Rose-ringed parakeet

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Rose-ringed parakeet or ringneck parrot
Rose-ringed parakeet (Psittacula krameri borealis) male Jaipur 2.jpg
Male P. k. borealis
Rose-ringed Parakeet (Psittacula krameri)- Female on a Neem (Azadirachta indica) tree at Hodal Iws IMG 1279.jpg
Female
Call
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Psittaciformes
Family: Psittaculidae
Genus: Psittacula
Species:
P. krameri
Binomial name
Psittacula krameri
(Scopoli, 1769)
Rose ringed parakeet range.PNG
Original (wild) range
Synonyms
  • Alexandrinus krameri (valid)

The rose-ringed parakeet (Psittacula krameri), also known as the ringneck parrot (in aviculture) or the Kramer parrot, is a medium-sized parrot in the genus Psittacula, of the family Psittacidae. It has disjunct native ranges in Africa and the Indian Subcontinent, [2] and is now introduced into many other parts of the world where feral populations have established themselves and are bred for the exotic pet trade.

Contents

One of the few parrot species that have successfully adapted to living in disturbed habitats, it has withstood the onslaught of urbanisation and deforestation. As a popular pet species, escaped birds have colonised a number of cities around the world, including populations in Northern and Western Europe. [3] These parakeets have also proven themselves capable of living in a variety of climates outside their native range, and are able to survive low winter temperatures in Northern Europe. [3] [4] The species is listed as least concern by the International Union for Conservation of Nature (IUCN) because its population appears to be increasing, but its popularity as a pet and unpopularity with farmers have reduced its numbers in some parts of its native range. [1]

Taxonomy

Four subspecies are recognised, though they differ little:

P. k. manillensis, Sri Lanka Rose-ringed parakeet (Psittacula krameri manillensis).jpg
P. k. manillensis, Sri Lanka
Rose-ringed parakeet near Chandigarh Rose-ringed parakeet near Chandigarh.jpg
Rose-ringed parakeet near Chandigarh

The Asian subspecies are both larger than the African subspecies. [5]

The genus name Psittacula is a diminutive of Latin psittacus, "parrot", and the specific krameri commemorates the Austrian naturalist Wilhelm Heinrich Kramer. [7]

In 2019, a genetic study revived the genus Alexandrinus , formerly viewed as a synonym of the current genus Psittacula . Some organisations, including the IUCN, have accepted the new taxonomy. [8]

Description

The rose-ringed parakeet is sexually dimorphic. The adult male sports a pink and black neck ring, [9] and the hen and immature birds of both sexes either show no neck rings, or display shadow-like pale to dark grey neck rings. Both sexes have a distinctive green colour in the wild with a red beak and blue tail, [9] and captive bred ringnecks have multiple colour mutations which include turquoise, cinnamon, olive, white, blue, violet, grey and yellow. Rose-ringed parakeets measure on average 40 cm (16 in) in length, including the tail feathers, a large portion of their total length. Their average single-wing length is about 15 to 17.5 cm (5.9 to 6.9 in). In the wild, this is a noisy species with an unmistakable squawking call. Captive individuals can be taught to speak. They are a herbivorous and non-migratory species.

Distribution

Rose-ringed parakeets in Garaboli National Park making a beak-lock - a common act in parakeet pairs Rose ringed parakeet.JPG
Rose-ringed parakeets in Garaboli National Park making a beak-lock – a common act in parakeet pairs

Since the 19th century, the rose-ringed parakeet has successfully colonised many other countries. It breeds further north than any other parrot species. It has established itself on a large scale in Germany, France, Belgium, the Netherlands, Italy, and the UK. [10] See Feral Birds section below.

The analyses show that the risk of parakeet establishment may rise further as a result of decreasing frost days due to global warming, rising urbanization, and rising human populations. Because of the significant separate parakeet imports in Europe, researchers are capable of investigating the widely held hypothesis of climate matching and human activity at the species level. [11]

Ecology and behaviour

Diet

In the wild, rose-ringed parakeets usually feed on buds, fruits, vegetables, nuts, berries, and seeds. Wild flocks also fly several miles to forage in farmlands and orchards, causing extensive damage. Feral parakeets will regularly visit gardens and other locations near human habitation, taking food from bird feeders.

