Toad bugs Temporal range: | |
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Gelastocoridae at the edge of the Eel River, California | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hemiptera |
Suborder: | Heteroptera |
Infraorder: | Nepomorpha |
Clade: | Tripartita |
Superfamily: | Ochteroidea |
Family: | Gelastocoridae |
Genera | |
Synonyms | |
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The Gelastocoridae (toad bugs) is a family of about 100 species of insects in the suborder Heteroptera. These fall into two genera, about 15 species of Gelastocoris from the New World and 85 of Nerthra from the Old World. [1] They are reminiscent of toads both in the warty appearance and hopping movements of some species.
Gelastocoridae are riparian insects, generally found at the margins of streams and ponds, where they are predators of small insects. Gelastocoridae catch their prey by leaping on top of them and grasping them with their modified front legs. [2] Members of the family are found throughout the world, but their highest diversity is in the tropics. [3]
Adults lay their eggs in sand. [4] There are 5 instars in their life cycle. [5] Nymphs of many species cover themselves with a layer of sand grains. Apart from the no doubt considerable physical protection that the armour affords them, the layer of sand renders them effectively invisible on the ground unless they move at the wrong moment. [6]
Gelastocoridae are short (6–15 mm or 0.24–0.59 in long) and stout, with large protuberant eyes and cryptic coloration. Many Gelastocoridae species can change their coloration to match their habitat. [4] Like other Heteroptera, they have hemelytra for their forewings and piercing-sucking mouthparts. Their antennae are hidden. [2] Males in the genera Nerthra have stridulatory mechanisms in the genital capsule, however sound produced by stridulation has never been observed. [7]
The oldest record of the family is the genus Cratonerthra from the Aptian aged Crato Formation of Brazil. Fossils assignable to both extant genera are known from the Cenomanian aged Burmese amber of Myanmar. [8]
Hemiptera is an order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from 1 mm (0.04 in) to around 15 cm (6 in), and share a common arrangement of piercing-sucking mouthparts. The name "true bugs" is often limited to the suborder Heteroptera.
The Pentatomoidea are a superfamily of insects in the suborder Heteroptera of the order Hemiptera. As hemipterans, they possess a common arrangement of sucking mouthparts. The roughly 7000 species under Pentatomoidea are divided into 21 families. Among these are the stink bugs and shield bugs, jewel bugs, giant shield bugs, and burrower bugs.
The Reduviidae is a large cosmopolitan family of the suborder Heteroptera of the order Hemiptera. Among the Hemiptera and together with the Nabidae almost all species are terrestrial ambush predators; most other predatory Hemiptera are aquatic. The main examples of non-predatory Reduviidae are some blood-sucking ectoparasites in the subfamily Triatominae, with a few species from South America noted for their ability to transmit Chagas disease. Though spectacular exceptions are known, most members of the family are fairly easily recognizable: they have a relatively narrow neck, sturdy build, and formidable curved proboscis. Large specimens should be handled with caution, if at all, because they sometimes defend themselves with a very painful stab from the proboscis.
Pyrrhocoridae is a family of insects with more than 300 species world-wide. Many are red coloured and are known as red bugs and some species are called cotton stainers because their feeding activities leave an indelible yellow-brownish stain on cotton crops. A common species in parts of Europe is the firebug, and its genus name Pyrrhocoris and the family name are derived from the Greek roots for fire "pyrrho-" and bug "coris". Members of this family are often confused with, but can be quickly separated from, Lygaeidae by the lack of ocelli on the top of the head.
Scutelleridae is a family of true bugs. They are commonly known as jewel bugs or metallic shield bugs due to their often brilliant coloration. With the name based on the Asian genus Scutellera, they are also known as shield-backed bugs due to the enlargement of the thoracic scutellum into a continuous shield over the abdomen and wings. This latter characteristic distinguishes them from most other families within Heteroptera, and may lead to misidentification as a beetle rather than a bug. These insects feed on plant juices from a variety of different species, including some commercial crops. Closely related to stink bugs, they may also produce an offensive odour when disturbed. There are around 450 species worldwide.
The insect family Nabidae contains the damsel bugs. There are over 500 species in 20 genera. They are soft-bodied, elongate, winged terrestrial predators. Many damsel bugs catch and hold prey with their forelegs, similar to mantids. They are considered helpful species in agriculture because of their predation on many types of crop pests.
