldi in Schmidt et al.	Carnian		Italy	A mite. The type species is T. fedelei.

Crustaceans

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Acanthocythereis inouei [21]	Sp. nov	Valid	Yamaguchi, Mashiba & Kamiya	Miocene	Osaki Formation	Japan	An ostracod, a species of Acanthocythereis.
Acanthophoenicides [22]	Gen. et sp. nov	Valid	Audo & Charbonnier	Late Cretaceous (Cenomanian)	Hadjoula Lagerstätte	Lebanon	A slipper lobster belonging to the subfamily Arctidinae. The type species is A. peterpani.
Acratia candyae [23]	Sp. nov	Valid	Forel	Late Permian (Changhsingian)	Wujiaping Formation	China	An acratiid ostracod, a species of Acratia .
Agnocarcinus [24]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Pilumnidae. The type species is A. zannatoi.
Alberticarcinus [24]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Atelecyclidae. The type species is A. eocaenus.
Ammopylocheles robertboreki [25]	Sp. nov	Valid	Fraaije et al.	Middle Oxfordian		Poland	A pylochelid, a species of Ammopylocheles .
Amphicytherura grandicribra [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A schizocytherid ostracod, a species of Amphicytherura .
Amphicytherura occulta [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	An ostracod. Originally described as a species of Amphicytherura ; Antonietto et al. (2013) transferred this species to the genus Dinglecythere. [27]
Amphissites gifuensis [28]	Sp. nov	Valid	Tanaka & Yuan in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Amphissites.
Antonioranina [29]	Gen. et comb. nov	Valid	Van Bakel et al.	Eocene		Italy   Spain   United States	A raninid crab. The type species is "Cyrtorhina" globosa Beschin, Busulini, De Angeli & Tessier (1988); genus also contains A. fusseli (Blow & Manning, 1996) and A. ripacurtae (Artal & Castillo, 2005). Van Bakel et al. (2012) tentatively assigned "Cyrtorhina" oblonga Beschin, Busulini, De Angeli & Tessier (1988) to the genus Antonioranina as well; [29] however, Karasawa et al. (2014) made it the type species of a separate genus Claudioranina . [30]
Archaeoniscus coreaensis [31]	Sp. nov	Valid	Park et al.	Early Cretaceous	Jinju Formation	South Korea	An isopod crustacean, a species of Archaeoniscus .
Asciocythere cabbagehillensis [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A cytherideid ostracod, a species of Asciocythere .
Asymmetricythere semoupathei [32]	Sp. nov	Valid	Sarr	Ypresian to Lutetian		Mauritania   Senegal	A trachyleberidid ostracod, a species of Asymmetricythere.
Aurikirkbya miyakei [28]	Sp. nov	Valid	Tanaka & Ono in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Aurikirkbya.
Aysegulina chapeltonensis [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A hemicytherid ostracod, a species of Aysegulina .
Aysegulina riominhoensis [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A hemicytherid ostracod, a species of Aysegulina .
Aysegulina sagitta [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A hemicytherid ostracod, a species of Aysegulina .
Aysegulina ventrocurva [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A hemicytherid ostracod, a species of Aysegulina .
Bairdia adelineae [23]	Sp. nov	Valid	Forel	Early Triassic (Griesbachian)	Daye Formation	China	A bairdiid ostracod, a species of Bairdia .
Bairdia fujisan [28]	Sp. nov	Valid	Tanaka & Maeda in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Bairdia.
Bairdia? huberti [23]	Sp. nov	Valid	Forel	Late Permian (Changhsingian)	Wujiaping Formation	China	A bairdiid ostracod, a possible species of Bairdia .
Bairdia ikeyai [28]	Sp. nov	Valid	Tanaka in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Bairdia.
Bairdia jeromei [23]	Sp. nov	Valid	Forel	Late Permian (Changhsingian) and Early Triassic (Griesbachian)	Daye Formation Wujiaping Formation	China	A bairdiid ostracod, a species of Bairdia .
Bairdia minoensis [28]	Sp. nov	Valid	Tanaka & Ono in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Bairdia.
Bairdia spindlica [28]	Sp. nov	Valid	Tanaka & Yuan in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Bairdia.
Balowella crassicostata [33]	Sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod, a species of Balowella.
Bathypluma pliocenica [34]	Sp. nov	Valid	Garassino, Pasini & Marini	Early Pliocene		Italy	A retroplumid crab, a species of Bathypluma.
Bournelyreidus [29]	Gen. et comb. et sp. nov	Valid	Van Bakel et al.	Late Cretaceous to Paleocene		Denmark   France   Germany   Greenland   United States	A lyreidid crab. The type species is "Hemioon" eysunesensis Collins & Wienberg Rasmussen (1992); according to the authors of its description genus also contains "Raninella" carlilensis Feldmann & Maxey (1980), "Palaeocorystes" laevis Schlüter in von der Marck & Schlüter (1868), "Raninella" manningi Bishop & Williams (2000), "Raninella" oaheensis Bishop (1978) and "Raninella" tridens Roberts (1962), as well as the new species Bournelyreidus teodorii. [29] On the other hand, Karasawa et al. (2014) transferred B. teodorii to the genus Lyreidus , "R." tridens to the genus Macroacaena and "R." manningi to the genus Heus . [30]
Brachycythere jamaicaensis [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod, a species of Brachycythere .
Bretonia [35]	gen et comb nov	Valid	Schweitzer et al.	Cenomanian	Sables du Perche Formation	France	crab, New genus for "Xanthosia" danielae (Collins and Breton, 2009).
Buntonia bodergati [32]	Sp. nov	Valid	Sarr	Thanetian to Ypresian		Mali   Senegal	A trachyleberidid ostracod, a species of Buntonia.
Buntonia nana [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod, a species of Buntonia .
Buntonia vulgaris [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod, a species of Buntonia .
Bythocypris antoniettae [36]	Sp. nov	Valid	Sciuto	Early Pleistocene		Italy	An ostracod, a species of Bythocypris .
Bythocythere solisdeus [37]	Sp. nov	Valid	Sciuto	Early Pleistocene		Italy	A bythocytherid ostracod, a species of Bythocythere .
Calliaxina altavillensis [38]	Sp. nov	Valid	Beschin & De Angeli	Eocene		Italy	A member of Callianassidae.
Carinokloedenia onusta carinata [39]	Subsp. nov	Valid	Becker & Franke	Devonian		Germany   Luxembourg	An ostracod belonging to the group Palaeocopida and the family Beyrichiidae.
Carcinoplax parsensis [40]	Sp. nov	Valid	Vega et al.	Miocene (Burdigalian or Langhian)	Mishan Formation	Iran	A goneplacid decapod crustacean, a species of Carcinoplax .