In India, they feed on cereal grains, and during winter also on pigeon peas. [12] In Egypt during the spring, they feed on mulberry and in summer they feed on dates and nest inside palm trees and eat from sunflower and corn fields.

In captivity, rose-ringed parakeets will take a large variety of food and can be fed on a number of fruits, vegetables, pellets, seeds, and even small amounts of cooked meat for protein. Oils, salts, chocolate, alcohol, and other preservatives should be avoided. [13] [14]

Reproduction

In northwestern India, Indian rose-ringed parakeets form pairs from September to December. They do not have life mates and often breed with another partner during the following breeding season. During this cold season, they select and defend nest sites, thus avoiding competition for sites with other birds. Feeding on winter pea crops provides the female with nutrients necessary for egg production. From April to June, they care for their young. Fledglings are ready to leave the nest before monsoon. [15]

Seasonal changes in testicular activity, plasma luteinizing hormone (LH), estradiol (E2), testosterone (T), and 5 α-dihydrotestosterone (5 α-DHT) were related to pair bond formation, nest building, nest defense, and parental behavior in free living Indian rose-ringed parakeets in northwest India. [15] The parakeets are able to reproduce in the winter because it allows them to avoid competing with other birds for nesting places, postpone having young during the monsoon season, and take use of the winter pea harvest, which provides the female with extra nutrients for egg formation. [15]

Aviculture

A blue colour morph mutation parakeet kept as a pet Rose-ringed Parakeet (Psittacula krameri) -blue mutation on perch.jpg
A blue colour morph mutation parakeet kept as a pet
Mimicry (talking)

Rose-ringed parakeets are popular as pets and they have a long history in aviculture. The ancient Greeks kept the Indian subspecies P. krameri manillensis, and the ancient Romans kept the African subspecies P. krameri krameri. Colour mutations of the Indian rose-ringed parakeet subspecies have become widely available in recent years. [16] A blue colour morph mutation of the rose-ringed parakeet is also commonly kept in aviculture. Birds that display this mutation have solid light blue feathers instead of green.

Mimicry

Both males and females have the ability to mimic human speech. First, the bird listens to its surroundings, and then it copies the voice of the human speaker. Some people hand-raise rose-ringed parakeet chicks for this purpose. Such parakeets then become quite tame and receptive to learning. [17] They have extremely clear speech and are one of the best talking parrots.

Feral birds

Rose-ringed parakeets feeding on stored grain Rose-ringed parakeet.JPG
Rose-ringed parakeets feeding on stored grain
Rose-ringed parakeet feeding on sunflowers, Kolkata, India Rose-ringed parakeet feeding on Sunflower 1, Salt Lake, Kolkata.jpg
Rose-ringed parakeet feeding on sunflowers, Kolkata, India

A popular pet, the rose-ringed parakeet has been released in a wide range of cities around the world, giving it an environment with few predators where their preferred diet of seeds, nuts, fruits, and berries is available from suburban gardens and bird feeders. [3]

Feral populations are established in Europe, the U.S., South Africa, the Middle East, Australia and Japan. There are stable populations in Florida, California and Hawaii). Self-sustaining populations are also found in Turkey, [18] mostly in Ankara, İzmir, Istanbul (concentrated in parks), Tunis, Tripoli and Tehran (concentrated in the north side of the city). It is also found throughout Lebanon, Israel, Iran, Jordan, the United Arab Emirates, Bahrain, Qatar, and Oman. A small number of escaped birds are present in Australia. [1]

The specimens in these naturalised populations often represent intra-specific hybrids, originally between varying numbers (according to locality) of the subspecies manillensis, borealis[ verification needed ], and/or (to a lesser extent) krameri, along with some inter-specific hybrids with naturalised Psittacula eupatria (the Alexandrine parakeet). [19]

Where introduced, rose-ringed parakeets may affect native biodiversity and human economy and wellness. [20] [21]

Europe

Its adaptations to cold winters in the Himalayan foothills allow it to easily withstand European winter conditions. [3] The European populations became established during the mid-to-late 20th century.

