Common myna | |
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Kokrebellur, India | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Family: | Sturnidae |
Genus: | Acridotheres |
Species: | A. tristis |
Binomial name | |
Acridotheres tristis (Linnaeus, 1766) | |
Subspecies | |
Acridotheres tristis melanosternus Contents
| |
Native distribution in blue, introduced in red | |
Synonyms | |
Paradisaea tristisLinnaeus, 1766 |
The common myna or Indian myna (Acridotheres tristis), sometimes spelled mynah, [2] is a bird in the family Sturnidae, native to Asia. An omnivorous open woodland bird with a strong territorial instinct, the common myna has adapted extremely well to urban environments.
The range of the common myna is increasing at such a rapid rate that in 2000 the IUCN Species Survival Commission declared it one of the world's most invasive species and one of only three birds listed among "100 of the World's Worst Invasive Species" that pose a threat to biodiversity, agriculture and human interests. [3] In particular, the species poses a serious threat to the ecosystems of Australia, where it was named "The Most Important Pest/Problem" in 2008. [4]
In 1760, the French zoologist Mathurin Jacques Brisson included a description of the common myna in his Ornithologie, based on a specimen that he mistakenly believed had been collected in the Philippines. He used the French name Le merle des Philippines and the Latin Merula Philippensis. [5] Although Brisson coined Latin names, they do not conform to the binomial system and are not recognised by the International Commission on Zoological Nomenclature. [6]
When the Swedish naturalist, Carl Linnaeus, updated his Systema Naturae in 1766, for the 12th edition, he added 240 species that had been previously described by Brisson. [6] One of them was the common myna. Linnaeus included a brief description, coined the binomial name Paradisea tristis and cited Brisson's work. [7] The type location was subsequently corrected to Pondicherry in southern India. [8] The specific name tristis is Latin for "sad" or "gloomy". [9] This species is now placed in the genus Acridotheres that was introduced by the French ornithologist Louis Pierre Vieillot in 1816. [10] The generic name Acridotheres is from the Greek ακριδος (akridos), meaning locust, and θηρας (theras), meaning hunter.
Two subspecies are recognised: [11]
The Sri Lankan subspecies melanosternus is darker than the Indian subspecies tristis and has half-black and half-white primary coverts and a larger yellow cheek-patch. [13] [14]
The common myna is readily identified by the brown body, black hooded head and the bare yellow patch behind the eye. The bill and legs are bright yellow. They have rounded wings as well, and a round square tipped tail. There is a white patch on the outer primaries and the wing lining on the underside is white, as well as having a white tail tip. The sexes are similar and birds are usually seen in pairs. [15] The common myna obeys Gloger's rule in that the birds from northwestern India tend to be paler than their darker counterparts in southern India. [13] [14]
The calls includes croaks, squawks, chirps, clicks, whistles and 'growls', and the bird often fluffs its feathers and bobs its head in singing. The common myna screeches warnings to its mate or other birds in cases of predators in proximity or when it is about to take off flying. [16] Common mynas are popular as cage birds for their singing and "speaking" abilities. Before sleeping in communal roosts, common mynas vocalise in unison, which is known as "communal noise". [17]
Morphometry. [13]
Parameter/sex | Male | Female |
---|---|---|
Average weight (g) | 109.8 | 120–138 |
Wing chord (mm) | 138–153 | 138–147 |
Bill (mm) | 25–30 | 25–28 |
Tarsus (mm) | 34–42 | 35–41 |
Tail (mm) | 81–95 | 79–96 |
The common myna is native to Asia, with its initial home range spanning Iran, Pakistan, India, Nepal, Bhutan, Bangladesh and Sri Lanka, Afghanistan, Uzbekistan, Tajikistan, Turkmenistan, Myanmar, Malaysia, Singapore, peninsular Thailand, Indochina, Japan (both mainland Japan and the Ryukyu Islands) and China. [13] [18]
The common myna has been introduced to many other parts of the world such as Canada, Australia, Israel, New Zealand, New Caledonia, Fiji, the United States (South Florida only [19] ), South Africa, Kazakhstan, Kyrgyzstan Uzbekistan, the Cayman Islands, islands in the Indian Ocean (the Seychelles – from which it was subsequently eradicated at great expense, [20] Mauritius, Réunion, Madagascar, the Maldives, the Andaman and Nicobar Islands and the Lakshadweep archipelago) and also in islands of the Atlantic (such as Ascension and Saint Helena, Pacific Ocean and Cyprus February 2022. [13] [ pages needed ] The range of the common myna is increasing to the extent that in 2000 the IUCN Species Survival Commission declared it among 100 of the world's worst invasive species. [3]
It is typically found in open woodland, cultivation and around habitation. Although it is an adaptable species, its population is abnormal and very much considered a pest in Singapore (where it is locally called as gembala kerbau, literally 'buffalo shepherd') due to competition with the related introduced Javan myna. [21]
The common myna thrives in urban and suburban environments; in Canberra, for instance, 110 common mynas were released between 1968 and 1971. By 1991, common myna population density in Canberra averaged 15 birds per square kilometer. [22] Only three years later, a second study found an average population density of 75 birds per square kilometer in the same area. [23]
The bird likely owes its success in the urban and suburban settings of Sydney and Canberra to its evolutionary origins; having evolved in the open woodlands of India, the common myna is pre-adapted to habitats with tall vertical structures and little to no vegetative ground cover, [24] features characteristic of city streets and urban nature preserves.