Carpopenaeus garassinoi [41]	Sp. nov	Valid	Charbonnier in Petit & Charbonnier	Late Cretaceous	Sahel Alma Lagerstätte	Lebanon
Cenocorystes bretoni [29]	Sp. nov	Valid	Van Bakel et al.	Late Cretaceous (late Cenomanian)		France	A palaeocorystid crab, a species of Cenocorystes.
Cenomanocarcinus cantabricus [29]	Sp. nov	Valid	Van Bakel et al.	Early Cretaceous (Albian)		Spain	A cenomanocarcinid crab, a species of Cenomanocarcinus.
Coleia martinlutheri [42]	sp nov	Valid	Schweigert & Ernst	Sinemurian		Germany	A coleiid polychelid, a species of Coleia.
Cretacolana [43]	Gen. et comb. nov	Valid	Schweitzer & Feldmann	Cenomanian		France	A porcelain crab, a new genus for "Porcellana" antiqua A. Milne-Edwards, 1862.
Cretatrizocheles [44]	gen et sp nov	Valid	Fraaije, Klompmaker, & Artal	Albian or Cenomanian		Spain	A hermit crab. The type species is Cretatrizocheles olazagutiensis.
Cromimaia [24]	Gen. et comb. nov	Valid	Beschin et al.	Eocene		Italy	A crab belonging to the family Majidae. The type species is "Micromaia" meneguzzoi Beschin, Busulini, De Angeli & Tessier (1985); genus also includes "Micromaia" elegans Beschin, Busulini, De Angeli & Tessier (1985).
Cushmanidea tessieri [32]	Sp. nov	Valid	Sarr	Ypresian to Lutetian		Senegal	A cushmanideid ostracod, a species of Buntonia.
Cytherelloidea guineacornensis [26]	Sp. nov	Valid	Puckett in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)	Guinea Corn Formation	Jamaica	A cytherellid ostracod, a species of Cytherelloidea .
Cytheropteron eleonorae [37]	Sp. nov	Valid	Sciuto	Early Pleistocene		Italy	A cytherurid ostracod, a species of Cytheropteron .
Cytheropteron italoi [36]	Sp. nov	Valid	Sciuto	Early Pleistocene		Italy	An ostracod, a species of Cytheropteron .
Cytheropteron rossanae [36]	Sp. nov	Valid	Sciuto	Early Pleistocene		Italy	An ostracod, a species of Cytheropteron .
Daira coccoi [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Dairidae; a species of Daira .
Dinochelus steeplensis [45]	Sp. nov	Valid	Tshudy & Saward	Eocene (Ypresian)	London Clay	United Kingdom	A lobster, a species of Dinochelus .
Distefania renefraaijei [46]	Sp. nov	Valid	Klompmaker, Feldmann & Schweitzer	Cretaceous (Albian or Cenomanian)		Spain	A goniodromitid crab, a species of Distefania.
Dolocythere tuberculata [33]	Sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod. Originally described as a species of Dolocythere ; Lord, Cabral & Danielopol (2020) transferred this species to the genus Minyocythere. [47]
Ebullitiocaris elatus [48]	Sp. nov	Valid	Womack et al.	Carboniferous		United Kingdom	A member of Cladocera, a species of Ebullitiocaris.
Eocytheropteron hazeli [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A cytherurid ostracod, a species of Eocytheropteron .
Eocytheropteron jamaicaensis [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A cytherurid ostracod, a species of Eocytheropteron .
Eouroptychus [49]	Gen. et sp. nov	Valid	De Angeli & Ceccon	Eocene (Ypresian)		Italy	A member of Chirostylidae. The type species is E. montemagrensis.
Eucytherura mitchelli [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A cytherurid ostracod, a species of Eucytherura .
Eomunidopsis aldoirarensis [50]	Sp. nov	Valid	Klompmaker et al.	Cretaceous (Albian or Cenomanian)		Spain	A galatheoid, a species of Eomunidopsis.
Eopaguropsis nidiaquilae [51]	Sp. nov	Valid	Fraaije et al.	Oxfordian		Poland	A diogenid, a species of Eopaguropsis .
Eosymethis [29]	Gen. et sp. nov	Valid	Van Bakel et al.	Eocene (Ypresian)	Roda Formation	Spain	A raninid crab. The type species is Eosymethis aragonensis.
Eohexapus orthogonius [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Hexapodidae.
Eucorystes iserbyti [29]	Sp. nov	Valid	Van Bakel et al.	Early Cretaceous (Albian)		France	A palaeocorystid crab, a species of Eucorystes.
Eucorystes navarrensis [29]	Sp. nov	Valid	Van Bakel et al.	Early Cretaceous (Albian)		Spain	A palaeocorystid crab, a species of Eucorystes.
Faksecarcinus [35]	gen et comb nov	Valid	Schweitzer et al.	Albian or Cenomanian to Danian		Denmark   Spain	A crab, a new genus for "Xanthosia" gracilis.
Ferroranina [29]	Gen. et comb. et sp. nov	Valid	Van Bakel et al.	Late Cretaceous (late Cenomanian to late Campanian)		Czech Republic [52]   India   Madagascar   Mexico   United States	A palaeocorystid crab. The type species is "Notopocorystes" dichrous Stenzel (1945); genus also contains "Notopocorystes" australis Secretan (1964) and "Notopocorystes" fritschi Glaessner (1929), [52] as well as the new species Ferroranina tamilnadu.
Floricythereis [26]	Gen. et sp. et comb. nov		Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica   United States	A trachyleberidid ostracod. Genus includes new species Floricythereis exquisita, as well as F. lixula (Crane, 1965) (the type species was not designated).
Galathea mainensis [53]	Sp. nov	Valid	Ceccon & De Angeli	Middle Eocene		Italy	A species of Galathea .
Gebiacantha tuscia [54]	Sp. nov	Valid	Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.	Early Pliocene		Italy	An upogebiid decapod, a species of Gebiacantha .
Geffenina mariebeatriceae [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Late Early Permian	Tak Fa Formation	Thailand	A knoxitid palaeocopidan ostracod, a species of Geffenina.
Geffenina mariebeatriceae [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Early Permian	Pha Nok Khao Formation	Thailand	A knoxitid palaeocopidan ostracod, a species of Geffenina.
Glyphea wiffenae [56]	Species	Valid	Feldmann and Stilwell	Late Cretaceous	Maungataniwa Sandstone	New Zealand	A glypheid lobster, a species of Glyphea .
Glyptopleura mugiensis [28]	Sp. nov	Valid	Tanaka & Yuan in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Glyptopleura.
Goniodromites kubai [57]	sp nov	Valid	Starzyk, Krzemiska, & Krzemiski	Oxfordian		Poland	A crab, a species of Goniodromites.