A Europe-wide count was held in 2015 and found 85,220 rose-ringed parakeets in 10 European countries. [10]

CountryNumber
Belgium 10,800
France 7,250
Germany 10,960
Greece 1,000
Italy 9,170
Netherlands 20,000
Portugal 800
Spain 3,000
Turkey 5,000 [22]
UK 31,100
Total99,080

Rose-ringed parakeets are seen as a direct threat to populations of Europe's largest bat, the greater noctule, as parakeets compete with the bats for nesting sites, and will attack and kill adults before colonising their habitat. [23]

Great Britain

There is a burgeoning population of feral parakeets in Great Britain which is centred around suburban London and the Home Counties of South-East England. [24] Parakeet numbers have been highest in the south-west of London, although the population has since spread rapidly, and large flocks of birds can be observed in places such as Crystal Palace Park, Battersea Park, Buckhurst Hill, Richmond Park, Wimbledon Common, Greenwich Park, and Hampstead Heath, as well as Surrey and Berkshire. Feral parakeets have also been observed in Abbey Wood, Bostall Heath, Bostall Woods and Plumstead Common. The winter of 2006 had three separate roosts of about 6000 birds around London. [25] They have also established themselves in Kensington Gardens, Hyde Park, and Regent's Park. [26] A smaller population occurs around Margate, Broadstairs and Ramsgate, Kent. There is also an established population to the North East of London in Essex at Loughton and Theydon Bois by Epping Forest. Elsewhere in Britain, smaller feral populations have become established from time to time throughout the Midlands, Northern England and even as far north as Edinburgh. [27] It has been suggested that feral parrots could endanger populations of native British birds, and that the rose-ringed parakeet should be culled as a result, [28] although this is not currently recommended by conservation organisations. [4] A major agricultural pest in locations such as India, as of 2011 the rose-ringed parakeet population was growing rapidly, but is generally limited to urban areas in southern England. [29]

In the United Kingdom and especially within London, parakeets face predation by native birds of prey and owls, including the peregrine falcon (Falco peregrinus), Eurasian hobby (F. subbuteo) and tawny owl (Strix aluco). [30]

Benelux

In the Netherlands, the feral population in the four largest urban areas (Amsterdam, Rotterdam, Utrecht and The Hague) was estimated at 20,000 birds in 2021, [31] double the number of birds estimated in 2010. [32] There also exists a feral population in Belgium, with as many as 5,000 pairs estimated in Brussels. [33] These originate from an original population that was set free in 1974 by the owner of the Meli Zoo and Attraction Park near the Atomium who wanted to make Brussels more colourful. [34] [35]

Germany

In Germany, these birds are found along the Rhine in all major urban areas such as Bonn, Cologne, Düsseldorf (about 800 birds), [36] Frankenthal, Heidelberg, Ladenburg, Ludwigshafen, Mainz, Mannheim, Speyer, Wiesbaden, Worms and Zweibrücken.

France

Large populations in France are found especially in and around Paris but also in other places.

Italy

In Italy, Rome is notable for parakeet populations in the gardens of the Palatine Hill, the trees of Trastevere and Janiculum and at Villa Borghese. There are also colonies in Orto Botanico di Palermo in Palermo and in the city of Genoa. [37]

Spain

In Spain there are populations in Barcelona. [38]

Portugal

There are breeding population in Lisbon [38] and on Madeira Island. [39]

Turkey

In Turkey, there are populations in Istanbul over 1000 parakeets [40] and also in İzmir, Manisa, Muğla, Balıkesir, Ankara, Antalya totally over 5,000. [22]

Japan

There is a feral population of the birds in Japan. In the 1960s many Japanese people became pet owners for the first time and the parakeet was widely imported as a pet. Some escaped or were released and formed populations around the country. By the 1980s groups could be found in Tokyo, Osaka, Nagoya, Niigata and Kyushu. Some groups since died out, but as of 2009 there was a large population residing at the Tokyo Institute of Technology's main campus at Ookayama, along with small groups in Maebashi and Chiba city. [41]

New Zealand

Feral rose-ringed parakeets have sporadically been observed around New Zealand, and are treated as a major potential threat to the country's native bird populations due to their potential to outcompete native parakeet species, and introduce diseases. [42]

Rose-ringed parakeet (female) in New Delhi Rose-ringed parakeet (female) in New Delhi.jpg
Rose-ringed parakeet (female) in New Delhi