The common myna (along with common starlings, house sparrows, and feral rock doves) is a nuisance to city buildings; its nests block gutters and drainpipes, causing water damage to building exteriors. [25]
Common mynas are believed to pair for life. They breed through much of the year depending on the location, building their nest in a hole in a tree or wall. They breed at elevations of 0–3,000 m (0–9,843 ft) in the Himalayas. [13]
The normal clutch size is 4–6 eggs. The average size of the egg is 30.8 by 21.99 millimetres (1+1⁄4 in × 3⁄4 in). The incubation period is 17 to 18 days and fledging period is 22 to 24 days. [13] The Asian koel is sometimes brood parasitic on this species. [26] Nesting material used by common mynas includes twigs, roots, tow and rubbish. Common mynas have been known to use tissue paper, tin foil and sloughed off snake-skin. [13]
During the breeding season, the daytime activity-time budget of the common myna in Pune in April to June 1978 has been recorded to comprise the following: nesting activity (42%), scanning the environment (28%), locomotion (12%), feeding (4%), vocalisation (7%) and preening-related activities, interactions and other activities (7%). [27]
The common myna uses the nests of woodpeckers, parakeets, etc. and easily takes to nest boxes; it has been recorded evicting the chicks of previously nesting pairs by holding them in the beak and later sometimes not even using the emptied nest boxes. This aggressive behaviour contributes to its success as an invasive species. [28]
There is also some evidence that shows that in introduced environments, the species chooses to nest in more modified and artificial structures than in natural tree cavities when compared to native species. [29]
Like most starlings, the common myna is omnivorous. It feeds on insects, grubs, earthworms, arachnids, crustaceans, reptiles, small mammals, seeds, grain, fruits, flower nectar and petals, and discarded waste from human habitation. [30] It forages on the ground among grass for insects, and especially for grasshoppers, from which it gets the generic name Acridotheres, "grasshopper hunter". It, however, feeds on a wide range of insects, mostly picked from the ground. [13] [31] It is a cross-pollinator of flowers such as Salmalia and Erythrina . It walks on the ground with occasional hops and is an opportunistic feeder on the insects disturbed by grazing cattle as well as fired grass fields. [13] They prey on eggs and young of other birds, such as Hawaiʻi ʻakepas (Loxops coccineus). [30] They sometimes even wade in shallow waters to catch fish. [30] Living in close proximity to human-made habitats, common mynas may also appear near roadsides to feed on roadkill. [32]
Common mynas roost communally throughout the year, either in pure or mixed flocks with jungle mynas, rosy starlings, house crows, jungle crows, cattle egrets and rose-ringed parakeets and other birds. The roost population can range from less than one hundred to thousands. [33] [34] The time of arrival of mynas at the roost starts before and ends just after sunset. The mynas depart before sunrise. The time and timespan of arrival and departure, time taken for final settlement at the roost, duration of communal sleep, flock size and population vary seasonally. [17] [35] [36]
The function of communal roosting is to synchronise various social activities, avoid predators, exchange information about food sources. [37]
Communal displays (pre-roosting and post-roosting) consist of aerial maneuvers which are exhibited in the pre-breeding season (November to March). It is assumed that this behaviour is related to pair formation. [38]
The IUCN declared the common myna as one of only three birds among the world's 100 worst invasive species [3] (the other two being the red-vented bulbul and the common starling). The French introduced it in the 18th century from Pondicherry to Mauritius with the aim of controlling insects, even levying a fine on anyone persecuting the bird. [39] It has since been introduced widely elsewhere, including adjacent areas in Southeast Asia, Madagascar, [40] the Middle East, South Africa, the United States, Argentina, Germany, Spain and Portugal, [41] the United Kingdom, Australia, New Zealand and various oceanic islands in the Indian and Pacific Oceans, including prominent populations in Fiji and Hawaii. [18] [42]
