Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families.
Perissodactyla is an order of ungulates. The order includes about 17 living species divided into three families: Equidae, Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). They typically have reduced the weight-bearing toes to three or one of the five original toes, though tapirs retain four toes on their front feet. The nonweight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, artiodactyls bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that perissodactyls digest plant cellulose in their intestines, rather than in one or more stomach chambers as artiodactyls, with the exception of Suina, do.
Ungulates are members of the diverse clade Euungulata, which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
Merychippus is an extinct proto-horse of the family Equidae that was endemic to North America during the Miocene, 15.97–5.33 million years ago. It had three toes on each foot and is the first horse known to have grazed.
Miohippus is an extinct genus of horse existing longer than most Equidae. It lived in what is now North America from 32 to 25 million years ago, during the late Eocene to late Oligocene. According to the Florida Museum of Natural History, Othniel Charles Marsh first believed Miohippus lived during the Miocene and thus named the genus using this incorrect conclusion. More recent research provides evidence that Miohippus actually lived during the Paleogene period.
Ailuridae is a family in the mammal order Carnivora. The family consists of the red panda and its extinct relatives.
Chalicotheriidae is an extinct family of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Pleistocene, existing from 48.6 to 1.806 mya. They are often called chalicotheres, a term which is also applied to the broader grouping of Chalicotherioidea. They are noted for their unusual morphology compared to other ungulates, such as their elongated clawed forelimbs. They are thought to have been browsers.
Chalicotherium is a genus of extinct odd-toed ungulates of the order Perissodactyla and family Chalicotheriidae. The genus is known from Europe and Asia, from the Middle Miocene to Late Miocene.
Ancylotherium is an extinct genus of the family Chalicotheriidae, subfamily Schizotheriinae, endemic to Europe, Asia, and Africa during the Late Miocene-Early Pleistocene, existing for approximately 9.8 million years.
Amphicyon is an extinct genus of large carnivorans belonging to the family Amphicyonidae, subfamily Amphicyoninae, from the Miocene epoch. Members of this family received their vernacular name for possessing bear-like and dog-like features. They ranged over North America, Europe, Asia, and Africa.
Hemicyon, also known as the "dog-bear", is an extinct genus of hemicyonine bear, which probably originated in Eurasia but was found in Europe, Asia and North America during the Miocene epoch, existing for approximately 16 to 13 mya. Hemicyon is the best-known genus in the Hemicyoninae, a subfamily intermediate between bears and their Caniform ancestors but most often classified as bears. Hemicyonid bears should not be confused with Amphicyonids (bear-dogs), which are their own separate family of carnivores.
The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the modern horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.
Simocyon is a genus of extinct carnivoran mammal in the family Ailuridae. Simocyon, which was about the size of a mountain lion, lived in the late Miocene and early Pliocene epochs, and has been found in Europe, Asia, and rarely, North America and Africa.
Protohippus is an extinct three-toed genus of horse. It was roughly the size of a modern donkey. Fossil evidence suggests that it lived during the Late Miocene, from about 13.6 Ma to 5.3 Ma.
Parahippus leonensis is an extinct proto-horse of the family Equidae that was endemic to North America during the Miocene from 23.030 to 16.3 Ma living for approximately 6.73 million years.
Nannippus is an extinct genus of three-toed horse endemic to North America during the Miocene through Pleistocene, about 13.3—1.8 million years ago (Mya), living around 11.5 million years. This ancient species of three-toed horse grew up to 3.5 feet and weighed between 165 pounds to 199 pounds, which was around the same size as a domestic sheep.
The Anchitheriinae are an extinct subfamily of the Perissodactyla family Equidae, the same family which includes modern horses, zebras and donkeys. This subfamily is more primitive than the living members of the family. The group first appeared with Mesohippus in North America during the middle Eocene and thrived until the late Miocene. The subfamily continued in Eurasia with the genus Sinohippus until the early Pliocene, when it finally became extinct. This subfamily are considered browsers with their more ancestral low-crowned molars, in contrast with the modern, specialized grazer species of modern Equinae.
Archaeohippus is an extinct three toed member of the family Equidae known from fossils of early Oligocene to middle Miocene age. The genus is noted for several distinct skeletal features. The skull possesses deeply pocketed fossa in a notably long preorbital region. The genus is considered an example of phyletic dwarfism with adults estimated at being on average 20 kilograms in weight. This is in contrast to the most common equid of the period, Miohippus. Characters of the teeth show a mix of both primitive and advanced traits. The advanced traits are very similar to those shown in the genus Parahippus. The noted similarities of Archaeohippus and Parahippus show them to be descended from a common ancestor. They are considered sister species.
Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos with some developing bony nose horns, and were some of the earliest mammals to have evolved large body sizes. They lived around 56–34 million years ago, until the very close of the Eocene. Brontotheres had a Holarctic distribution, with the exception of Western Europe: they occupied North America, Asia, and Eastern Europe. They were the first fossilized mammals to be discovered west of the Mississippi, and were first discovered in South Dakota.
Cerro de los Batallones is a hill at Torrejón de Velasco, Madrid, Spain where a number of fossil sites from the Upper Miocene (MN10) have been found. Nine sites have been discovered with predominantly vertebrate fossils, invertebrates and plants being less represented. The first deposits were discovered accidentally in July 1991.