Aggression towards other animals

In the María Luisa Park in Seville, the population of the greater noctule declined sharply when the population of rose-ringed parakeets increased 20-fold. The rose-ringed parakeets attacked the greater noctule at tree cavities and occupied most of the cavities previously used by the bats. The attacks by the rose-ringed parakeets often led to the death of the greater noctule. In 14 years, the number of tree cavities occupied by the greater noctule decreased by 81%. A spatial analysis of tree cavity use showed that the greater noctule tried to avoid cavities near parakeets. [23] In the Rhineland, conspicuous bite wounds have been found in bats caught near rose-ringed parakeet nesting cavities. However, more detailed studies in the Rhineland on this are missing so far. [43] Several authors have reported negative behaviors of the ring-necked parakeet near their nest sites: lethal attacks on a Leisler's bat (Nyctalus leisleri) in Italy (Menchetti et al. 2014), on black rats (Rattus rattus) in Spain (Hernández-Brito et al. 2014b) and on several competitor and predator species in Spain (Hernández-Brito et al. 2014a). In the Paris area in France, an attack by a ring-necked parakeet on an adult red squirrel (Sciurus vulgaris) has been reported (Clergeau et al. 2009). [44]

Related Research Articles

<span class="mw-page-title-main">Parakeet</span> Small to medium-sized species of parrot

A parakeet is any one of many small to medium-sized species of parrot, in multiple genera, that generally has long tail feathers.

<span class="mw-page-title-main">Newton's parakeet</span> Extinct parrot from the island Rodrigues

Newton's parakeet, also known as the Rodrigues parakeet or Rodrigues ring-necked parakeet, is an extinct species of parrot that was endemic to the Mascarene island of Rodrigues in the western Indian Ocean. Several of its features diverged from related species, indicating long-term isolation on Rodrigues and subsequent adaptation. The rose-ringed parakeet of the same genus is a close relative and probable ancestor. Newton's parakeet may itself have been ancestral to the endemic parakeets of nearby Mauritius and Réunion.

<span class="mw-page-title-main">Plum-headed parakeet</span> Species of bird

The plum-headed parakeet is a species of parakeet in the family Psittacidae. It is endemic to the Indian Subcontinent and was once thought to be conspecific with the blossom-headed parakeet before being elevated to a full species. Plum-headed parakeets are found in flocks, the males having a pinkish purple head and the females, a grey head. They fly swiftly with twists and turns accompanied by their distinctive calls.

<span class="mw-page-title-main">Alexandrine parakeet</span> Species of bird

The Alexandrine parakeet, also known as the Alexandrine parrot, is a medium-sized parrot in the genus Psittacula of the family Psittaculidae, native to South Asia and Southeast Asia. It is named after Alexander the Great, who transported numerous birds from Punjab to various European and Mediterranean countries and regions, where they were prized by the royalty, nobility and warlords.

<i>Psittacula</i> Genus of birds

Psittacula, also known as Afro-Asian ring-necked parrots, is a genus of parrots from Africa and Southeast Asia. It is a widespread group with a clear concentration of species in south Asia, but also with representatives in Africa and the islands of the Indian Ocean. This is the only genus of parrot which has the majority of its species in continental Asia. Of all the extant species only Psittacula calthropae, Psittacula caniceps and Psittacula echo do not have a representative subspecies in any part of mainland continental Asia. The rose-ringed parakeet, Psittacula krameri, is one of the most widely distributed of all parrots.

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<span class="mw-page-title-main">Orange-winged amazon</span> Species of bird

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<span class="mw-page-title-main">Slaty-headed parakeet</span> Species of bird

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<span class="mw-page-title-main">Blue-winged parakeet</span> Species of bird

The blue-winged parakeet, also known as the Malabar parakeet is a species of parakeet endemic to the Western Ghats of southern India. Found in small flocks, they fly rapidly in forest clearings while making screeching calls that differ from those of other parakeet species within their distribution range. Their long blue tails tipped in yellow and the dark wings with blue contrast with the dull grey of their head and body. Adult males and females can be easily told apart from the colour of their beak.