The common myna is regarded as a pest in South Africa, North America, the Middle East, Australia, New Zealand and many Pacific islands. It is particularly problematic in Australia. [43] Several methods have been tried to control the bird's numbers and protect native species. [44]
In Australia, the common myna is an invasive pest. They are often the predominant bird in urban areas along the whole east coast. In a 2008 popular vote, the bird was named "The Most Important Pest/Problem" in Australia. They have earned the nickname "flying rats", due to their numbers and their scavenging behaviour. They are also known as "the cane toad of the sky". [4] However, there is little scientific consensus concerning the extent of its impact on native species. [45] [46]
The common myna was first introduced to Australia between 1863 and 1872, in Victoria, to control insects in the market gardens of Melbourne. The bird is likely to have spread to New South Wales (where it is currently most populous) at around the same time, but documentation is uncertain. [47] The bird was later introduced to Queensland as a predator of grasshoppers and cane beetles. Common myna populations in Australia are now concentrated along the eastern coast around Sydney and its surrounding suburbs, [48] with sparser populations in Victoria and a few isolated communities in Queensland. [49] During 2009 several municipal councils in New South Wales began trials of catching myna birds in an effort to reduce numbers. [50]
The myna can live and breed in a wide range of temperatures, ranging from the frosty winters of Canberra to the tropical climate of Cairns. Self-sustaining populations have been found in regions with a mean monthly highest temperature no less than 23.2 °C (73.8 °F) and a mean monthly lowest temperature no less than −0.4 °C (31.3 °F), implying that the common myna could spread from Sydney northwards along the eastern coast to Cairns, and westwards along the southern coast to Adelaide, but not to Tasmania, Darwin or the arid outback regions. [49]
In 2019, common mynas were added to the List of Invasive Alien Species of Union concern. [51] They have established in Spain and Portugal [52] and were introduced to France, where they occasionally bred. [53]
The common myna was introduced to both the North Island and South Island of New Zealand in the 1870s. However, the cooler summer temperatures in the South Island appear to have impeded the breeding success rate of the southern populations, preventing the proliferation of the species, which was largely non-existent there by the 1890s. In contrast, the North Island population was able to breed more successfully and large portions of the North Island are now populated. However, in the southern reaches of the North Island, the cooler summer temperatures, like those of the South Island, have prevented the establishment of large myna populations. [54] [32] Since the 1950s, mynas have spread northwards and presently inhabit beyond the Waikato region, [55] leading to a majority of its successful population thriving upon lower latitude regions due to the warmer climate. [55] At present, mynas have become especially common in regions of lower latitude, particularly the Northland region, [32] but rarely found south of Whanganui. [56]
In South Africa where it escaped into the wild in 1902, it has become very common and its distribution is greater where human populations are greater or where there is more human disturbance. [57] The bird is also notorious for being a pest, kicking other birds out of their nests and killing their young due to the myna's strong territorial instinct. In South Africa it is considered somewhat of a major pest and disturbance of the natural habitat; as a result, it has been declared an invasive species, [58] requiring it to be controlled.
Morphological studies show that the process of spatial sorting is at work on the range expansion of A. tristis in South Africa. [59] Dispersal-relevant traits are significantly correlated with distance from the range core, with strong sexual dimorphism, indicative of sex-biased dispersal. Morphological variations are significant in wing and head traits of females, suggesting females as the primary dispersing sex. In contrast, traits not related to dispersal such as those associated with foraging show no signs of spatial sorting but are significantly affected by environmental variables such as vegetation and intensity of urbanisation.
