2012 in paleobotany

This paleobotany list records new fossil plant taxa that were described during the year 2012, as well as other significant paleobotany discoveries and events which occurred during 2012.

Bryophytes

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Bryokhutuliinia crassimarginata [1]	Sp nov		Ignatov et al	Late Jurassic	Ukurey formation	Russia	A moss of uncertain placement.

Lycophytes

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Winslowia [2]	Gen. et sp. nov	Valid	Dunn et al.	Carboniferous (Serpukhovian)	Pride Mountain Formation	United States	A member of Isoetales belonging to the group Chaloneriaceae. Genus includes new species W. tuscumbiana.

Marchantiophytes

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Riccardiothallus [3]	Gen. et sp. nov	Valid	Guo et al.	Devonian (Pragian)	Posongchong Formation	China	A liverwort belonging to the family Aneuraceae. Genus includes new species R. devonicus.

Ferns and fern allies

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Korallipteris [4]	Gen. et comb. nov	Valid	Vera & Passalia	Triassic to Miocene		Antarctica Argentina Chile New Zealand [5]	A fern of uncertain phylogenetic placement. A new genus for "Gleichenia" argentinica (1962); Additional species were originally placed in Gleichenia, Gleichenites and Microphyllopteris .
Millerocaulis tekelili [6]	Sp. nov	Valid	Vera	Early Cretaceous (Aptian)	Cerro Negro Formation	Antarctica (Livingston Island)	An Osmundaceae fern. Originally placed in Millerocaulis ; Moved to Claytosmunda tekelili in (2017). [7]
Osmundopsis rafaelii [8]	Sp. nov	Valid	Escapa & Cúneo	Early Jurassic		Argentina	A member of Osmundaceae.
Todites cacereii [8]	Sp. nov	Valid	Escapa & Cúneo	Early Jurassic		Argentina	A member of Osmundaceae.

Bennettitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Anomozamites harrisi [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Anomozamites heeri [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Anomozamites villosus [10]	Sp. nov	Valid	Pott et al.	Probably late Middle Jurassic	Daohugou Beds	China	A member of Bennettitales. Originally described as a species of Anomozamites Moved to the Wielandiella villosus (2015). [11] [12]
Cycadolepis barnardii [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis karatauensis [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis kirillii [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis kokalensis [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis lata [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis maija [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis multistomata [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis pachydermatica [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis perfecta [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis prynadае [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Cycadolepis schairensis [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Nilssoniopteris chirchilica [9]	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Кokala Suite	Kazakhstan
Nilssoniopteris localis [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Nilssoniopteris saveljevii [9]	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Tonasha Formation	Kazakhstan
Pterophyllum chagaense [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Pterophyllum doludenkoae [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Pterophyllum originalum [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Pterophyllum pаrvisegmentatum [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Pterophyllum pinniforme [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Pterophyllum valentinaeа [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Pterophyllum vsevolodii [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Williamsonia antonia [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Williamsonia baranovaeа [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Williamsonia chagaensе [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Williamsonia kokalensis [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Williamsonia vialovaea [9]	Sp. nov	Valid	Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan

"Pteridospermatophytes"

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ctenozamites schweitzerii [9]	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation	Kazakhstan	A member of Pteridospermatophyta of uncertain affinities.
Eretmonia macloughlinii [13]	Sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Upper Buckley Formation	Antarctica	A member of Glossopteridales.
Kuvakospermum [14]	Gen. et sp. nov	Valid	Naugolnykh & Sidorov	Permian (Wordian)		Russia	A seed fern belonging to the family Peltaspermaceae. Genus includes new species K. pedatum.
Lidgettoniopsis [15]	Gen. et sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Beacon Supergroup (upper Buckley Formation)	Antarctica	A seed fern belonging to the group Dictyopteridiales and the family Eretmoniaceae. Genus includes new species L. ramulus.
Scutum leiophyllum [15]	Sp. nov	Valid	Ryberg, Taylor & Taylor	Late Permian	Beacon Supergroup (upper Buckley Formation)	Antarctica	A member of Glossopteridales belonging to the family Dictyopteridiaceae.
Vetlugospermum [16]	Gen. et sp. nov	Valid	Naugolnykh	Early Triassic		Russia	A seed fern belonging to the group Peltaspermales. Genus includes new species V. rombicum.