<span class="mw-page-title-main">Echo parakeet</span> Species of parrot endemic to Mauritius

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<span class="mw-page-title-main">Red-breasted parakeet</span> Species of bird

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<span class="mw-page-title-main">Talking bird</span> Bird that can mimic human speech

Talking birds are birds that can mimic the speech of humans. There is debate within the scientific community over whether some talking parrots also have some cognitive understanding of the language. Birds have varying degrees of talking ability: some, like the corvids, are able to mimic only a few words and phrases, while some budgerigars have been observed to have a vocabulary of almost 2,000 words. The hill myna, a common pet, is well known for its talking ability and its relative, the European starling, is also adept at mimicry. Wild cockatoos in Australia have been reported to have learned human speech by cultural transmission from ex-captive birds that have integrated into the flock.

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<span class="mw-page-title-main">Feral parrot</span> Parrot in non-native environment

A feral parrot is a parrot that has adapted to life in an ecosystem to which it is not native. The birds are often descended from pets that have escaped or been deliberately released. Many species of parrots are highly social, and like to gather in large flocks. Not all feral parrots are able to adapt to life outside of captivity, although having a pre-existing nearby parrot colony can assist with the adaptation process.

<span class="mw-page-title-main">Feral parakeets in Great Britain</span> Population of feral parakeets in Great Britain

Feral parakeets in Great Britain are wild-living, non-native parakeets that are an introduced species into Great Britain. The population mainly consists of rose-ringed parakeets, a non-migratory species of bird native to Africa and the Indian Subcontinent, with a few, small breeding populations of monk parakeets, and other occasional escaped cage birds. The origins of these birds are subject of speculation, but they are generally thought to have bred from birds that escaped from captivity or were released.

<span class="mw-page-title-main">Parrot</span> Order of birds

Parrots (Psittaciformes), also known as psittacines, are birds with a strong curved beak, upright stance, and clawed feet. They are conformed by four families that contain roughly 410 species in 101 genera, found mostly in tropical and subtropical regions. The four families are the Psittaculidae, Psittacidae, Cacatuoidea (cockatoos), and Strigopidae. One-third of all parrot species are threatened by extinction, with a higher aggregate extinction risk than any other comparable bird group. Parrots have a generally pantropical distribution with several species inhabiting temperate regions as well. The greatest diversity of parrots is in South America and Australasia.

<span class="mw-page-title-main">Mascarene grey parakeet</span> Extinct parrot from Mauritius and Réunion

The Mascarene grey parakeet, Mauritius grey parrot, or Thirioux's grey parrot, is an extinct species of parrot which was endemic to the Mascarene Islands of Mauritius and Réunion in the western Indian Ocean. It has been classified as a member of the tribe Psittaculini, along with other parrots from the Islands.

<span class="mw-page-title-main">Monk parakeet</span> South American true parrot

The monk parakeet, also known as the monk parrot or Quaker parrot, is a species of true parrot in the family Psittacidae. It is a small, bright-green parrot with a greyish breast and greenish-yellow abdomen. Its average lifespan is 20–30 years. It originates from the temperate to subtropical areas of South America. Self-sustaining feral populations occur in many places, mainly in areas of similar climate in North America and Europe.