In Hawaii, it is out-competing many native birds for food and nesting areas. [60]
To study the invasion genetics and landscape-scale dynamics of A. tristis, scientists have recently developed 16 polymorphic nuclear microsatellite markers [61] using the next generation sequencing (NGS) approach.
The common myna is a hollow-nesting species; that is, it nests and breeds in protected hollows found either naturally in trees or artificially on buildings (for example, recessed windowsills or low eaves). [62] Compared to native hollow-nesting species, the common myna is extremely aggressive, and breeding males will actively defend areas ranging up to 0.83 hectares in size (though males in densely populated urban settings tend to only defend the area immediately surrounding their nests). [63]
This aggressiveness has enabled the common myna to displace many breeding pairs of native hollow-nesters, thereby reducing their reproductive success. In Australia, their aggressiveness has enabled them to chase native birds as large as galahs out of their nests.
The common myna is also known to maintain up to two roosts simultaneously; a temporary summer roost close to a breeding site (where the entire local male community sleeps during the summer, the period of highest aggression), and a permanent all-year roost where the female broods and incubates overnight. Both male and female common mynas will fiercely protect both roosts at all times, leading to further exclusion of native birds. [63]
The common myna (which feeds mostly on ground-dwelling insects, tropical fruits such as grapes, plums and some berries and, in urban areas, discarded human food) [64] poses a serious threat to Australian blueberry crops, though its main threat is to native bird species. [65]
In Hawaii, where the common myna was introduced to control pest armyworms and cutworms in sugarcane crops, the bird has helped to spread the robust Lantana camara weed across the islands' open grasslands. [66] It also has been recorded as the fourth-ranking avian pest in the fruit industry by a 2004 survey of the Hawaiian Farm Bureau and the sixth in number of complaints of avian pests overall. [67]
Common mynas can cause considerable damage to ripening fruit, particularly grapes, but also figs, apples, pears, strawberries, blueberries, guava, mangoes and breadfruit. Cereal crops such as maize, wheat and rice are susceptible where they occur near urban areas. Roosting and nesting commensal with humans create aesthetic and health concerns. Common mynas are known to carry avian malaria and exotic parasites such as the Ornithonyssus bursia mite, which can cause dermatitis in humans. The common myna can help spread agricultural weeds: for example, it spreads the seeds of Lantana camara, which has been classed as a Weed of National Significance because of its invasiveness. Common mynas are regularly observed to usurp nests and hollows, destroy the eggs and kill the young of native bird species, including seabirds and parrots. There is evidence that common mynas have killed small land mammals such as mice, squirrels and possums, but further research on these occurrences is under consideration. [68]
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The common myna, being a major agricultural pest and posing a threat to native species in non-native countries, is controlled by various factors. Mynas are either killed or chased away as control. Poison, [55] shooting, [55] cage traps, [55] and bird-scaring devices [55] are currently used for control.
In Sanskrit literature, the common myna has a number of names, most are descriptive of the appearance or behaviour of the bird. In addition to saarika, the names for the common myna include kalahapriya, which means "one who is fond of arguments" referring to the quarrelsome nature of this bird; chitranetra, meaning "picturesque eyes"; peetanetra (one with yellow eyes) and peetapaad (one with yellow legs). [69]
The bird called śārikā (Sanskrit : सारिक) [a] seems to refer to the common myna, [70] [b] though there are other candidates.
The house crow, also known as the Indian, greynecked, Ceylon or Colombo crow, is a common bird of the crow family that is of Asian origin but now found in many parts of the world, where they arrived assisted by shipping. It is between the jackdaw and the carrion crow in size but is slimmer than either. The forehead, crown, throat and upper breast are a richly glossed black, whilst the neck and breast are a lighter grey-brown in colour. The wings, tail and legs are black. There are regional variations in the thickness of the bill and the depth of colour in areas of the plumage.
Starlings are small to medium-sized passerine birds in the family Sturnidae, common name of Sturnid. The Sturnidae are named for the genus Sturnus, which in turn comes from the Latin word for starling, sturnus. The family contains 128 species which are divided into 36 genera. Many Asian species, particularly the larger ones, are called mynas, and many African species are known as glossy starlings because of their iridescent plumage. Starlings are native to Europe, Asia, and Africa, as well as northern Australia and the islands of the tropical Pacific. Several European and Asian species have been introduced to these areas, as well as North America, Hawaii, and New Zealand, where they generally compete for habitats with native birds and are considered to be invasive species. The starling species familiar to most people in Europe and North America is the common starling, and throughout much of Asia and the Pacific, the common myna is indeed common.
