Cormohipparion

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Cormohipparion
Temporal range: Mid Miocene-Early Pliocene
(Hemphillian-Blancan)
~15.97–3.6  Ma
Cormohipparion occidentale (sturdy three-toed horse) (Ash Hollow Formation, Miocene, 11.83 Ma; Ashfall Fossil Beds, Nebraska, USA) 4.jpg
C. occidentale skeleton
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Subfamily: Equinae
Tribe: Hipparionini
Genus: Cormohipparion
Skinner & MacFadden, 1977
Type species
Hipparion occidentale
Subgenera and species

Cormohipparion

  • C. (C.) cappadociumBernor, 2024
  • C. (C.) frickiWoodburne, 2007
  • C. (C.) goorisiMacFadden & Skinner, 1981
  • C. (C.) johnsoniWoodburne, 2007
  • C. (C.) matthewiWoodburne, 2007
  • C. (C.) merriamiWoodburne, 2007
  • C. (C.) occidentale(Leidy, 1856) (type)
  • C. (C.) quinniWoodburne, 1996
  • C. (C.) skinneriWoodburne, 2007

Notiocradohipparion

  • C. (N.) plicatile(Leidy, 1887)
  • C. (N.) ingenuum(Leidy, 1885)
  • C. (N.) emslieiHulbert, 1988

Cormohipparion (Greek: "noble" (cormo), "pony" (hipparion) [1] is an extinct genus of horse belonging to the tribe Hipparionini that lived in North America and Eurasia during the late Miocene to Pliocene (Hemphillian to Blancan in the NALMA classification). [2] They grew up to 3 feet (0.91 meters) long. [3] [4]

Contents

Taxonomy

Skull Cormohipparion occidentale.jpg
Skull

The genus Cormohipparion was coined for the extinct hipparionin horse "Equus" occidentale, described by Joseph Leidy in 1856. [5] However it was soon argued that the partial material fell within the range of morphological variation seen in Hipparion , and that the members of Cormohipparion belonged instead within Hipparion. [6] [7] This rested on claims that pre-orbital morphology did not have any taxonomic significance, a claim that detailed study of quarry sections later showed to be false. [8] The genus was originally identified by a closed off preorbital fossa, but later examinations of the cheek teeth, specifically the lower cheek teeth, of Cormohipparion specimens found that they were indeed valid and distinct from Hipparion. [9] A reappraisal of many horse genera was thus conducted in 1984, [10] and the proposed synonymy was not acknowledged by later literature. [11] C. ingenuum holds the distinction for being the first prehistoric horse to be described in Florida, as well as being one of the most common species of extinct three-toed horses found to be in Florida, lasting until the early Pliocene. [12] [13] Cormohipparionemsliei has the distinction of being the last hipparion horse known from the fossil record. [14]

The genus is considered to represent an ancestor to Hippotherium . [15] Its fossils have been recovered from as far south as Mexico. [16] Fossils have been found in the Great Plains and Rio Grande regions of North America, Mexico, Florida, and Texas, which shows that they were herding animals. [17] [18] [19] [20] Fossils have been unearthed in California, [21] Louisiana, [22] [23] Nebraska, [24] [25] South Dakota, [26] Honduras, [27] Costa Rica, [28] and Panama. [29] Fossils have also been found in India and Turkey. [30] [31]

Evolution

Skeletons of Cormohipparion and Teleoceras from the Ashfall Fossil Beds Ashfall Fossil Beds - Teleoceras and Cormohipparion.jpg
Skeletons of Cormohipparion and Teleoceras from the Ashfall Fossil Beds

A species of Cormohipparion closely related to C. occidentale is thought to have crossed the Bering land Bridge over into Eurasia around 11.4-11 million years ago, becoming the ancestor to Old World hipparionines. [32]