Ginkgoales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Baiera kazachstanica [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Кokala Suite	Kazakhstan
Baieroxylon rocablanquense [17]	Sp. nov	Valid	Gnaedinger	Early Jurassic	Roca Blanca Formation	Argentina
Ginkgo cranei [18]	Sp. nov	Valid	Zhou, Quan, & Liu	upper Paleocene	Sentinel Butte Formation	USA North Dakota	An extinct species in the modern genus Ginkgo
Ginkgo gomolitzkyana [19]	Sp. nov	Valid	Nosova	Middle Jurassic	Angren Formation	Uzbekistan	A member of Ginkgoales.
Ginkgo huolinhensis [20]	Sp. nov	Valid	Dong & Sun	Early Cretaceous	Huolinhe Formation	China
Ginkgoites barnardianus [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Кokala Suite	Kazakhstan
Ginkgoites irdalicus [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic	Ergozi Suite	Kazakhstan
Ginkgoites kelendensis [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Aalenian)	Tonasha Formation	Kazakhstan
Ginkgoites ketovianus [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Bathonian)	Sarydiirmeny Suite	Kazakhstan
Leptotoma asiatica [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation	Kazakhstan
Pseudotorellia karadiirmenica [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation	Kazakhstan

Conifers

Araucariaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Emwadea [21]	Gen. et sp. nov	Valid	Dettmann, Clifford & Peters	Early Cretaceous (Albian)	Winton Formation	Australia	A member of the family Araucariaceae. Genus includes new species E. microcarpa.

Cheirolepidiaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Duartenia [22]	Gen. et sp. nov	Valid	Mohr et al.	Early Cretaceous	Crato Formation	Brazil	A possible member of Cheirolepidiaceae. The type species is Duartenia araripensis.
Glenrosa falcata [23]	Sp. nov	Valid	Gomez, Ewin & Daviero-Gomez	Early Cretaceous (Barremian)	Calizas de La Huergina Formation	Spain	A conifer.

Cupressaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Calocedrus huashanensis [24]	Sp. nov	Valid	Shi, Zhou, & Xie	Oligocene	Ningming Formation	China	A Calocedrus species
Fokienia shengxianensis [25]	Sp. nov	Valid	He, Sun & Liu	Late Miocene	Shengxian Formation Xiaolongtan Formation	China	Originally placed in Fokienia . Moved to Calocedrus shengxianensis in (2015). [26]

Pinaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Eathiestrobus [27]	Gen. et sp. nov	Valid	Rothwell et al.	Late Jurassic (Kimmeridgian)	Kimmeridge Clay Formation	United Kingdom	A member of Pinaceae. The type species is Eathiestrobus mackenziei.
Picea burtonii [28]	Sp. nov	Valid	Klymiuk & Stockey	Early Cretaceous (Valanginian)		Canada	A spruce species seed cone.
Pinus yorkshirensis [29]	Sp. nov	Valid	Ryberg et al.	Early Cretaceous (Hauterivian-Barremian)	Speeton Clay Formation	United Kingdom	A pine.

Podocarpaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Dacrycarpus puertae [30]	Sp. nov	Valid	Wilf	Eocene		Argentina	A species of Dacrycarpus

Taxaceae

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Marskea kazachstanica [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Kokala Suite	Kazakhstan
Torreya kazachstanica [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Kokala Suite	Kazakhstan

Other conifers

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Elatocladus bulakus [9]	Sp. nov	Valid	Nosova & Kiritchkova in Kiritchkova & Nosova	Early Jurassic (Toarcian)	Kokala Suite	Kazakhstan	A conifer of uncertain affinities.
Plyophyllioxylon [31]	Gen. et sp. nov	Valid	Feng et al.	Permian (Roadian/Wordian)	Lower Shihhotse Formation	China	A conifer of uncertain phylogenetic placement. Genus includes new species P. hulstaiense.

Other gymnosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Nilssonia prosvirjakovaea [9]	Sp. nov	Valid	Kiritchkova & Nosova	Middle Jurassic (Bajocian)	Karadiirmen' Formation	Kazakhstan	A cycad.
Rehezamites [32]	Gen. et sp. nov	Valid	Pott et al.	Early Cretaceous (Aptian)	Yixian Formation	China	A seed plant of uncertain phylogenetic placement. Genus includes new species R. anisolobus.