The mynas are a group of birds in the starling family (Sturnidae). This is a group of passerine birds which are native to Iran and Southern Asia, especially Afghanistan, India, Pakistan, Bangladesh, Nepal and Sri Lanka. Several species have been introduced to areas like North America, Australia, South Africa, Fiji and New Zealand, especially the common myna, which is often regarded as an invasive species. It is often known as "Selarang" and "Teck Meng" in Malay and Chinese respectively in Singapore, due to their high population there.
The common starling, also known as the European starling in North America and simply as the starling in Great Britain and Ireland, is a medium-sized passerine bird in the starling family, Sturnidae. It is about 20 cm (8 in) long and has glossy black plumage with a metallic sheen, which is speckled with white at some times of the year. The legs are pink and the bill is black in winter and yellow in summer; young birds have browner plumage than the adults. It is a noisy bird, especially in communal roosts and other gregarious situations, with an unmusical but varied song. Its gift for mimicry has been noted in literature including the Mabinogion and the works of Pliny the Elder and William Shakespeare.
The palmchat is a small, long-tailed passerine bird, the only species in the genus Dulus and the family Dulidae endemic to the Caribbean island of Hispaniola. It is related to the waxwings, family Bombycillidae. Its name reflects its strong association with palms for feeding, roosting, and nesting.
The red-vented bulbul is a member of the bulbul family of passerines. It is a resident breeder across the Indian subcontinent, including Bangladesh, Sri Lanka extending east to Burma and parts of Bhutan and Nepal. It has been introduced in many other parts of the world and has established itself in New Zealand, Argentina, Tonga and Fiji, as well as parts of Samoa, Australia, USA and Cook Islands. It is included in the list of the world's 100 worst invasive alien species.
The jungle myna is a myna, a member of the starling family. It is found patchily distributed across much of the mainland of the Indian Subcontinent but absent in the arid zones of India. It is easily recognized by the tuft of feathers on its forehead that form a frontal crest, a feature also found in the closely related Javan myna and the pale-bellied myna which were treated as a subspecies in the past. The eyes are pale, yellow or blue depending on the population and the base of the orange-yellow bill is dark. It has also been introduced either intentionally or accidentally into many other parts of the world including Fiji, Taiwan, the Andaman Islands, and parts of Japan. The species has also spread out on its own to some islands in the Pacific.
The common hill myna, sometimes spelled "mynah" and formerly simply known as the hill myna or myna bird, is the myna most commonly sighted in aviculture, where it is often simply referred to by the latter two names. It is a member of the starling family (Sturnidae), resident in hill regions of South Asia and Southeast Asia. The Sri Lanka hill myna, a former subspecies of G. religiosa, is now generally accepted as a separate species G. ptilogenys. The Enggano hill myna and Nias hill myna are also widely accepted as specifically distinct, and many authors favor treating the southern hill myna from the Nilgiris and elsewhere in the Western Ghats of India as a separate species.
The brahminy starling or brahminy myna is a member of the starling family of birds. It is usually seen in pairs or small flocks in open habitats on the plains of the Indian subcontinent.
The yellow-billed oxpecker is a passerine bird in the family Buphagidae. It was previously placed in the starling and myna family, Sturnidae.
The black drongo is a small Asian passerine bird of the drongo family Dicruridae. It is a common resident breeder in much of tropical southern Asia from southwest Iran through Pakistan, India, Bangladesh and Sri Lanka east to southern China and Indonesia and accidental visitor of Japan. It is an all black bird with a distinctive forked tail and measures 28 cm (11 in) in length. It feeds on insects, and is common in open agricultural areas and light forest throughout its range, perching conspicuously on a bare perch or along power or telephone lines.
The crested myna, also known as the Chinese starling, is a species of starling in the genus Acridotheres native to southeastern China and Indochina. It is named after the tuft of feathers on its forehead that resembles a crest.
