Stilt bugs | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hemiptera |
Suborder: | Heteroptera |
Infraorder: | Pentatomomorpha |
Superfamily: | Lygaeoidea |
Family: | Berytidae Fieber, 1851 |
Synonyms [1] | |
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Berytidae is a family of the order Hemiptera ("true bugs"), commonly called stilt bugs [2] or thread bugs [3] (not to be confused with the thread-legged bugs, Emesinae). Most berytids are brown to yellow, with species that are plant sap feeders, a few being predaceous. About 200 species are known from all around the world and they are classified into three subfamilies.
The Berytidae are extremely gracile insects with legs so long and slender as to suggest common names such as "thread bugs" and "stilt bugs". In this they resemble the Emesinae, with which they are easily confused, though they are in different families. They may be distinguished most readily by the forelegs, that in the Emesinae are raptorial in a way resembling those of the Mantodea, Mantispidae and certain other invertebrate predators. In form and function the forelegs of the Berytidae are roughly similar to those of their other legs. Other differences are subtler and not fully consistent. For one thing, the antennae of most Berytidae though long, geniculate, and in other ways generally similar to Emesinae, tend to have a more or less obvious swelling at the tip. Some members of the family also have slight swellings at the distal ends of the femora of their legs, though in many species this is either absent or not obvious. The head often has a forward-facing protrusion between the antennae bases. The wing membrane has five veins and can be hard to examine in short-winged forms. Some species have spine like protrusions emerging at the base of the forewings which have a pore through which chemicals are secreted. Many species have split or toothed claws which apparently allow, along with the long legs, these bugs to overcome plant defenses involving stick hair on their surfaces. Many species are not purely plant-sap feeding and will opportunistically scavenge on insects trapped on the surfaces of sticky-haired plants. [4] They typically probe all surfaces and can inflict a painful prick on the human skin as well. [5]
The habits of most species are not well known. Most are believed to be sap-suckers like most other Hemiptera, but some also feed on mites and small insects. [6] [7] [8]
Three subfamilies are usually recognised: [6]
Southwood and Leston, 1959
Douglas and Scott, 1865
Hemiptera is an order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from 1 mm (0.04 in) to around 15 cm (6 in), and share a common arrangement of piercing-sucking mouthparts. The name "true bugs" is often limited to the suborder Heteroptera.
The Reduviidae is a large cosmopolitan family of the suborder Heteroptera of the order Hemiptera. Among the Hemiptera and together with the Nabidae almost all species are terrestrial ambush predators; most other predatory Hemiptera are aquatic. The main examples of non-predatory Reduviidae are some blood-sucking ectoparasites in the subfamily Triatominae, with a few species from South America noted for their ability to transmit Chagas disease. Though spectacular exceptions are known, most members of the family are fairly easily recognizable: they have a relatively narrow neck, sturdy build, and formidable curved proboscis. Large specimens should be handled with caution, if at all, because they sometimes defend themselves with a very painful stab from the proboscis.
Pyrrhocoridae is a family of insects with more than 300 species world-wide. Many are red coloured and are known as red bugs and some species are called cotton stainers because their feeding activities leave an indelible yellow-brownish stain on cotton crops. A common species in parts of Europe is the firebug, and its genus name Pyrrhocoris and the family name are derived from the Greek roots for fire "pyrrho-" and bug "coris". Members of this family are often confused with, but can be quickly separated from, Lygaeidae by the lack of ocelli on the top of the head.
Coreidae is a large family of predominantly sap-sucking insects in the Hemipteran suborder Heteroptera. The name "Coreidae" derives from the genus Coreus, which derives from the Ancient Greek κόρις (kóris) meaning bedbug.
The insect family Nabidae contains the damsel bugs. There are over 500 species in 20 genera. They are soft-bodied, elongate, winged terrestrial predators. Many damsel bugs catch and hold prey with their forelegs, similar to mantids. They are considered helpful species in agriculture because of their predation on many types of crop pests.
The Lygaeoidea are a sizeable superfamily of true bugs, containing seed bugs and allies, in the order Hemiptera. There are about 16 families and more than 4,600 described species in Lygaeoidea, found worldwide. Most feed on seeds or sap, but a few are predators.
The Emesinae, or thread-legged bugs, are a subfamily of the Reduviidae. They are conspicuously different from the other reduviids by their very slender body form. They are stalking, predatory insects that can be collected on palm fronds, cliffs, spider webbing, or near lights at night. They walk on their mid and hind legs; the front pair is raptorial. Some groups specialize on spiders. Very little is known about emesines except that many species are found in the tropics. Pedro Wygodzinsky wrote the most recent revision of this group.
Macrotylus is a genus of plant bugs in the family Miridae. There are at least 60 described species in Macrotylus.
Tytthus is a genus of insects in family Miridae, the plant bugs. They are carnivorous, feeding upon the eggs of various planthoppers in the family Delphacidae, and thus are important in the biological control of pests. The genus is distributed throughout the Holarctic of the Northern Hemisphere, but species are also found in the tropics, in China, South America, Australia, and the Indo-Pacific.
The Blissidae are a family in the Hemiptera, comprising nearly 50 genera and 400 species. The group has often been treated as a subfamily of the Lygaeidae but was resurrected as a full family by Thomas Henry (1997).
Orthops is a genus of plant bugs in the family Miridae. There are at least 30 described species in Orthops.
Macrolophus is a genus of plant bugs in the family Miridae. There are at least 20 described species in Macrolophus.
Jalysus is a genus of stilt bugs in the family Berytidae. There are about 12 described species in Jalysus.
Metacanthinae is a subfamily of stilt bugs in the family Berytidae. There are about 12 genera and 80 described species in Metacanthinae.
Hoplinus is a genus of stilt bugs in the family Berytidae. There are about six described species in Hoplinus.
Gampsocorinae is a subfamily of stilt bugs in the family Berytidae. There are about 60 described species in Gampsocorinae.
Yemma is a genus of stilt bugs in the family Berytidae. There are about five described species in Yemma.
Mecomma is a genus of plant bugs in the family Miridae. There are more than 30 described species in Mecomma.
Metacanthus is a genus of stilt bugs in the family Berytidae. There are more than 20 described species in Metacanthus.
Dufouriellus is a monotypic genus of minute pirate bugs in the tribe Dufouriellini. The described species is Dufouriellus ater, which has been recorded from much of western Europe through to Bohemia, Moravia, Slovakia, southern Scandinavia and including the British Isles.