Notonectidae is a cosmopolitan family of aquatic insects in the order Hemiptera, commonly called backswimmers because they swim "upside down" (inverted). They are all predators and typically range from 0.5 to 1.5 cm (0.2–0.6 in) in length. They are similar in appearance to Corixidae, but can be separated by differences in their dorsal-ventral coloration, front legs, and predatory behavior. Their dorsum is convex, lightly colored without cross striations. Their front tarsi are not scoop-shaped and their hind legs are fringed for swimming. There are about 350 species in two subfamilies: Notonectinae with seven genera, and Anisopinae with four genera. Members in the former subfamily are often larger than those in the latter.
Enicocephaloidea is a sole superfamily within the infraorder Enicocephalomorpha of the hemipteran suborder Heteroptera. The group was formerly thought to be related to the Reduviidae because of similarities in head structure but they are now considered different enough from other Heteroptera to constitute a separate infraorder and represents a sister group of the Leptopodomorpha. They are predaceous and some species are known to swarm but little is known of their life history. A few species of enicocephalids are known to shed their wings before entering the surface of subsoil.
Nepomorpha is an infraorder of insects in the "true bug" order (Hemiptera). They belong to the "typical" bugs of the suborder Heteroptera. Due to their aquatic habits, these animals are known as true water bugs. They occur all over the world outside the polar regions, with about 2,000 species altogether. The Nepomorpha can be distinguished from related Heteroptera by their missing or vestigial ocelli. Also, as referred to by the obsolete name Cryptocerata, their antennae are reduced, with weak muscles, and usually carried tucked against the head.
The Lygaeidae are a family in the Hemiptera, with more than 110 genera in four subfamilies. The family is commonly referred to as seed bugs, and less commonly, milkweed bugs, or ground bugs. Many species feed on seeds, some on sap or seed pods, others are omnivores and a few, such as the wekiu bug, are insectivores. Insects in this family are distributed across the world.
Hydrometridae is a family of semiaquatic insects, known as marsh treaders or water measurers. They have a characteristic elongated head and body which makes them resemble a yardstick for measuring the water surface.
The Ochteridae comprise a small family of insects. Eight genera with about 80 species have been described. They occur worldwide along the shore of various types of water and the greatest diversity is in tropical regions. They are "true bugs", being members of the order Hemiptera, and are in the suborder Heteroptera. Ochteridae commonly are known as the velvety shore bugs. They resemble the Saldidae shore bugs and have lengths ranging from 4.5 to 9 mm (0.18–0.35 in).
The Tingidae are a family of very small insects in the order Hemiptera that are commonly referred to as lace bugs. This group is distributed worldwide with about 2,000 described species.
Coleorrhyncha or Peloridiomorpha, also known as moss bugs or beetle bugs, are a suborder of Hemiptera and represent an ancient lineage of moss-feeding insects. They show some similarities to the Heteroptera but have been considered distinct. It has a single extant family, the Peloridiidae. They are 2 to 5 millimetres in length, and feed on moss and liverworts. They have wings in some species which are reduced in others but all species are flightless and live in damp moss habitats and are associated with the distribution of Nothofagus trees in Australia, New Zealand, New Caledonia, and South America, which all were formerly part of the supercontinent Gondwana.
Tessaratomidae is a family of true bugs. It contains about 240 species of large bugs divided into 3 subfamilies and 56 genera.
Schizopteridae is the largest family in the infraorder Dipsocoromorpha and comprises 56 genera and approximately 255 species. Schizopterids are some of the smallest (0.5–2.0 mm) true bugs. Members of this family can be distinguished by their small size, enlarged forecoxae and varying degree of abdominal and genitalic asymmetry in males. Schizopteridae exhibit a wide range of simple and complex wing venation patterns. The group is currently divided into three subfamilies: Schizopterinae, Ogeriinae and Hypselosomatinae.
Nerthra fuscipes is a species of toad bug in the family Gelastocoridae. It is found in the Caribbean, Central America, North America, and South America.
Nerthra is a genus of toad bugs in the family Gelastocoridae. There are at least 90 described species in Nerthra.
Mesoveliidae is a family of water treaders in the order Hemiptera. There are about 16 genera and at least 50 described species in Mesoveliidae.
Gelastocoris oculatus, the big-eyed toad bug, is a species of toad bug in the family Gelastocoridae. It is found in Central America and North America. Due to their coloration they can be difficult to find. Females lay their eggs under rocks or in sand or mud and both the larva and adults are carnivorous: feeding on other insects along the shorelines.
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