Dictyopharidae Temporal range: | |
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Dictyophara species | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hemiptera |
Suborder: | Auchenorrhyncha |
Infraorder: | Fulgoromorpha |
Superfamily: | Fulgoroidea |
Family: | Dictyopharidae Spinola, 1839 |
Dictyopharidae is a family of planthoppers, related to the Fulgoridae. The family comprises nearly 760 species in more than 150 genera [1] which are grouped into two subfamilies, Dictyopharinae and Orgeriinae.
Like all other fulgoroids, they have the antennae arising on the side of the head below the compound eye (not between the eyes as in the Cicadoidea). Many species have an elongated frons. Those that do not have this elongation may have 2 or 3 carinae (keels). The median ocellus is absent. [2]
Genera are placed in two subfamilies:
Authority: Onuki, 1901; selected genera include:
Authority: Fieber, 1872; the following genera, in four tribes, are included by BioLib.cz: [3]
The following genera are incertae sedis :
A number of species are known from the fossil record, which reaches back to the Santonian age of the Late Cretaceous. The oldest fossil, Netutela annunciator belonging to the extinct dictyopharine tribe Netutelini, was described from Taymyr amber on the Taymyr Peninsula of Russia. [4] Younger amber fossils include the amber genus Alicodoxa described from Eocene Baltic and Rovno ambers [5] Compression fossil species include the Ypresian Limfjordia breineri from the Fur Formation in Denmark and the Priabonian Florissantia elegans from the Florissant Formation, Colorado. [4]
A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment and that they often "hop" for quick transportation in a similar way to that of grasshoppers. However, planthoppers generally walk very slowly. Distributed worldwide, all members of this group are plant-feeders, though few are considered pests. The infraorder contains only a single superfamily, Fulgoroidea. Fulgoroids are most reliably distinguished from the other Auchenorrhyncha by two features; the bifurcate (Y-shaped) anal vein in the forewing, and the thickened, three-segmented antennae, with a generally round or egg-shaped second segment (pedicel) that bears a fine filamentous arista.
The Cixiidae are a family of fulgoroid insects, one of many families commonly known as planthoppers, distributed worldwide and comprising more than 2,000 species from over 150 genera. The genera are placed into three subfamilies, Borystheninae, Bothriocerinae and Cixiinae with sixteen tribes currently accepted in Cixiinae.
Delphacidae is a family of planthoppers containing about 2000 species, distributed worldwide. Delphacids are separated from other "hoppers" by the prominent spur on the tibia of the hindleg.
Eurybrachidae is a small family of planthoppers with species occurring in parts of Asia, Australia and Africa. They are remarkable for the sophistication of their automimicry.
Derbidae is a family of insects in the order Hemiptera, the true bugs. It is one of the largest and most diverse families of planthoppers. It is widely distributed around the world, especially in the tropics, and with many species in subtropical and temperate regions.
Flatidae are a family of fulgoroid planthoppers. They are cosmopolitan in distribution and are distinguished from others in the superfamily by a combination of characters. Like all other planthoppers, they suck phloem sap of plants. Some species are known to communicate with vibrations through the plant stems. Communication may be with mates, or with ants that tend the nymphs, protecting them and gathering honeydew secretions. Adults of some species have brightly coloured forewings which are tougher and known as tegmina unlike the membranous hindwings which are used for flight. Although a few can be identified by their coloration, most species requires dissection and examination under a microscope with access to literature on already described species.
Glisachaemus is an extinct monotypic genus of planthopper in the Cixiidae subfamily Cixiinae and at present, it contains the single species Glisachaemus jonasdamzeni. The genus is solely known from the Early Eocene Baltic amber deposits in the Baltic Sea region of Europe.
Issidae is a family of planthoppers described by Spinola in 1839, belonging to the order Hemiptera, suborder Auchenorrhyncha superfamily Fulgoroidea.
Ordralfabetix is an extinct genus of planthoppers in the family Lophopidae and containing the single species Ordralfabetix sirophatanis. The species is known only from the Early Eocene, Ypresian stage Oise amber from the Quesnoy locality, Oise Department, France.
Nogodinidae is a family of planthoppers. They have membranous wings with delicate venation and can be confused with members of other Fulgoroid families such as the Issidae and Tropiduchidae. Some authors treat it as a subfamily of the Issidae. Some of their key features are a frons ("face") that is longer than wide and a reticulate wing venation. They are less than 2 cm long. The antenna arises well below the eye, has the base clubbed and flagellum unsegmented. The lateral ocelli are outside the margins of the face. The face has carinae on the edge. On the hind leg, the second tarsal segment has an apical spine arising from it. The tibia of the hind leg also has spines towards the tip. An important family character is found in the shape of the male genital structure, a style that is longer than broad. Most members of this family are forest species.
Caliscelidae is a family of planthoppers, sap-sucking insects that belong to the order Hemiptera, suborder Auchenorrhyncha and superfamily Fulgoroidea. They are somewhat anomalous and have often been included within the family Issidae. Studies made in 2013 of the phylogeny of the Issidae and other groups using molecular techniques support the treatment of the group as a separate family. Sexual dimorphism can be marked. Some members of the family are called piglet bugs due to the shape of their snout. A particularly aberrant genus described in 2011 from India, Formiscurra, has males that resemble ants.
Cixiinae is a planthopper subfamily in the family Cixiidae. It is one of three such subfamilies, the other two being the Bothriocerinae and the Borystheninae. While a few species had been tested in a larger study of the Fulgoroidea, neither the Cixiinae nor its tribes were analysed cladistically until 2002. Resolution of tribal relationships is incomplete and additional testing of the tribes with samples larger than one per tribe is needed.
Cixiini is a planthopper tribe in the family Cixiidae. This tribe is non-monophyletic.
Athysanini is a tribe of leafhoppers in the subfamily Deltocephalinae. The type genus of the tribe is Athysanus. The tribe has a cosmopolitan distribution. It is the largest tribe in the subfamily Deltocephalinae and has 228 genera and at least 1120 species.
Alicodoxa is an extinct genus of planthopper in the fulgoroid family Dictyopharidae. The genus contains a single described species, Alicodoxa rasnitsyni and is known from several Late Eocene fossils which were found in Europe.
Achilidae is a family of planthoppers, sometimes called "achilids" in the order Hemiptera. There are at least 520 described species in Achilidae.
Dictyopharinae is a subfamily of dictyopharid planthoppers in the family Dictyopharidae. There more than 100 genera and 500 described species in Dictyopharinae.
Dictyophara is the type genus of planthoppers belonging to the family Dictyopharidae and tribe Dictyopharini, containing five subgenera. The scientific genus name Dictyophara derives from the Greek and can be translated "who wears a net".
The Hysteropterinae are a subfamily of bugs in the family Issidae, based on the type genus Hysteropterum. Species in 59 genera have been recorded in Europe, Africa, and Asia. One speices, Agalmatium bilobum, is now also found as an invasive species in the United States, especially in California.
Florissantia is an extinct monotypic genus of planthopper in the dictyopharid subfamily Dictyopharinae. The single species, Florissantia elegans, was described by Samuel Hubbard Scudder (1890) from fossils found in the Florissant Formation of Colorado.