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Commelinales Temporal range: Late Cretaceous - Recent | |
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Tradescantia pallida | |
Scientific classification ![]() | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Commelinales Mirb. ex Bercht. & J.Presl [1] |
Families | |
Commelinaceae Contents | |
Synonyms | |
Haemodorales |
Commelinales is an order of flowering plants. It comprises five families: Commelinaceae, Haemodoraceae, Hanguanaceae, Philydraceae, and Pontederiaceae. All the families combined contain over 885 species in about 70 genera; the majority of species are in the Commelinaceae. Plants in the order share a number of synapomorphies that tie them together, such as a lack of mycorrhizal associations and tapetal raphides. Estimates differ as to when the Commelinales evolved, but most suggest an origin and diversification sometime during the mid- to late Cretaceous. Depending on the methods used, studies suggest a range of origin between 123 and 73 million years, with diversification occurring within the group 110 to 66 million years ago. The order's closest relatives are in the Zingiberales, which includes ginger, bananas, cardamom, and others. [2]
According to the most recent classification scheme, the APG IV of 2016, the order includes five families: [3]
This is unchanged from the APG III of 2009 and the APG II of 2003, but different from the older APG system of 1998, which did not include Hanguanaceae.
The older Cronquist system of 1981, which was based purely on morphological data, placed the order in subclass Commelinidae of class Liliopsida and included the families Commelinaceae, Mayacaceae, Rapateaceae and Xyridaceae. These families are now known to be only distantly related. [4] [1] In the classification system of Dahlgren the Commelinales were one of four orders in the superorder Commeliniflorae (also called Commelinanae), and contained five families, of which only Commelinaceae has been retained by the Angiosperm Phylogeny Group.
Canellales is the botanical name for an order of flowering plants, one of the four orders of the magnoliids. It is recognized by the most recent classification of flowering plants, the APG IV system. It is defined to contain two families: Canellaceae and Winteraceae, which comprise 136 species of fragrant trees and shrubs. The Canellaceae are found in tropical America and Africa, and the Winteraceae are part of the Antarctic flora. Although the order was defined based on phylogenetic studies, a number of possible synapomorphies have been suggested, relating to the pollen tube, the seeds, the thickness of the integument, and other aspects of the morphology.
Piperales is an order of flowering plants. It necessarily includes the family Piperaceae but other taxa have been included or disincluded variously over time. Well-known plants which may be included in this order include black pepper, kava, the many Peperomias, pepper elder, lizard's tail, birthwort, and wild ginger.
Ranunculales is an order of flowering plants. Of necessity it contains the family Ranunculaceae, the buttercup family, because the name of the order is based on the name of a genus in that family. Ranunculales belongs to a paraphyletic group known as the basal eudicots. It is the most basal clade in this group; in other words, it is sister to the remaining eudicots. Widely known members include poppies, barberries, hellebores, and buttercups.
Proteales is an order of flowering plants consisting of three families. The Proteales have been recognized by almost all taxonomists.
The Francoaceae are a small family of flowering plants in the order Geraniales, including the genera Francoa, commonly known as bridal wreaths, and Tetilla. The Francoaceae are recognized as a family under various classification schemes but under the APG III system the Francoaceae are included within the Melianthaceae. In the APG IV system the Francoaceae are again recognized as a family, with Melianthaceae included in the circumscription of Francoaceae.
The Aquifoliales are an order of flowering plants, including the Aquifoliaceae (holly) family, and also the Helwingiaceae and the Phyllonomaceae. In 2001, the families Stemonuraceae and Cardiopteridaceae were added to this order. This circumscription of Aquifoliales was recognized by the Angiosperm Phylogeny Group when they published the APG II system in 2003. In the Cronquist system, there is no Aquifoliales order: the Aquifoliaceae are placed within the order Celastrales and the others are in other families.
The Zygophyllales are an order of dicotyledonous plants, comprising the following two families:
The Xyridaceae are a family of flowering plants. This family has been recognized by many taxonomists and is known as the yellow-eyed grass family.
Haemodoraceae is a family of perennial herbaceous flowering plants with 14 genera and 102 known species. It is sometimes known as the "bloodroot family". Primarily a Southern Hemisphere family, they are found in South Africa, Australia and New Guinea, and in the Americas. Perhaps the best known are the widely cultivated and unusual kangaroo paws from Australia, of the two closely related genera Anigozanthos and Macropidia.
Pontederiaceae is a family of flowering plants.
Saururaceae is a plant family comprising four genera and seven species of herbaceous flowering plants native to eastern and southern Asia and North America. The family has been recognised by most taxonomists, and is sometimes known as the "lizard's-tail family". The APG IV system assigned it to the order Piperales in the clade magnoliids.
In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade. Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total flowering plant species. Well-known plants in this clade include the common daisy, forget-me-nots, nightshades, the common sunflower, petunias, yacon, morning glory, lettuce, sweet potato, coffee, lavender, lilac, olive, jasmine, honeysuckle, ash tree, teak, snapdragon, sesame, psyllium, garden sage, table herbs such as mint, basil, and rosemary, and rainforest trees such as Brazil nut.
Myrothamnus is a genus of flowering plants, consisting of two species of small xerophytic shrubs, in the southern parts of tropical Africa and in Madagascar. Myrothamnus is recognized as the only genus in the family Myrothamnaceae.
In plant taxonomy, commelinids is a clade of flowering plants within the monocots, distinguished by having cell walls containing ferulic acid.
Hanguana is a genus of flowering plants with a dozen known species. It is the only genus in the family Hanguanaceae.
Philydraceae is a family of flowering plants composed of three genera and a total of six known species. Such a family has not been recognized by many taxonomists.
Tecophilaeaceae is a family of flowering plants, placed in the order Asparagales of the monocots. It consists of nine genera with a total of 27 species.
Huaceae is a family of plant in the rosids group, which has been classed in the orders Malpighiales, Malvales, and Violales or in its own order Huales. The APG II system placed it in the clade eurosids I, whereas the APG III system of 2009 and APG IV (2016) place it within the Oxalidales. The family is endemic to central Africa. It contains four species in the following two genera:
Icacinales is an order of Angiosperms.