Larus

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Larus
Great Black Backed Gull Ireland's Eye.jpg
Great black-backed gull Larus marinus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Charadriiformes
Family: Laridae
Subfamily: Larinae
Genus: Larus
Linnaeus, 1758
Type species
Larus marinus
Linnaeus, 1758
Species

See list

Larus is a large genus of gulls with worldwide distribution (by far the greatest species diversity is in the Northern Hemisphere).

Contents

Many of its species are abundant and well-known birds in their ranges. Until about 2005–2007, most gulls were placed in this genus, but this arrangement is now known to be polyphyletic, leading to the resurrection of the genera Chroicocephalus , Ichthyaetus , Hydrocoloeus , and Leucophaeus for many other species formerly included in Larus.

They are in general medium-large birds, typically pale grey to black above and white below and on the head, often with black markings with white spots ("mirrors") on their wingtips and in a few species also some black on the tail. They have stout, longish bills and webbed feet; in winter, the head is often streaked or smudged dark grey. The young birds are brown, and take three to five years to reach adult plumage, with subadult plumages intermediate between the young and adult. [1]

The taxonomy of the large gulls in the herring and lesser black-backed complex is complicated, with different authorities recognising from two species in the past, increasingly up to eight species more recently.

Taxonomy

The genus Larus was introduced in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his Systema Naturae . [2] The genus name is from Ancient Greek laros (λάῥος) or Latin larus, which appears to have referred to a gull or other large seabird. [3] The type species is the great black-backed gull (Larus marinus). [4] [5] The Latin name Larus marinus translates as "sea gull", and the gulls in this genus generally are the species most often known colloquially as "seagulls".

Species

The genus contains 24 extant species. [6]

ImageCommon NameScientific nameDistribution
Larus pacificus - Derwent River Estuary.jpg Pacific gull L. pacificusAustralia.
Gaviota peruana, Playa La Mina, Paracas, Ica, Peru.JPG Belcher's gull L. belcherithe Pacific coast of South America.
Larus crassirostris -Japan-8.jpg Black-tailed gull L. crassirostriscoasts of the East China Sea, Japan, Manchuria and the Kuril Islands, to western Alaska in North America.
Larus atlanticus1.jpg Olrog's gull L. atlanticusAtlantic coast of southern Brazil, Uruguay, and northern Argentina.
Larus heermanni at Richardson Bay.jpg Heermann's gull L. heermanniPacific coast of the United States, Mexico and extreme southwestern British Columbia.
Larus canus winter plumage.jpg Common gull L. canusnorthern Palearctic (northern Europe, northern Asia), wintering south to the Mediterranean and China.
Mew Gull RWD1.jpg Short-billed gull or mew gullL. brachyrhynchusnorthwestern North America.
Larus-delawarensis-021.jpg Ring-billed gull L. delawarensisCanada and the northern United States, wintering south to the Caribbean.
Larus livens 2910976.jpg Yellow-footed gull L. livensGulf of California in Mexico.
Larus occidentalis (Western Gull), Point Lobos, CA, US - May 2013.jpg Western gull L. occidentalisfrom British Columbia, Canada to Baja California, Mexico.
Larus cachinnans 3 (Marek Szczepanek).jpg Caspian gull L. cachinnansBlack and Caspian Seas and adjacent river basins, extending east into Central Asia; wintering west into Europe and south to the coasts of Arabia.
Kelp Gull (Larus dominicanus) -Hot Water Beach -NZ-8.jpg Kelp gull (called southern black-backed gull or karoro in New Zealand)L. dominicanusAntarctic Peninsula, southern South America, southern Africa, Southern Ocean Islands, southeastern Australia, and New Zealand.
Silbermowe - Larus argentatus.jpg European herring gull L. argentatusnorthern and northwestern Europe.
GULL, VEGA (9-5-08) Gambell, Ak -03 (2844589388).jpg Vega gull (or East Siberian gull)L. vegaenortheastern Siberia, wintering in Japan, Korea, southern and eastern China, and Taiwan.
Gaivota-Ferragudo.JPG Yellow-legged gull L. michahellisMediterranean Sea and Macaronesia, dispersing north as far as the British Isles after breeding.
Armenian gull near Sevanavank, side view.jpg Armenian gull L. armenicusinterior Turkey, the Caucasus and the Middle East, wintering in the eastern Mediterranean and the Persian Gulf.
Great Black-backed Gull Larus marinus.jpg Great black-backed gull L. marinusnorthwest European coasts, northeast North American coasts, and islands of the North Atlantic.
Glaucous Gull (5532943864).jpg Glaucous gull L. hyperboreusArctic Ocean coasts, wintering in the North Atlantic and North Pacific as far south as the British Isles and northernmost states of the United States, also on the Great Lakes.
Larus-fuscus-taxbox.jpg Lesser black-backed gull L. fuscusAtlantic coasts of Europe and northwest Asia, wintering south to central Africa and India.
Larus californicus Palo Alto May 2011 009.jpg California gull L. californicuswestern interior North America from the Northwest Territories, Canada south to eastern California and Colorado, wintering south to western Mexico.
American Herring Gull - natures pics.jpg American herring gull L. smithsonianusNorth America from central and southern Alaska to the Great Lakes and northeast coast of the United States from Maine south to North Carolina, wintering south to the Caribbean.
Glaucous-winged Gull RWD1.jpg Glaucous-winged gull L. glaucescensPacific Ocean coasts of Russia and Alaska to the coast of Washington, wintering south to Japan and northwest Mexico.
Ooseguro-kamome.jpg Slaty-backed gull L. schistisagusnortheastern coasts of Asia, wintering south to eastern China.
Adult Larus glaucoides, Swallow Pond 2.jpg Iceland gull L. glaucoidesArctic Ocean coasts of Canada and Greenland, wintering from Iceland south to the British Isles and the far northeast USA, with subspecies thayeri wintering on the Pacific coast of North America.