Angiosperms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Archaeampelos senonica [33]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Baasia [34]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation	United States	A member of the family Celastraceae. Genus includes new species B. armendarisense.
Bertilanthus [35]	Gen. et sp. nov	Valid	Friis & Pedersen	Late Cretaceous		Sweden	An asterid. Genus includes new species B. scanicus.
Camptodromites sibiricus [33]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Celastrinites lanceolatus [33]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Chaneya hainanensis [36]	Sp. nov	Valid	Feng & Jin	Eocene	Changchang Basin	China	A fossil fruit.
Cissites basicordatus [33]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Cissus lombardii [37]	Sp. nov	Valid	Manchester, Chen & Lott	Early Oligocene	Mancora Formation	Peru	A species of Cissus .
Eucalyptus caldericola [38]	Sp. nov	Valid	Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)		Argentina	A species of Eucalyptus .
Eucalyptus frenguelliana [38]	Sp. nov	Valid	Gandolfo & Zamaloa in Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)		Argentina	A species of Eucalyptus .
Eucalyptus lynchiae [38]	Sp. nov	Valid	Gandolfo & Hermsen in Hermsen, Gandolfo & Zamaloa	Eocene (Ypresian)		Argentina	A species of Eucalyptus .
Fulleroxylon [34]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation	United States	A member of the family Myrtaceae. Genus includes new species F. armendarisense.
Juglandiphyllites microdentatus [33]	Sp. nov	Valid	Alekseev	Late Cretaceous		Russia
Laurinoxylon stichkai [39]	Sp. nov	Valid	Boonchai & Manchester	Early Eocene	Bridger Formation	United States	A member of Lauraceae described on the basis of fossil wood.
Laurophyllum calicarioides [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Litsea .
Laurophyllum lacustris [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia .
Laurophyllum maarensis [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya .
Laurophyllum microphyllum [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya .
Laurophyllum otagoensis [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia .
Laurophyllum sylvestris [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia .
Laurophyllum taieriensis [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya .
Laurophyllum vulcanicola [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Beilschmiedia .
Laurophyllum waipiata [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae with affinities to members of the genus Cryptocarya .
Litseopsis nova-zelandiae [40]	Sp. nov	Valid	Bannister, Conran & Lee	Early Miocene		New Zealand	A member of Lauraceae.
Mahonia grimmii [41]	Sp. nov	Valid	Güner & Denk	Miocene	Eskihisar Formation	Turkey	A species of Mahonia .
Mahonia somaensis [41]	Sp. nov	Valid	Güner & Denk	Miocene	Soma Formation	Turkey	A species of Mahonia .
Morinda chinensis [42]	Sp. nov	Valid	Shi et al.	Eocene	Changchang Formation	China	A species of Morinda .
Newtonia mushensis [43]	Sp. nov	Valid	Pan et al.	Early Miocene		Ethiopia	A species of Newtonia .
Palaeophytocrene hammenii [44]	Sp. nov	Valid	Stull in Stull et al.	Middle-late Paleocene	Bogotá Formation	Colombia	A member of Icacinaceae belonging to the tribe Phytocreneae.
Palaeophytocrene piggae [44]	Sp. nov	Valid	Stull in Stull et al.	Late Paleocene	Sentinel Butte Formation	United States	A member of Icacinaceae belonging to the tribe Phytocreneae.
Pygmaeoxylon [34]	Gen. et sp. nov	Valid	Estrada-Ruiz et al.	Late Cretaceous (probably early to middle Maastrichtian)	McRae Formation	United States	A magnoliid of uncertain phylogenetic placement. Genus includes new species P. paucipora.
Pyrenacantha austroamericana [44]	Sp. nov	Valid	Stull in Stull et al.	Late early Oligocene		Peru	A species of Pyrenacantha .
Schenkeriphyllum glanduliferum [45]	Gen. et sp. nov	Valid	Mohr, Coiffard & Bernardes-de-Oliveira	Early Cretaceous (Aptian)	Crato Formation	Brazil	Fossil magnolialean angiosperm.
Wilsonoxylon [39]	Gen. et sp. nov	Valid	Boonchai & Manchester	Early Eocene	Bridger Formation	United States	A member of Canellaceae described on the basis of fossil wood. Genus includes new species W. edenense.

References

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