Gonioplacoides [58]	Gen. et sp. nov	Valid	Quayle & Collins	Eocene	Headon Hill Formation	United Kingdom	A goneplacid. The type species is Gonioplacoides minuta.
Graptocarcinus urbasaensis [59]	Sp. nov	Valid	Van Bakel et al.	Late Cretaceous		Spain	A dynomenid crab, a species of Graptocarcinus.
Grinioneis pachycosta [60]	Sp. nov	Valid	Guernet et al.	Eocene (Lutetian)	Paris Basin	France	A trachyleberidid ostracod, a species of Grinioneis.
Grolamaia [24]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the subfamily Pisinae. The type species is G. vicariottoi.
Hadrocarcinus [61]	gen et comb et sp nov	Valid	Schweitzer, Feldmann & Lamanna	Late Cretaceous		Antarctica	A necrocarcinid crab, a new genus for "Necrocarcinus" wrighti. The genus also contains "Necrocarcinus" carinatus and a new species Hadrocarcinus tectilacus.
Harbinia alta [62]	Sp. nov	Valid	Antonietto et al.	Aptian or early Albian	Santana Formation	Brazil	An ostracod crustacean, a species of Harbinia .
Headonipus [58]	Gen. et sp. nov	Valid	Quayle & Collins	Late Eocene	Headon Beds	United Kingdom	A hexapodid. The type species is Headonipus tuberculatus.
Hemiparacytheridea exquisita [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A paracytherideid ostracod, a species of Hemiparacytheridea .
Hispanigalathea [50]	Gen. et 2 sp. nov	Valid	Klompmaker et al.	Cretaceous (Albian or Cenomanian)		Spain	A galatheoid. Genus contains two species: Hispanigalathea pseudolaevis and H. tuberosa.
Huanghestheria linxiensis [63]	Sp. nov	Valid	Niu in Zhang et al.	Late Permian	Linxi Formation	China	A vertexiine lioestheriid spinicaudatan clam shrimp, a species of Huanghestheria.
Joeranina [29]	Gen. et comb. et sp. nov	Valid	Van Bakel et al.	Cretaceous (Albian to early Campanian)		Canada   France   Greenland   Japan   Madagascar   Spain    Switzerland   Syria   United Kingdom   United States	A palaeocorystid crab. The type species is "Corystes" broderipii Mantell (1844). According to the authors of its description genus also contains "Palaeocorystes" harveyi Woodward (1896), "Eucorystes" japonicus Jimbô (1894), "Notopocorystes" paututensis Collins & Wienberg Rasmussen (1992), "Eucorystes" platys Schweitzer & Feldmann (2002) and "Notopocorystes" syriacus Withers (1928), as well as the new species Joeranina gaspari. [29] On the other hand, Karasawa et al. (2014) transferred "P." harveyi, E. platys and "N." paututensis to the genus Eucorystes; the authors also transferred the species "Notopocorystes" xizangensis Wang (1981) to the genus Joeranina. [30]
Juracrista [64]	Gen. et 2 sp. nov	Valid	Robins, Feldmann & Schweitzer	Late Jurassic (Tithonian)	Ernstbrunn Limestone	Austria	A munidid. The type species is Juracrista perculta; genus also contains Juracrista costaspinosa.
Jurapylocheles iwonae [25]	Sp. nov	Valid	Fraaije et al.	Middle Oxfordian		Poland	A pylochelid, a species of Jurapylocheles .
Kinkelinella circumcostata [33]	Sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod, a species of Kinkelinella.
Kloedenella poschmanni [39]	Sp. nov	Valid	Becker & Franke	Devonian		Germany	An ostracod.
Koryncheiros [65]	Gen. et sp. nov	Valid	Feldmann et al.	Middle or late Anisian	Guanling Formation	China	A decapod crustacean; a clytiopsid erymoid. The type species is Koryncheiros luopingensis.
Kroemmelbeincypris [66]	Gen. et comb. nov	Valid	Poropat & Colin	Aptian			An ostracod crustacean, a new genus for the species "Hourcqia" symmetrica Krömmelbein & Weber and "Hourcqia" angulata Krömmelbein & Weber.
Langdaia meesooki [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Late Early to Middle Permian	Tak Fa Formation	Thailand	A knoxitid palaeocopidan ostracod, a species of Langdaia.
Leiocythereis [26]	Gen. et sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod. The type species is Leiocythereis polita.
Lepas sattmanni [67]	Sp. nov	Valid	Harzhauser & Schlögl	Early Miocene	Ebelsberg Formation	Austria	A lepadid barnacle, a species of Lepas .
Lessinicarcinus [68]	Gen. et comb. nov	Valid	De Angeli	Eocene		Croatia [69]   Italy	A crab belonging to the family Pilumnidae. The type species is "Titanocarcinus" euglyphos Bittner (1875).
Lessinipagurus [24]	Gen. et sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A hermit crab. Originally assigned to the family Paguridae, [24] subsequently transferred to the separate family Xylopaguridae by Gašparič et al. (2016). [70] The type species is L. granulatus.
Lienenklausicythere [71]	Gen. et comb. nov	Valid	Uffenorde, Monostori & Tóth	Eocene to Oligocene		Europe	An ostracod, a new genus for "Hermanites" acuticosta and "Hermanites" camela; genus also contains a new, yet-unnamed species.
Ljubimovella debilicostatus [33]	Sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod, a species of Ljubimovella.
Lobonotus beschini [72]	Sp. nov	Valid	De Angeli & Checchi	Eocene (Lutetian)		Italy	A crab belonging to the family Tumidocarcinidae.
Loxoconcha pindarsensis [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A loxoconchid ostracod, a species of Loxoconcha .
Marylyreidus [29]	Gen. et comb. nov	Valid	Van Bakel et al.	Cretaceous (late Albian–early Cenomanian)	Pawpaw Formation	United States	A lyreidid crab. The type species is "Notopocorystes" punctatus Rathbun (1935).
Megokkos patagoniensis [73]	Sp. nov	Valid	Schweitzer et al.	Eocene	Rio Turbio Formation	Argentina	A crab, a species of Megokkos.
Mesoparapylocheles [44]	gen et sp nov	Valid	Fraaije, Klompmaker, & Artal	Albian or Cenomanian		Spain	A hermit crab. The type species is Mesoparapylocheles michaeljacksoni .
Microcheilinella subovata [28]	Sp. nov	Valid	Tanaka & Maeda in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Microcheilinella.
Microcheilinella tsubogawensis [28]	Sp. nov	Valid	Tanaka & Ichida in Tanaka et al.	Early Permian		Japan	An ostracod, a species of Microcheilinella.
Microcoelonella takfaensis [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Middle Permian	Tak Fa Formation	Thailand	A coelonellid palaeocopidan ostracod, a species of Microcoelonella.