References

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  2. ↑ Stephen Pruett-Jones (2021). Naturalized Parrots of the World: Distribution, Ecology, and Impacts of the World's Most Colorful Colonizers. Princeton University Press. p. 55. ISBN   978-0-691-20441-3.
  3. 1 2 3 4 "How do parakeets survive in the UK?". BBC News Online. 22 March 2007. Retrieved 14 October 2017.
  4. 1 2 "Ring-Necked Parakeets in the UK". Royal Society for the Protection of Birds (RSPB). Archived from the original on 24 March 2019. Retrieved 20 August 2019.
  5. 1 2 Pithon, Josephine; Dytham, Calvin (2001). "Determination of the origin of British feral Rose-ringed Parakeets". British Birds: 74–79. Retrieved 14 October 2017.
  6. ↑ Morgan, David (1993). "Feral Rose-ringed Parakeets in Britain". British Birds: 561–4. Retrieved 14 October 2017.
  7. ↑ Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp.  215, 321. ISBN   978-1-4081-2501-4.
  8. ↑ Braun, Michael P.; Datzmann, Thomas; Arndt, Thomas; Reinschmidt, Matthias; Schnitker, Heinz; Bahr, Norbert; Sauer-GüRth, Hedwig; Wink, Michael (2019). "A molecular phylogeny of the genus Psittacula sensu lato (Aves: Psittaciformes: Psittacidae: Psittacula, Psittinus, Tanygnathus, †Mascarinus) with taxonomic implications". Zootaxa. 4563 (3): 547–562. doi:10.11646/zootaxa.4563.3.8. PMID   31716534. S2CID   91969786.
  9. 1 2 Bikram Grewal; Bill Harvey; Otto Pfister (2014). Photographic Guide to the Birds of India: And the Indian Subcontinent, Including Pakistan, Nepal, Bhutanh, Bangladesh, Sri Lanka & the Maldives. Tuttle Publishing. p. 109. ISBN   978-1-4629-1485-2.
  10. 1 2 Pârâu, Liviu; Strubbe, Diederik; Mori, Emiliano; Menchetti, Mattia; Ancillotto, Leonardo; van Kleunen, André; White, Rachel; Luna, Álvaro; Hernández-Brito, Dailos; Le Louarn, Marine; Clergeau, Philippe; Albayrak, Tamer; Franz, Detlev; Braun, Michael; Schroeder, Julia; Wink, Michael (2016). "Rose-ringed parakeet populations and numbers in Europe: A complete overview". The Open Ornithology Journal. 9: 1–13. doi: 10.2174/1874453201609010001 . hdl: 10261/133407 .
  11. ↑ Strubbe, Diederik; Matthysen, Erik (2009). "Establishment success of invasive ring-necked and monk parakeets in Europe". Journal of Biogeography. 36 (12): 2264–2278. Bibcode:2009JBiog..36.2264S. doi:10.1111/j.1365-2699.2009.02177.x. S2CID   85979303.
  12. ↑ Sailaja, R., Kotak, V. C., Sharp, P. J., Schmedemann, R., Haase, E. (1988). Environmental, dietary, and hormonal factors in the regulation of seasonal breeding in free-living female Indian rose-ringed parakeets (Psittacula krameri). Hormones and Behavior 22 (4): 518–527.
  13. ↑ "Feeding Your Indian Ringneck or Asiatic Parrot". Indianringneck.com.
  14. ↑ "Indian Ring-Necked Parakeet". Lafeber.com.
  15. 1 2 3 Krishnaprasadan, T. N.; Kotak, Vibhakar C.; Sharp, Peter J.; Schmedemann, Reiner; Haase, Eberhard (1988). "Environmental and hormonal factors in seasonal breeding in free-living male Indian rose-ringed parakeets (Psittacula krameri)". Hormones and Behavior. 22 (4): 488–496. doi:10.1016/0018-506X(88)90053-0. PMID   3235065. S2CID   51919.
  16. ↑ Alderton, David (2003). The Ultimate Encyclopedia of Caged and Aviary Birds. London, England: Hermes House. pp. 189–190. ISBN   1-84309-164-X.
  17. ↑ "Rose-Ringed Parakeet". Calm Zoo. 20 December 1977. Retrieved 27 May 2020.
  18. ↑ "İstanbul'un sevimli misafirleri: Papağanlar". Anadolu Ajansı . 24 November 2018. Retrieved 2 February 2024.
  19. ↑ Ancillotto, Leonardo; Strubbe, Diederik; Menchetti, Mattia; Mori, Emiliano (2015). "An overlooked invader? Ecological niche, invasion success and range dynamics of the Alexandrine parakeet in the invaded range" (PDF). Biological Invasions. 18 (2): 1–13. doi:10.1007/s10530-015-1032-y. hdl: 10067/1292210151162165141 . S2CID   17533751.
  20. ↑ Menchetti, Mattia; Mori, Emiliano (2014). "Worldwide impact of alien parrots (Aves Psittaciformes) on native biodiversity and environment: a review". Ethology Ecology & Evolution. 26 (2–3): 172–194. Bibcode:2014EtEcE..26..172M. doi:10.1080/03949370.2014.905981. S2CID   84294580.