Fossils

Fossils of Larus gulls are known from the Middle Miocene, about 20-15 million years ago; allocation of earlier fossils to this genus is generally rejected. Biogeography of the fossil record suggests that the genus evolved in the northern Atlantic and spread globally during the Pliocene, when species diversity seems to have been highest, as with most seabirds.

  • Larus sp. (Middle Miocene of Grund, Austria)
  • Larus sp. (Middle Miocene of Romania) [7]
  • Larus sp. (Late? Miocene/Early Pliocene of Lee Creek Mine, U.S.) - several species [7]
  • Larus elmorei (Middle Pliocene of Bone Valley, southeastern U.S.)
  • Larus lacus (Late Pliocene of Pinecrest, southeastern U.S.)
  • Larus perpetuus (Pliocene of southeastern U.S.)
  • Larus sp. (San Diego Late Pliocene of the southwestern U.S.)
  • Larus oregonus (Late Pliocene - Late Pleistocene of the west-central U.S.)
  • Larus robustus (Late Pliocene - Late Pleistocene of the west-central U.S.)
  • Larus sp. (Late Pleistocene of Lake Manix western U.S.)

"Larus" raemdonckii (Early Oligocene of Belgium) is now at least tentatively believed to belong in the procellariiform genus Puffinus . "L." elegans (Late Oligocene?/Early Miocene of St-Gérand-le-Puy, France) and "L." totanoides (Late Oligocene?/Early Miocene of southeastern France) are now in Laricola , while "L." dolnicensis (Early Miocene of the Czech Republic) was actually a pratincole; it is now placed in Mioglareola .

The Early Miocene "Larus" desnoyersii (southeastern France) and "L." pristinus (John Day Formation, Willow Creek, U.S.) probably do not belong in this genus; the former may be a skua. [8]

Ring species

The Larus gulls formerly interpreted as a ring around the Arctic: 1. L. fuscus graellsii & L. f. intermedius (Western European lesser black-backed gulls); 2. L. fuscus fuscus (North East European lesser black-backed gull); 3. L. fuscus heuglini (Heuglin's gull); 4. L. vegae birulai (Birula's gull); 5. L. vegae sensu stricto (Vega gull); 6. L. smithsonianus (American herring gull); 7. L. argentatus (European herring gull) Ring species seagull.svg
The Larus gulls formerly interpreted as a ring around the Arctic: 1. L. fuscus graellsii & L. f. intermedius (Western European lesser black-backed gulls); 2. L. fuscus fuscus (North East European lesser black-backed gull); 3. L. fuscus heuglini (Heuglin's gull); 4. L. vegae birulai (Birula's gull); 5. L. vegae sensu stricto (Vega gull); 6. L. smithsonianus (American herring gull); 7. L. argentatus (European herring gull)

The circumpolar group of Larus gull species has often been cited as a classic example of the ring species. The range of these gulls forms a ring around the North Pole. The European herring gull, which lives primarily in Great Britain and Northern Europe, can hybridize with the American herring gull (living in North America), which can also interbreed with the Vega or East Siberian gull, the western subspecies of which, Birula's gull, can hybridize with Heuglin's gull which, in turn, can interbreed with the Siberian lesser black-backed gull (all four of these live across the north of Siberia). The last is the eastern representative of the lesser black-backed gulls back in northwestern Europe, including Great Britain. However, the lesser black-backed gulls and herring gull are sufficiently different that they rarely interbreed; thus, the group of gulls forms a continuum except in Europe, where the two lineages meet. However, a recent genetic study has shown that this example is far more complicated than presented here, and probably does not constitute a true ring species. [9]

See also

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References

  1. Olsen, Klaus Malling; Larsen, Hans (2003). Gulls of Europe, Asia and North America. London: Helm. ISBN   978-0-7136-7087-5.
  2. Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 136.
  3. Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names . London: Christopher Helm. p.  219. ISBN   978-1-4081-2501-4.
  4. Selby, Prideaux John (1840). A Catalogue of the Generic and Sub-Generic Types of the Class Aves, Birds, Arranged According to the Natural System. Newcastle: T. and J. Hodgson. pp. 48–49.
  5. Peters, James Lee, ed. (1934). Check-List of Birds of the World. Vol. 2. Cambridge, Massachusetts: Harvard University Press. p. 313.
  6. Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Noddies, gulls, terns, skimmers, skuas, auks". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 16 August 2021.
  7. 1 2 Olson, Storrs L. (1985): Section X.D.2.j. Laridae. In: Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): Avian Biology8: 181-182. Academic Press, New York.
  8. (Olson, 1985).
  9. Liebers, Dorit; de Knijff, Peter & Helbig, Andreas J. (2004). "The herring gull complex is not a ring species". Proc. R. Soc. B . 271 (#1542): 893–901. doi:10.1098/rspb.2004.2679. PMC   1691675 . PMID   15255043. Supplemental material: Electronic appendices