Microcoelonella takliensis [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Middle Permian	Tak Fa Formation	Thailand	A coelonellid palaeocopidan ostracod, a species of Microcoelonella.
Munida deangelii [74]	Sp. nov	Valid	Robins, Feldmann & Schweitzer	Late Oligocene to early Miocene	Río Foyel Formation	Argentina	A munidid, a species of Munida .
Myra gurii [40]	Sp. nov	Valid	Vega et al.	Miocene (Burdigalian or Langhian)	Mishan Formation	Iran	A leucosiid decapod crustacean, a species of Myra .
Navarradromites [46]	Gen. et sp. nov	Valid	Klompmaker, Feldmann & Schweitzer	Cretaceous (Albian or Cenomanian)		Spain	A goniodromitid crab. The type species is Navarradromites pedroartali.
Navarrahomola [75]	Gen. et sp. nov.	Valid	Artal et al.	Cretaceous (Albian or Cenomanian)		Spain	A homolid crab. The type species is Navarrahomola hispanica.
Neocallichirus bacatus [76]	Sp. nov	Valid	Busulini, Beschin & Tessier	Eocene (Priabonian)		Italy	A member of Callianassidae.
Neocytherideis labyrinthoidea [60]	Sp. nov	Valid	Guernet et al.	Eocene (Lutetian)	Paris Basin	France	A neocytherideid ostracod, a species of Neocytherideis.
Neocytheromorpha priscipacifica [21]	Sp. nov	Valid	Yamaguchi, Mashiba & Kamiya	Miocene	Osaki Formation	Japan	An ostracod, a species of Neocytheromorpha.
Neurocythere jakovlevae [77]	Sp. nov	Valid	Kolpenskaya in Tesakova et al.	Late Jurassic (Early Kimmeridgian)		Russia	An ostracod, a species of Neurocythere.
Nitotacarcinus antipodes [73]	Sp. nov	Valid	Schweitzer et al.	Eocene	Rio Turbio Formation	Argentina	A crab, a species of Nitotacarcinus.
Nodosculda [78]	Gen. et sp. nov	Valid	Franţescu	Early Cretaceous (late Albian)		United States	A mantis shrimp. The type species is Nodosculda fisherorum.
Noetlingocarcinus messinai [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the superfamily Dromioidea and the family Basinotopidae, a species of Noetlingocarcinus .
Nogarhomola [79]	Gen. et sp. nov	Valid	De Angeli & Alberti	Eocene (Lutetian)		Italy	A homolid crab. The type species is Nogarhomola aurorae.
Notopocorystes kerri [80]	Sp. nov	Valid	Luque et al.	Early Cretaceous (late Aptian)	Paja Formation	Colombia	A palaeocorystid raninoid crab. Initially described as a species of Notopocorystes , [80] but subsequently reassigned to the genus Joeranina . [81]
Nykteripteryx [50]	Gen. et sp. nov	Valid	Klompmaker et al.	Cretaceous (Albian or Cenomanian)		Spain	A galatheoid. The type species is Nykteripteryx rostrata.
Ochtholambrus gambillarai [76]	Sp. nov	Valid	Busulini, Beschin & Tessier	Eocene (Priabonian)		Italy	A crab belonging to the family Parthenopidae.
Odontodactylus italicus [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A mantis shrimp belonging to the family Odontodactylidae; a species of Odontodactylus .
Ommaciria [24]	Gen. et comb. nov	Valid	Beschin et al.	Eocene		Italy	A crab belonging to the family Majidae. The type species is "Micromaia" mainensis Beschin, Busulini, De Angeli & Tessier (1985).
Orthobairdia jeanlouisi [23]	Sp. nov	Valid	Forel	Early Triassic (Griesbachian)	Daye Formation	China	A bairdiid ostracod, a species of Orthobairdia .
Ovocytheridea rotunda [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A cytherideid ostracod, a species of Ovocytheridea .
Paguristes gagnaisoni [82]	Sp. nov	Valid	Gagnaison	Miocene		France	A species of Paguristes .
Pagurus valdagnensis [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A hermit crab belonging to the family Paguridae, a species of Pagurus .
Palaega picena [83]	Sp. nov	Valid	Pasini & Garassino	Miocene		Italy	A cirolanid isopod, a species of Palaega.
Palaega pisana [84]	Sp. nov	Junior synonym	Pasini & Garassino	Pliocene		Italy	A cirolanid isopod. Originally described as a species of Palaega; considered to be a junior synonym of Bathynomus sismondai (Ristori, 1891) by Hyžný, Pasini & Garassino (2019). [85]
Palaega steatopigia [86]	Sp. nov	Nomen dubium	Pasini & Garassino	Pliocene		Italy	A cirolanid isopod. Originally described as a species of Palaega ; Hyžný, Pasini & Garassino (2019) considered this species to be a nomen dubium and reassigned it to the genus Bathynomus . [85]
Palaeohomarus pacificus [87]	Sp. nov	Valid	Aguirre-Urreta, Lazo & Rawson	Late Hauterivian or early Barremian	Agrio Formation	Argentina	A decapod crustacean, a species of Palaeohomarus.
Paracypris keiji [60]	Sp. nov	Valid	Guernet et al.	Eocene (Lutetian)	Paris Basin	France	A candonid ostracod, a species of Paracypris.
Paranecrocarcinus balla [29]	Sp. nov	Valid	Van Bakel et al.	Late Cretaceous (Cenomanian)		France	A necrocarcinid crab, a species of Paranecrocarcinus.
Periosocypris [88]	Gen. et sp. nov	Valid	Whatley, Khosla & Rathore	Late Cretaceous (Maastrichtian)	Lameta Formation	India	A cypridid ostracod. The type species is Periosocypris megistus.
Petrochirus savii [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A hermit crab belonging to the family Diogenidae, a species of Petrochirus .
Phrixocythere [26]	Gen. et sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A hemicytherid ostracod. The type species is Phrixocythere unionensis.
Planocarcinus [80]	Gen. et comb. nov	Valid	Luque et al.	Early Cretaceous (late Aptian)	Paja Formation	Colombia	A necrocarcinid raninoid crab, a new genus for "Dakoticancer" olssoni Rathbun (1937).
Podocallichirus laepaensis [89]	Sp. nov	Valid	Hyžný & Muñiz	Late Miocene		Spain	A callianassid ghost shrimp. Originally described as a species of Podocallichirus ; subsequently transferred to the genus Balsscallichirus by Hyžný (2016). [90]
Protobuntonia warorei [32]	Sp. nov	Valid	Sarr	Paleocene to Ypresian		Senegal   Togo	A trachyleberidid ostracod, a species of Buntonia.
Pterygocythereis babinoti [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod, a species of Pterygocythereis .