References

  1. C. Hulbert Jr., Richard (October 17, 2013). "Cormohipparion ingenuum". Florida Vertebrate Fossils. Retrieved 2025-04-09.
  2. MacFadden, Bruce J.; Skinner, Morris F. (1982). "Hipparion Horses and Modern Phylogenetic Interpretation: Comments on Forsten's View of Cormohipparion". Journal of Paleontology. 56 (6): 1336–1342. ISSN   0022-3360. JSTOR   1304670.
  3. "Region 4: The Great Plains". geology.teacherfriendlyguide.org. Retrieved 2021-06-23.
  4. Czaplewski, Nicholas J.; Thurmond, J. Peter; Wyckoff, Don G. (Fall 2001). "Wild Horse Creek #1: A Late Miocene (Clarendonian-Hemphillian) Vertebrate Fossil Assemblage in Roger Mills County, Oklahoma" (PDF). Oklahoma Geology. 61 (3): 60–67.
  5. Skinner, M. F.; MacFadden, B. J. (1977). "Cormohipparion n. gen. (Mammalia, Equidae) from the North American Miocene (Barstovian-Clarendonian)". Journal of Paleontology. 51 (5). Paleontological Society: 912–926. JSTOR   1303763.
  6. Forsten, A. (1982). "The Status of the Genus Cormohipparion Skinner and MacFadden (Mammalia, Equidae)". Journal of Paleontology. 56 (6). Paleontological Society: 1332–1335. JSTOR   1304669.
  7. MacFadden, Bruce J. (1985). "Patterns of Phylogeny and Rates of Evolution in Fossil Horses: Hipparions from the Miocene and Pliocene of North America". Paleobiology. 11 (3): 245–257. Bibcode:1985Pbio...11..245M. doi:10.1017/S009483730001157X. ISSN   0094-8373. JSTOR   2400665. S2CID   89569817.
  8. MacFadden, B. J.; Skinner, M. F. (1982). "Hipparion Horses and Modern Phylogenetic Interpretation_ Comments on Forsten's View of Cormohipparion". Journal of Paleontology. 56 (6). Paleontological Society: 1336–1342. JSTOR   1304670.
  9. Forsten, Ann (1982). "The Status of the Genus Cormohipparion Skinner and MacFadden (Mammalia, Equidae)". Journal of Paleontology. 56 (6): 1332–1335. ISSN   0022-3360. JSTOR   1304669.
  10. MacFadden, BJ (1984). "Systematics and phylogeny of Hipparion, Neohipparion, Nannippus, and Cormohipparion (Mammalia, Equidae) from the Miocene and Pliocene of the new world". American Museum of Natural History.
  11. Hulbert Jr, R. C. (1988). "A New Cormohipparion (Mammalia, Equidae) from the Pliocene (Latest Hemphillian and Blancan) of Florida". Journal of Vertebrate Paleontology. 7 (4). The Society of Vertebrate Paleontology: 451–468. Bibcode:1988JVPal...7..451H. doi:10.1080/02724634.1988.10011675. JSTOR   4523166.
  12. "Cormohipparion ingenuum". Florida Museum. 2017-03-30. Retrieved 2021-06-16.
  13. "Late Pliocene (Late Blancan) vertebrates from the St. Petersburg Times site, Pinellas County, Florida, with a brief review of Florida Blancan faunas | WorldCat.org". search.worldcat.org. Retrieved 2025-04-09.
  14. "Neohipparion". Florida Museum. 2018-02-16. Retrieved 2021-06-26.
  15. Woodburne, M. O. (2005). "A New Occurrence of Cormohipparion, with Implications for the Old World Hippotherium Datum". Journal of Vertebrate Paleontology. 25: 256–257. doi:10.1671/0272-4634(2005)025[0256:ANOOCW]2.0.CO;2. ISSN   0272-4634. S2CID   86241044.
  16. Bravo-Cuevas, V. M.; Ferrusquía-Villafranca, I. (2008). "Cormohipparion (Mammalia, Perissodactyla, Equidae) from the Middle Miocene of Oaxaca, Southeastern Mexico". Journal of Vertebrate Paleontology. 28: 243–250. doi:10.1671/0272-4634(2008)28[243:CMPEFT]2.0.CO;2. ISSN   0272-4634. S2CID   131613568.
  17. Skinner, Morris F.; MacFadden, Bruce J. (1977). "Cormohipparion n. gen. (Mammalia, Equidae) from the North American Miocene (Barstovian-Clarendonian)". Journal of Paleontology. 51 (5): 912–926. ISSN   0022-3360. JSTOR   1303763.
  18. Bravo-Cuevas, Victor Manuel; Ferrusquía-Villafranca, Ismael (2008). "Cormohipparion (Mammalia, Perissodactyla, Equidae) from the Middle Miocene of Oaxaca, Southeastern Mexico". Journal of Vertebrate Paleontology. 28 (1): 243–250. doi:10.1671/0272-4634(2008)28[243:CMPEFT]2.0.CO;2. ISSN   0272-4634. JSTOR   30126348. S2CID   131613568.
  19. Hulbert, Richard C. (1988). "A New Cormohipparion (Mammalia, Equidae) from the Pliocene (Latest Hemphillian and Blancan) of Florida". Journal of Vertebrate Paleontology. 7 (4): 451–468. Bibcode:1988JVPal...7..451H. doi:10.1080/02724634.1988.10011675. ISSN   0272-4634. JSTOR   4523166.