The Indian pied myna is a species of starling found in the Indian subcontinent. It is usually found in small groups mainly on the plains and low foothills. It is often seen within cities and villages although it is not as bold as the common myna. It produces a range of calls made up of liquid notes.
The baya weaver is a weaverbird found across the Indian Subcontinent and Southeast Asia. Flocks of these birds are found in grasslands, cultivated areas, scrub and secondary growth and they are best known for their hanging retort shaped nests woven from leaves. These nest colonies are usually found on thorny trees or palm fronds and the nests are often built near water or hanging over water where predators cannot reach easily. They are widespread and common within their range but are prone to local, seasonal movements mainly in response to rain and food availability.
The bank myna is a myna found in the northern parts of South Asia. It is smaller but similar in colouration to the common myna, only differing in having brick-red naked skin behind the eyes instead of yellow. It is greyer on the underside and in this and in the presence of a slight tuft of feathers bears some resemblance to the jungle myna. They are found in flocks on the plains of northern and central India, often within towns and cities. Their range appears to be extending southwards into India. The name is derived from their habit of nesting almost exclusively in the earthen banks of rivers, where they excavate burrows and breed in large colonies.
The black-winged myna is a species of starling in the family Sturnidae. The species is also known as the black-winged starling or the white-breasted starling. It is endemic to Indonesia. There are three recognised subspecies: the nominate race, which occurs across much of the island of Java; tricolor, which is restricted to south east Java; and tertius, which is found on Bali and possibly Lombok. The validity of the records on Lombok has been called into question, as there are only a few records and those may represent escapees from the caged-bird trade or natural vagrants. The species has often been assigned to the starling genus Sturnus, but is now placed in Acridotheres because it is behaviourally and vocally closer to the birds in that genus.
The Tahiti monarch, or Tahiti flycatcher, is a rare species of bird in the monarch flycatcher family. It is endemic to Tahiti in French Polynesia. There are between 25 and 100 individuals remaining with an increasing population trend. Adults boast a striking black plumage with pale blue pills and feet. Recognized for its melodious flute-like song and distinctive "tick-tick-tick" call, it thrives in dense forest, primarily inhabiting canopy and understory amidst native mara trees. Despite facing threats from introduces predators such as ship rats and invasive bird species, conservation efforts have been implemented to safeguard its population.
Starlicide or gull toxicant is a chemical avicide that is highly toxic to European starlings and gulls, but less toxic to other birds or to mammals such as humans and pets.
The Ascension scrub and grasslands ecoregion covers the dormant volcano, Ascension Island in the Atlantic Ocean. As well as shrubs and grasses wildlife on the island includes a range of unique flora and fauna. In particular the surrounding islets are important havens for many seabirds. However the seabird populations on Ascension Island itself have been severely affected by introduced species, especially cats, which were the subject of an eradication campaign between 2002 and 2006.
The Canberra Indian Myna Action Group (CIMAG) is a voluntary public action group started in Canberra in 2006 to combat the invasive Indian myna that since its introduction to Canberra in 1968 had become the third most prevalent bird species, competing with and crowding out native birds. In November 2013 the action group won the 2014 Keep Australia Beautiful Award for 'Environmental Innovation and Protection'. As of July 2014, the action group has captured and humanely disposed an estimated 49,000 Indian mynas. The prevalence of the Indian myna in Canberra has been reduced to the sixteenth most prevalent species, with anecdotal evidence of increased observations and nesting by native bird species. The action group has been associated with scientific research in Canberra into the effect of Indian mynas on native bird species and whether controlling Indian myna populations affects the prevalence of native bird species.
No introduced species in Hawaii has elicited so much opinion as the Common Myna, perhaps in part due to its intelligence and amusing anthropomorphic qualities. Although they were thought to be of "great value to the aviculturist" in Hawaii for controlling pests (Bryan 1937b), it was also generally vilified for its noisy habits, "quarrelsome" and opportunistic nature, disturbance to domestic pigeons, fruit-eating and nest-robbing habits, and the possibility of its adversely affecting native bird populations (e.g., Finsch 1880; Wilson 1890a; Rothschild 1900; Perkins 1901, in Evenhuis 2007:75)
..Thurdus Salica, F. (B.) Hamilton, MS. I., p. 12 / Paradise Grackle, Latham. / The Common Hill Mynah.