Pylochelitergites [91]	Gen. et 2 sp. nov	Valid	Fraaije et al.	Oxfordian to Kimmeridgian		Germany   Poland	A pylochelid hermit crab. Genus contains two species: P. westerbergensis (the type species) and P. gelasinus.
Quasilaeviranina minuscula [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Raninidae, a species of Quasilaeviranina .
Raninoides rioturbiensis [73]	Sp. nov	Valid	Schweitzer et al.	Eocene	Rio Turbio Formation	Argentina	A crab, a species of Raninoides.
Rhinolambrus elongatus [24]	Sp. nov	Valid	Beschin et al.	Eocene (Lutetian)		Italy	A crab belonging to the family Parthenopidae; a species of Rhinolambrus .
Ristoria [54]	Gen. et comb. nov	Valid	Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.	Early Pliocene		Italy	A leucosiid crab, a new genus for "Ilia" pliocaenica Ristori, 1891.
Samarella sonei [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Middle Permian	Tak Fa Formation	Thailand	A paraparchitid palaeocopidan ostracod, a species of Samarella.
Samarella viscusforma [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Early to Middle Permian	Pha Nok Khao Formation Tak Fa Formation	Thailand	A paraparchitid palaeocopidan ostracod, a species of Samarella.
Sargentina chantarameei [55]	Sp. nov	Valid	Chitnarin in Chitnarin et al.	Early to Middle Permian	Pha Nok Khao Formation Tak Fa Formation	Thailand	A knoxitid palaeocopidan ostracod, a species of Sargentina.
Scalpellum paratethyianum [67]	Sp. nov	Valid	Harzhauser & Schlögl	Early Miocene	Lakšárska Nová Ves Formation	Slovakia	A scalpellid barnacle, a species of Scalpellum .
Schizoptocythere nana [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod, a species of Schizoptocythere .
Schizoptocythere pinna [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A trachyleberidid ostracod, a species of Schizoptocythere .
Schuleridea exigua [26]	Sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica	A cytherideid ostracod, a species of Schuleridea .
Semicytherura carioui [32]	Sp. nov	Valid	Sarr	Ypresian to Lutetian		Senegal	A cytherurid ostracod, a species of Semicytherura.
Sineruga [92]	gen et sp nov	Valid	Perrier	Silurian (uppermost Ludlow to lower Pridoli)	Le Val Formation	France	myodocope ostracod. type species is Sineruga insolita.
Southcavea (Saxosouthcavea) [33]	Subgen. et 3 sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod, a subgenus of Southcavea containing three new species: Southcavea (Saxosouthcavea) extendata, Southcavea (Saxosouthcavea) bartenbrandi and Southcavea (Saxosouthcavea) malzi (with two subspecies: S. (S.) malzi malzi and S. (S.) malzi reticulata).
Spinicytheridea [26]	Gen. et sp. nov	Valid	Puckett & Colin in Puckett, Colin & Mitchell	Late Cretaceous (Maastrichtian)		Jamaica   Mexico	A cytherideid ostracod. The type species is Spinicytheridea compta from Jamaica; genus also includes a second, unnamed species from the Maastrichtian Ocozocoautla Formation of Chiapas, Mexico.
Spinirostrimaia [24]	Gen. et comb. nov	Valid	Beschin et al.	Eocene		Italy   Spain	A crab belonging to the family Majidae. The type species is "Micromaia" margaritata Fabiani (1910).
Stagmacaris adielklompmakeri [91]	Sp. nov	Valid	Fraaije et al.	Early late Kimmeridgian		Germany	A pylochelid hermit crab, a species of Stagmacaris.
Stagmacaris biarcus [91]	Sp. nov	Valid	Fraaije et al.	Middle Oxfordian		Poland	A pylochelid hermit crab, a species of Stagmacaris.
Steorrosia [35]	gen et comb nov	Valid	Schweitzer et al.	Late Albian to Cenomanian		United States	Crab, New genus for "Xanthosia" aspera. Other species S. pawpawensis and S. reidi.
Styracocarcinus [43]	Gen. et comb. nov	Valid	Schweitzer & Feldmann	Late Cretaceous (Senonian)		Morocco	A tumidocarcinid decapod, a new genus for "Titanocarcinus" meridionalis Secretan, 1961.
Superpendicythere [33]	Gen. et sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod. The type species is S. binodosa.
Symethoides [29]	Gen. et sp. nov	Valid	Van Bakel et al.	Paleocene (early Danian)	Tinton Formation	United States	A raninid crab. The type species is Symethoides monmouthorum.
Szaboa deppermanni [93]	Sp. nov	Valid	Fraaije et al.	Oligocene	Ratzeburg Formation	Germany	A matutid crab, a species of Szaboa .
Thalamita italica [54]	Sp. nov	Valid	Garassino, Pasini, De Angeli & Charbonnier in Garassino et al.	Early Pliocene		Italy	A portunid crab, a species of Thalamita .
Tridactylastacus [65]	Gen. et sp. nov	Valid	Feldmann et al.	Middle or late Anisian	Guanling Formation	China	A decapod crustacean; a glypheoid belonging to the family Litogastridae. [94] The type species is Tridactylastacus sinensis.
Turbiocheir [73]	Gen. et sp. nov	Disputed	Schweitzer et al.	Eocene	Rio Turbio Formation	Argentina   Chile   Venezuela [95]	A thalassinidean. The type species is Turbiocheir minutospinata. Hyžný et al. (2020) considered the genus Turbiocheir to be a junior synonym of the genus Anacalliax , creating a new combination Anacalliax minutospinata. [96]
Vecticallichirus [58]	Gen. et sp. nov	Valid	Quayle & Collins	Late Eocene and Oligocene [97]	Headon Beds	United Kingdom	A callichirid [97] ghost shrimp. The type species is Vecticallichirus abditus. Hyžný (2020) considered V. abditus to be a junior synonym of "Callianassa" batei Woodward (1869), resulting in a new combination Vecticallichirus batei. [97]
Vegaranina [29]	Gen. et comb. nov	Valid	Van Bakel et al.	Late Cretaceous (Campanian–Maastrichtian)		Cuba   France   Mexico   United States (  Puerto Rico)	A raninid crab. The type species is "Lophoranina" precocious Feldmann, Vega, Tucker, García-Barrera & Avendaño (1996).
Vernoniella tenuipora [33]	Sp. nov	Valid	Luppold	Middle Jurassic		Germany	An ostracod, a species of Vernoniella.
Viaia [75]	Gen. et sp. nov.	Valid	Artal et al.	Cretaceous (Albian or Cenomanian)		Spain	A viaiid glaessneropsoid crab. The type species is Viaia robusta.
Yunnanopalinura [65]	Gen. et sp. nov	Valid	Feldmann et al.	Middle or late Anisian	Guanling Formation	China	A decapod crustacean; a spiny lobster. The type species is Yunnanopalinura schrami.