  20. MacFadden, Bruce J.; Skinner, Morris F. (1981). "Earliest Holarctic Hipparion, Cormohipparion goorisi n. sp. (Mammalia, Equidae), from the Barstovian (Medial Miocene) Texas Gulf Coastal Plain". Journal of Paleontology. 55 (3): 619–627. ISSN   0022-3360. JSTOR   1304276.
  21. WHISTLER, DAVID P.; BURBANK, DOUGLAS W. (1992-06-01). "Miocene biostratigraphy and biochronology of the Dove Spring Formation, Mojave Desert, California, and characterization of the Clarendonian mammal age (late Miocene) in California" . GSA Bulletin. 104 (6): 644–658. Bibcode:1992GSAB..104..644W. doi:10.1130/0016-7606(1992)104<0644:MBABOT>2.3.CO;2. ISSN   0016-7606.
  22. Schiebout, Judith A.; Ting, Suyin; Williams, Michael; Boardman, Grant; Gose, Wulf (2004-04-01). Paleofaunal and Environmental Research on Miocene Fossil Sites TVOR SE and TVOR S on Fort Polk, Louisiana, with Continued Survey, Collection, Processing, and Documentation of other Miocene Localities (Report). Fort Belvoir, VA: Defense Technical Information Center. doi:10.21236/ada422018.
  23. "Abstract: TERRESTRIAL CARNIVORES, ARTIODACTYLS, PERISSODACTYLS, AND PROBOSCIDEANS OF THE FORT POLK MIOCENE SITES OF LOUISIANA (South-Central Section - 45th Annual Meeting (27–29 March 2011))". gsa.confex.com. Retrieved 2025-04-09.
  24. Otto, Rick (2017-01-01). "Five Species of Fossil Equids Preserved In-situ at Ashfall Fossil Beds". University of Nebraska State Museum: Programs Information.
  25. Tucker, S. T.; Otto, R. E.; Joeckel, R. M.; Voorhies, M. R. (2014-01-01), Korus, Jesse T. (ed.), "The geology and paleontology of Ashfall Fossil Beds, a late Miocene (Clarendonian) mass-death assemblage, Antelope County and adjacent Knox County, Nebraska, USA", Geologic Field Trips along the Boundary between the Central Lowlands and Great Plains: 2014 Meeting of the GSA North – Central Section, vol. 36, Geological Society of America, p. 0, ISBN   978-0-8137-0036-6 , retrieved 2025-04-09
  26. Pagnac, Darrin. "Additions to the Vertebrate Faunal Assemblage of the Middle Miocene Fort Randall Formation in the Vicinity of South Bijou Hill, South Dakota, U.S.A".{{cite journal}}: Cite journal requires |journal= (help)
  27. Webb, S. David; Perrigo, Stephen C. (1984). "Late Cenozoic Vertebrates from Honduras and El Salvador" . Journal of Vertebrate Paleontology. 4 (2): 237–254. Bibcode:1984JVPal...4..237W. doi:10.1080/02724634.1984.10012006. ISSN   0272-4634. JSTOR   4522986.
  28. Miller, Stephanie; Valerio, Ana L.; Sudasassi, Arturo; Laurito, César A. (2022). "Cormohipparion quinni (Mammalia, Equidae, Hipparionini) del Mioceno Medio (Barstoviano Tardío) del Norte de Costa Rica". Revista geológica de América central (in Spanish). 66: 1–8. doi: 10.15517/rgac.v66i0.49763 . ISSN   2215-261X.
  29. MacFadden, Bruce J.; Jones, Douglas S.; Jud, Nathan A.; Moreno-Bernal, Jorge W.; Morgan, Gary S.; Portell, Roger W.; Perez, Victor J.; Moran, Sean M.; Wood, Aaron R. (2017). "Integrated Chronology, Flora and Faunas, and Paleoecology of the Alajuela Formation, Late Miocene of Panama". PLOS ONE. 12 (1): e0170300. Bibcode:2017PLoSO..1270300M. doi: 10.1371/journal.pone.0170300 . ISSN   1932-6203. PMC   5249130 . PMID   28107398.
  30. Basu, Prabir Kumar (2004-01-01). "Siwalik mammals of the Jammu Sub-Himalaya, India: an appraisal of their diversity and habitats" . Quaternary International. 5th International conference on the cenozoic evolution of the Asia-Pacific environment. 117 (1): 105–118. Bibcode:2004QuInt.117..105B. doi:10.1016/S1040-6182(03)00120-4. ISSN   1040-6182.
  31. Bernor, Raymond Louis; Ataabadi, Majid Mirzaie; Basoglu, Oksan; Cirilli, Omar; Kaya, Ferhat; Pehlevan, Cesur; Niknahad, Mansoureh; Vaziri, Mohammad Reza; Arab, Ahmad Lotfabad (November 2024). "Cormohipparion cappadocium, a new species from the Late Miocene of Yeniyaylacık, Türkiye, and the emergence of western Eurasian hipparion bioprovinciality". Annales Zoologici Fennici. 61 (1). doi:10.5735/086.061.0119. ISSN   0003-455X. Archived from the original on 2025-03-29.
  32. Bernor, Raymond L.; Kaya, Ferhat; Kaakinen, Anu; Saarinen, Juha; Fortelius, Mikael (October 2021). "Old world hipparion evolution, biogeography, climatology and ecology". Earth-Science Reviews. 221: 103784. Bibcode:2021ESRv..22103784B. doi:10.1016/j.earscirev.2021.103784.