  21. ↑ Menchetti, Mattia; Mori, Emiliano; Angelici, Francesco Maria (2016). Effects of the recent world invasion by ring-necked parakeets Psittacula krameri. Springer International Publishing. pp. 253–266. doi:10.1007/978-3-319-22246-2_12. ISBN   978-3-319-22246-2.
  22. 1 2 "5000 PAPAĞAN ARAMIZDA". Atlas Dergisi. 15 February 2022. Retrieved 2 February 2024.
  23. 1 2 Hernández-Brito, Dailos; Carrete, Martina; Ibáñez, Carlos; Juste, Javier; Tella, José L. (2018). "Nest-site competition and killing by invasive parakeets cause the decline of a threatened bat population". Royal Society Open Science. 5 (5): 172477. Bibcode:2018RSOS....572477H. doi:10.1098/rsos.172477. PMC   5990744 . PMID   29892437.
  24. ↑ Attenborough, D. 1998. The Life of Birds. p.298. BBC ISBN   0563-38792-0
  25. ↑ London Bird Report 2006. London Natural History Society. 2006. p. 93. ISBN   978-0-901009-22-7.
  26. ↑ Hunt, Nick (6 June 2019). "The great green expansion: how ring-necked parakeets took over London". The Guardian. Retrieved 8 June 2023.
  27. ↑ "Scotsman – Parakeets Mystery is Causing a Bit of a Flap".
  28. ↑ "Parakeet 'threat' to native birds". BBC. 22 March 2007.
  29. ↑ Rosenthal, Elisabeth (13 May 2011). "British Parakeet Boom Is a Mystery, and a Mess". The New York Times. Retrieved 14 May 2011.
  30. ↑ McCarthy, Michael (8 June 2015). "Nature Studies: London's beautiful parakeets have a new enemy to deal with". The Independent . Retrieved 20 December 2017.
  31. ↑ Meershoek, Patrick (22 December 2021). "Halsbandparkiet verspreidt zich over Amsterdam, met kolonies overal". Het Parool (in Dutch). Retrieved 22 December 2021.
  32. ↑ "Duizenden halsbandparkieten in grote steden". nos.nl (in Dutch). 14 January 2010. Retrieved 22 December 2021.
  33. ↑ "Halsbandparkieten". De Standaard. 19 June 2009.
  34. ↑ Joyce Mesdag (8 February 2019). "Exotische parkieten veroveren Park van Heule, en dat is géén goeie zaak voor onze inheemse vogels". Het Laatste Nieuws.
  35. ↑ "Invasive Alien Species in Belgium: Psittacula krameri". ias.biodiversity.be.
  36. ↑ "Information page". Umweltamt Düsseldorf.
  37. ↑ Verner, Aldo (17 May 2012). "Pappagalli verdi, ex "prigionieri" che a Genova volano in libertà". Genova.it (in Italian). Archived from the original on 14 July 2014. Retrieved 6 July 2015.
  38. 1 2 Geraldes, Helena (25 December 2011). "Que misteriosas aves verdes e estridentes são estas que invadiram Lisboa". Público. Retrieved 25 December 2011.
  39. ↑ Rocha, Ricardo; et al. (2020). "Introduced population of ring-necked parakeets Psittacula krameri in Madeira Island, Portugal – Call for early action" (PDF). Management of Biological Invasions. 11 (3): 576–587. doi: 10.3391/mbi.2020.11.3.15 .
  40. ↑ "İstanbul'un sevimli misafirleri: Papağanlar". Anadolu Ajansı . 24 November 2018. Retrieved 2 February 2024.
  41. ↑ Gordenker, Alice (19 March 2009). "Feral parakeets" . The Japan Times. Retrieved 8 February 2023.
    Raillan Brooks (8 December 2014). "Tokyo's Got a Parrot Problem". Audubon Magazine. Retrieved 8 February 2023.
    Kail, Ellyn (22 August 2014). "Eerie Photos of Feral Parrots in Tokyo". Feature Shoot. Retrieved 8 February 2023.
  42. ↑ Ministry for Primary Industries. "Indian ring-necked parakeets in NZ: Reporting and identification". mpi.govt.nz. New Zealand Government. Retrieved 8 February 2023.
  43. ↑ "Esther Koch: Die Halsbandsittich-Population im Rheinland/NRW. NWO-Mitteilungen 55, 2022, pp. 57-58" (PDF).
  44. ↑ Le Louarn, Marine; Couillens, Bertrand; Deschamps-Cottin, Magali; Clergeau, Philippe (2016). "Interference competition between an invasive parakeet and native bird species at feeding sites". Journal of Ethology. 34 (3): 291–298. doi:10.1007/s10164-016-0474-8. PMC   5080312 . PMID   27829702.
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