Insects

Main article: 2012 in paleoentomology

Eurypterids

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Adelophthalmus dubius [98]	Sp. nov	Valid	Shpinev	Middle Devonian		Russia	A eurypterid, a species of Adelophthalmus.
Adelophthalmus kamyshtensis [98]	Sp. nov	Valid	Shpinev	Middle Devonian		Russia	A eurypterid, a species of Adelophthalmus.
Drepanopterus odontospathus [99]	Sp. nov	Valid	Lamsdell	Early Devonian		Canada	A eurypterid.
Nanahughmilleria notosibirica [100]	Sp. nov	Valid	Shpinev	Devonian		Russia	A eurypterid, a species of Nanahughmilleria .
Parahughmilleria longa [100]	Sp. nov	Valid	Shpinev	Devonian		Russia	A eurypterid, a species of Parahughmilleria .
Stylonuroides orientalis [100]	Sp. nov	Valid	Shpinev	Devonian		Russia	A eurypterid, a species of Stylonuroides .

Trilobites

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Abanomocarella [101]	Gen. et sp. et comb. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae. Genus includes new species A. quadrata, as well as A. humilis (Resser & Endo in Endo & Resser, 1937) and A. pyriformis (Resser & Endo in Endo & Resser, 1937).
Acmarhachis kindlei [102]	Sp nov	Valid	Westrop & Eoff	Late Cambrian	Cow Head Group	Canada	An agnostid, a species of Acmarhachis .
Acmarhachis whittingtoni [102]	Sp nov	Valid	Westrop & Eoff	Late Cambrian	Cow Head Group	Canada	An agnostid, a species of Acmarhachis .
Angsiduoa jiulongshanensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Ignotogregatidae.
Angsiduoa shanxiensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Ignotogregatidae.
Angsiduoa transversa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Ignotogregatidae.
Anomocarella? antiqua [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae.
Anomocarella jiulongshanensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae.
Anomocarella similis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae.
Aponileus aasei [103]	Sp nov	Valid	Adrain & McAdams	Early Ordovician (late Floian)	Fillmore Formation	United States	A bathyurid trilobite, a species of Aponileus.
Aponileus belkaae [103]	Sp nov	Valid	Adrain & McAdams	Early Ordovician (late Floian)	Wah Wah Formation	United States	A bathyurid trilobite, a species of Aponileus.
Aponileus laikaae [103]	Sp nov	Valid	Adrain & McAdams	Early Ordovician (late Floian)	Fillmore Formation	United States	A bathyurid trilobite, a species of Aponileus.
Aponileus strelkaae [103]	Sp nov	Valid	Adrain & McAdams	Early Ordovician (late Floian)	Wah Wah Formation	United States	A bathyurid trilobite, a species of Aponileus.
Aponileus ugolekae [103]	Sp nov	Valid	Adrain & McAdams	Early Ordovician (late Floian)	Wah Wah Formation	United States	A bathyurid trilobite, a species of Aponileus.
Aponileus? veterokae [103]	Sp nov	Valid	Adrain & McAdams	Early Ordovician (late Floian)	Wah Wah Formation	United States	A bathyurid trilobite, a possible species of Aponileus.
Archaeagnostus typicus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Peronopsidae.
Archegonus (Phillibole) nehdenensis africanus [104]	Subsp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Phillipsiidae belonging to the subfamily Archegoninae, a subspecies of Archegonus nehdenensis .
Asaphellus cuervoae [105]	Sp. nov	Disputed	Corbacho & López-Soriano	Ordovician (Floian)	Upper Fezouata Formation	Morocco	A member of Asaphidae, a species of Asaphellus . Gutiérrez-Marco et al. (2017) considered this species to be likely based on poorly preserved and probably retouched specimens of Asaphellus stubbsi. [106]
Astycoryphe arduinnae [107]	Sp nov	Valid	Van Viersen, van Rossum & Prescher	Middle Devonian		Belgium	A member of Proetida.
Australoscutellum iota [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Australoscutellum .
Australoscutellum microps [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Australoscutellum.
Australoscutellum pulex [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Australoscutellum.
Australoscutellum trica [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Australoscutellum.
Baltagnostus yaojiayuensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Diplagnostidae.
Baojingia (Eobaojingia) [101]	Subgen. et 2 sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Lisaniidae. The subgenus includes Baojingia (Eobaojingia) dubia and Baojingia (Eobaojingia) ezhuangensis, as well as Baojingia miaogouensis Mong in Zhou et al. (1977).
Borenoria [108]	gen et sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite. The type species is Borenoria cirrita.
Bumastoides graffhami [109]	Sp nov	Valid	Carlucci et al.	Ordovician		United States	An illaenid trilobite, a species of Bumastoides.
Bumastoides kimmswickensis [109]	Sp nov	Valid	Carlucci et al.	Ordovician		United States	An illaenid trilobite, a species of Bumastoides.
Bumastoides moundensis [109]	Sp nov	Valid	Carlucci et al.	Ordovician		United States	An illaenid trilobite, a species of Bumastoides.
Calymene harringtoni [110]	Sp. nov	Valid	Tortello in Tortello et al.	Early Silurian	Vargas Peña Formation	Paraguay	A species of Calymene .
Chlupacula (Chlupacula) erfoudensis [104]	Sp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Chlupacula.
Crepicephalina exsculpta [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Crepicephalina obscura [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Crepicephalina pulchra [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Crepicephalina? rara [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Crepicephalina woqishanensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Cummingella boikoi [111]	sp nov	Valid	Mychko	Sakmarian		Russia	A phillipsiid, a species of Cummingella.
Cyrtoproetus (Crassibole) acrops [104]	Sp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Cyrtoproetus.
Dalejeproetus owensi [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Dalejeproetus.
Dalejeproetus sagaouii [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Dalejeproetus.
Dameselloides [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Damesellidae. Genus includes new species D. sinensis.
Danjiangella shandongensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Dechenelloides tazoultensis [104]	Sp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Dechenelloides.
Decoroscutellum planes [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Decoroscutellum .
Derikaspis conicus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Derikaspis ruichengensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Diacoryphe (Archaeocoryphe) maiderensis [104]	Sp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Diacoryphe.
Dindymene whittingtoni [113]	Sp nov	Valid	Owen & Bruton	Late Ordovician		United States	A species of Dindymene.
Dorypyge cylindrica [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Dorypyge zhangxianensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Dunopyge seiberti [114]	Sp. nov	Valid	Basse & Müller	Early Devonian	Seifen Beds	Germany	A member of Acastidae belonging to the subfamily Asteropyginae, a species of Dunopyge.
Eilura trapezialis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Elaphraella? taebaeksanensis [115]	Sp. nov	Valid	Park & Choi	Cambrian (middle Furongian)	Sesong Formation	South Korea	A member of the family Shumardiidae.
Emmrichella punctata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ptychopariidae.
Eokoldinia [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Plethopeltidae. Genus includes new species E. porifera.
Eomapania [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Mapaniidae. Genus includes new species E. subquadrata.
Eoscutellum annosum [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Eoscutellum .
Eymekops pristina [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Eymekops transversa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Grandioculus macroculus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Grandioculus (Protohedina) quadraticephala [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Gumunsoia [116]	Gen. et sp. nov	Valid	Park, Sohn & Choi	Cambrian (middle Furongian)	Sesong Formation	South Korea	A member of Leiostegioidea belonging to the family Kaolishaniidae. Genus includes new species G. triangularis.
Gushanella [101]	Gen. et 2 sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Solenopleuridae. Genus includes new species G. quadrata and G. truncata.
Heyelingella crassa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Honania porifera [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Huayuanaspis angustilimbata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Idioura brevispina [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Illaenoscutellum tryo [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Illaenoscutellum .
Inouyella rara [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Inouyella shiliuyuanensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Inouyia (Bulbinouyia) [101]	Subgen. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Inouyiidae. The subgenus includes Inouyia kangzhuangensis (Zhang & Wang, 1985) and I. lubrica (Zhou in Zhou et al., 1982).
Inouyia latilimbata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Iranoleesia pustulosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Iranoleesia (Proasaphiscina) rogosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Japonoscutellum borenorense [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum .
Japonoscutellum crassum [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum diascetum [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum gephyricum [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum guttulatum [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum serangicum [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum strabo [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Japonoscutellum.
Jiulongshania shandongensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Diceratocephalidae.
Koptura (Eokoptura) [101]	Subgen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Anomocaridae. The subgenus includes new species Koptura (Eokoptura) bella, as well as Koptura quadrata (Resser & Endo in Endo & Resser, 1937).
Koptura intermedia [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Anomocaridae.
Kosovopeltis trepida [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Kosovopeltis .
Kosovopeltis xynon [108]	sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite, a species of Kosovopeltis.
Lepidoproetus lahceni [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Lepidoproetus.
Lepidoproetus maharchianus [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Lepidoproetus.
Lepidoproetus splendens [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Lepidoproetus.
Liowutaishania laevigata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Lisania elliptica [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Longiproetus meiopsifius [107]	Sp nov	Valid	Van Viersen, van Rossum & Prescher	Middle Devonian		Belgium	A member of Proetida.
Lunania [101]	Gen. nov	Junior homonym	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Menocephalidae. The generic name is preoccupied by Lunania Chow (1957).
Maotunia transversa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Mapania (Mapanella) [101]	Subgen. et 2 sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Mapaniidae. The subgenus includes new species M. (Mapanella) transversa and M. (Mapanella) prisca, as well as Mapania blackwelderi (Resser & Endo in Endo & Resser, 1937).
Mapania subcylindrica [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Mapanopsis conicus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Maurotarion megacephalum [117]	Sp. nov.	Valid	Rustán & Vaccari	Early Devonian (Pragian)	Talacasto Formation	Argentina	An otarionine aulacopleurid, a species of Maurotarion.
Megagraulos similis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Menocephalina [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Menocephalidae. Genus includes new species M. longispinosa.
Menocephalites (Hutoushania) [101]	Subgen. et 2 sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Menocephalidae. The subgenus includes Menocephalites (Hutoushania) triangulata and M. (Hutoushania) zaozhuangansis.
Menocephalites porosus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Menocephalites primitivus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Menocephalites simplex [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Metanomocarella primitiva [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Micagnostus semicircularis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Mirrabooka [108]	gen et sp nov	Valid	Holloway and Lane	Silurian		Australia	A scutelluid trilobite. The type species is Mirrabooka harryi.
Monkaspis conica [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Ohleum magreani [118]	Sp nov	Valid	Taghon, Bonino & Mottequin	Devonian (Eifelian)	Jemelle Formation	Belgium	A member of Lichida belonging to the family Lichidae.
Pagetides qianensis [119]	Sp nov	Valid	Zhang & Clarkson	Cambrian		China	An eodiscoid trilobite, a species of Pagetides.
Paleonelsonia granulosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ptychopariidae.
Paleonelsonia sulcata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ptychopariidae.
Paracoosia langgongsiensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Anomocaridae.
Paralevisia subglobosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Parayujinia pulchra [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Peishania abnormis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Peishania obscura [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Peronopsis jiulongshanensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Peronopsis universalis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Persia [120]	Gen. et sp. nov	Junior homonym	Lerosey-Aubril	Carboniferous (Tournaisian)		Iran	The type species is Persia praecox. The generic name is preoccupied by Persia Repin (1996); Lerosey-Aubril & Deshmukh (2022) coined a replacement name Persiax. [121]
Pilletina seifenensis [114]	Sp. nov	Valid	Basse & Müller	Early Devonian	Seifen Beds	Germany	A member of Acastidae belonging to the subfamily Asteropyginae, a species of Pilletina.
Pingluia granulosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Plectocrania elegans [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Plectocrania intermedia [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Plesigangderria latilimbata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Podoliproetus mirdani [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Podoliproetus.
Podoliproetus sinespina [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Podoliproetus.
Praeymekops [101]	Gen. et sp. et comb. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Proasaphiscidae. Genus includes new species P. minutus, as well as P. sinensis (Mansuy, 1916).
Proasaphiscus laiwuensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Proetina ihamadii [112]	Sp nov	Valid	Johnson & Fortey	Early Devonian (Pragian)	Ihandar Formation	Morocco	A proetid trilobite, a species of Proetina.
Pseudoolenoides fossilmountainensis [122]	Sp nov	Valid	Adrain, McAdams & Karim	Middle Ordovician	Kanosh Formation	United States	A bathyurid, a species of Pseudoolenoides.
Pseudoolenoides oilcreekensis [122]	Sp nov	Valid	Adrain, McAdams & Karim	Middle Ordovician	Oil Creek Formation	United States	A bathyurid, a species of Pseudoolenoides.
Pseudoolenoides pogonipensis [122]	Sp nov	Valid	Adrain, McAdams & Karim	Middle Ordovician	Kanosh Formation	United States	A bathyurid, a species of Pseudoolenoides.
Pseudopeishania [101]	Gen. et 3 sp. et comb. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae. Genus includes new species P. brevis, P. similis and P. woqishanensis, as well as P. angustata (Endo, 1944).
Pseudoperonopsis elegans [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Pseudophillipsia parvizii [120]	Sp. nov	Valid	Lerosey-Aubril	Permian		Iran	A species of Pseudophillipsia sensu lato.
Pseudotaitzuia angustilimbata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Asaphida belonging to the family Ordosiidae.
Pseudotaitzuia rara [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Asaphida belonging to the family Ordosiidae.
Pudoproetus zhorae [104]	Sp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Proetidae, a species of Pudoproetus.
Raragnostus intermedius [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Raymondaspis grandigena [123]	sp nov	Valid	Stein & Bergström	Middle Ordovician		Sweden	A styginid, a species of Raymondaspis.
Reediella mitchelli tafilaltensis [104]	Subsp. nov	Valid	Hahn, Müller & Becker	Carboniferous (Mississippian)		Morocco	A member of Phillipsiidae belonging to the subfamily Bollandiinae, a subspecies of Reediella mitchelli .
Ruichengocephalus [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ptychopariidae. Genus includes new species R. typicus.
Saimachia yichangensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Schopfaspis? graciai [124]	sp nov	Valid	Esteve et al..	Middle Cambrian		Spain	An alokistocarid, possibly a species of Schopfaspis .
Sdzuyella conica [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belongig to the family Holocephalidae.
Securifrons [108]	gen et comb nov	Valid	Holloway and Lane	Ordovician to Silurian			Scutelluid trilobite. species- S. borealis, S. cunctatum, S. major and ?S. umbonatus.
Shiliuyuania [101]	Gen. et 2 sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Crepicephalidae. Genus includes new species S. cylindrica and S. quadrata.
Solenoparia zhangxianensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Solenoparina bella [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Sphaerophthalmus arcus [125]	Sp nov	Valid	Høyberget & Bruton	Cambrian (Furongian)	Alum Shale Formation	Norway	A member of the family Olenidae.
Spiniscabrella albertii [126]	Sp. nov	Valid	Müller & Franke	Devonian		Germany	A member of the family Homalonotidae.
Sudanomocarina elliptica [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Sudanomocarina hutoushanensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Symphysops stevaninae [127]	Sp. nov	Valid	López-Soriano & Corbacho	Late Ordovician	Lower Ktaoua Formation	Morocco	A member of Cyclopygidae, a species of Symphysops .
Szeaspis (Pseudoszeaspis) [101]	Subgen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae. The subgenus includes new species Szeaspis (Pseudoszeaspis) conicus, as well as Szeaspis huananensis Peng, Babcock & Lin (2004) and Szeaspis pustulosa (Resser & Endo in Endo & Resser, 1937).
Szeaspis crassus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae.
Taitzehoia huolianzhaiensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Damesellidae.
Teinistion jiulongshanense [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Damesellidae.
Teinistion quadratum [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Damesellidae.
Teinistion yangjiazhuangense [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Damesellidae.
Tomagnostus (Paratomagnostus) [101]	Subgen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ptychagnostidae. The subgenus includes Tomagnostus (Paratomagnostus) ezhuangensis.
Trachoparia elongata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Trachoparia intermedia [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Trachoparia spinosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Triangulopyge [125]	Gen. et comb. nov	Valid	Høyberget & Bruton	Cambrian		Norway   Sweden   United Kingdom	A member of the family Olenidae. The type species is "Olenus" humilis Phillips (1848); genus also includes "Sphaerophthalmus" majusculus Linnarsson (1880) and T. major (Lake, 1913).
Tsunyidiscus pengshuiensis [119]	Sp nov	Valid	Zhang & Clarkson	Cambrian		China	An eodiscoid trilobite, a species of Tsunyidiscus.
Ulania concava [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Utagnostus primitivus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Clavagnostidae.
Utagnostus sulcatus [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Clavagnostidae.
Woqishanaspis [101]	Gen. et comb. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ignotogregatidae. Genus includes W. trapezoidalis (Lin, 1984).
Wutaishanaspis yangjiazhuangensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Xenolorenzella granulosa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Yichengocephalus [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Solenopleuridae. Genus includes new species Y. latus.
Yongwolia quadrata [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Yushugouia ezhuangensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Zhangxiaspis [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Ptychopariidae. Genus includes new species Z. subcylindrica.
Zhongtiaoshanella [101]	Gen. et sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Crepicephalidae. Genus includes new species Z. brevispinosa.
Zhongweia convexa [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China
Zhujia yangjiazhuangensis [101]	Sp. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of Ptychopariida belonging to the family Anomocarellidae.
Zichuanaspis [101]	Gen. et 2 sp. et comb. nov	Valid	Yuan et al.	Cambrian	Changhia Formation	China	A member of the family Leiostegiidae. Genus includes Zichuanaspis acerius (Walcott, 1905) and Z. catinus (Resser & Endo in Endo & Resser, 1937), as well as new species Z. yangjiazhuangensis and possibly Z? latilimbata.

Others

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Australimicola [128]	Gen. et sp.	Valid	Paterson, García-Bellido and Edgecombe	Early Cambrian	The Emu Bay Shale Konservat-Lagerstätte	Australia	An artiopodan arthropod. The type species is Australimicola spriggi.
Kangacaris shui [129]	Sp. nov	Valid	Zhang, Fu & Dai	Lower Cambrian	Chengjiang lagerstätte	China	A species of Kangacaris.
Luohuilinella [130]	Gen. et sp. nov	Valid	Zhang, Fu & Dai	Lower Cambrian	Chengjiang Lagerstätte	China	A possible member of Xandarellida. The type species is Luohuilinella rarus.
Nereocaris [131]	Gen. et sp. nov	Valid	Legg et al.	Cambrian	Burgess Shale Formation	Canada	A bivalved arthropod, one of the basalmost known arthropods. The type species is Nereocaris exilis.
Pleuralata [132]	Gen. et sp. nov	Valid	McCoy, Strother and Briggs	Silurian (Llandovery)	Tuscarora Formation	United States	An arthropod of uncertain phylogenetic affinities, a probable tracemaker of Arthrophycus alleghaniensis trace fossils. The type species is Pleuralata spinosa.
Scolopocryptops simojovelensis [133]	Sp. nov	Valid	Edgecombe et al.	Miocene		Mexico	A scolopocryptopid scolopendromorph centipede, a species of Scolopocryptops.
Sinoleperditia hamadai [134]	Sp. nov	Valid	Tanaka, Ono and Maeda	Early Devonian		Japan	A member of Leperditicopida (a group of arthropods of uncertain affinities, classified as ostracods or excluded from this group by different authors), a species of Sinoleperditia.
Sinoleperditia huyeni [135]	Sp. nov	Valid	Tanaka, Phong and Komatsu	Early Devonian		Vietnam	A member of Leperditicopida, a species of Sinoleperditia.
Squamacula buckorum [128]	Species	Valid	Paterson, García-Bellido and Edgecombe	Early Cambrian	The Emu Bay Shale Konservat-Lagerstätte	Australia	An artiopodan arthropod, a species of Squamacula .

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