Palaeoscinis Temporal range: Middle Miocene | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Suborder: | Passeri |
Family: | † Palaeoscinidae Howard, 1957 |
Genus: | † Palaeoscinis Howard, 1957 |
Species: | †P. turdirostris |
Binomial name | |
†Palaeoscinis turdirostris Howard, 1957 | |
Palaeoscinis (meaning "ancient oscine") is an extinct genus of songbird described in 1957 from the middle Miocene of the Monterey Formation in Santa Barbara, California. It is assigned to the extinct monotypic family Palaeoscinidae, and contains the type and only species P. turdirostris.
The fossil was first discovered in 1955 on two slabs of limestone intended for use as flagstones before being recognised for their significance. It was preserved partly as an imprint of the skeleton with some of bones still intact and in articulation with only slight separation of individual bones, including scattered tracheal rings. The fossil was found in marine sediments, but the anatomy of Palaeoscinis indicates that it was a songbird rather than a seabird. The specific name was chosen for the thrush-like shape of its beak.
The beak is long and slender, similar to that of the modern varied thrush, and its body proportions are also generally similar to thrushes. The legs are relatively short, though, unlike thrushes, and are shorter than the bones of the wing. Feather imprints on the slab suggest the wings may have been relatively short, although it is possible that they were not fully preserved. The foot is typical of songbirds, with a reversed 1st toe and three forward facing ones, of which the middle is the longest and the 2nd is the shortest, with the 1st and 4th toes roughly equal in length. The three forward facing claws are short, but the claw of the reversed 1st toe is long, approximately 70% the length of the preceding toe bone.
Palaeoscinus was assigned to its own family due to the combination of skeletal characteristics (mostly of the sternum, shoulder, and humerus) and its proportions being unlike any modern or fossil family of songbirds. However, it was regarded as similar and possibly related to the Pycnonotidae (bulbuls), Bombycillidae (waxwings), Corvidae, or Cinclidae (dippers) by Howard (1957). It was provisionally allocated as closest to the Pycnonotidae over the other families by Wetmore (1960) based on its proportions, although this relationship is not certain.
The area has produced fossils of other marine animals, including fossils of a shearwater and the "pseudo-toothed" pelagornithid Osteodontornis , as well as a porpoise and fish, alongside fossils of palms.
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Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous flightless birds that were among the largest apex predators in South America during the Cenozoic era; their conventionally accepted temporal range covers from 53 to 0.1 million years (Ma) ago.
Dromornis is a genus of large to enormous prehistoric birds native to Australia during the Oligocene to Pliocene epochs. The species were flightless, possessing greatly reduced wing structures but with large legs, similar to the modern ostrich or emu. They were likely to have been predominantly, if not exclusively, herbivorous browsers. The male of the largest species, Dromornis stirtoni, is a contender for the tallest and heaviest bird, and possibly exhibited aggressive territorial behaviour. They belong to the family Dromornithidae, extinct flightless birds known as mihirungs.
Teratornithidae is an extinct family of very large birds of prey that lived in North and South America from the Late Oligocene to Late Pleistocene. They include some of the largest known flying birds.
Titanis is a genus of phorusrhacid, an extinct family of large, predatory birds, in the order Cariamiformes that inhabited the United States during the Pliocene and earliest Pleistocene. The first fossils were unearthed by amateur archaeologists Benjamin Waller and Robert Allen from the Santa Fe River in Florida and were named Titanis walleri by ornithologist Pierce Brodkorb in 1963, the species name honoring Waller. The holotype material is fragmentary, consisting of only an incomplete right tarsometatarsus and phalanx, but comes from one of the largest phorusrhacid individuals known. In the years following the description, many more isolated elements have been unearthed from sites from other areas of Florida, Texas, and California. It was classified in the subfamily Phorusrhacinae, which includes some of the last and largest phorusrhacids like Devincenzia and Kelenken.
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The Pelagornithidae, commonly called pelagornithids, pseudodontorns, bony-toothed birds, false-toothed birds or pseudotooth birds, are a prehistoric family of large seabirds. Their fossil remains have been found all over the world in rocks dating between the Early Paleocene and the Pliocene-Pleistocene boundary.
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Devincenzia is an extinct genus of giant flightless predatory birds in the family Phorusrhacidae or "terror birds" that lived during the Early Miocene (Deseadan) Fray Bentos Formation of Uruguay and Late Miocene (Huayquerian) Ituzaingó Formation to Early Pliocene (Montehermosan) of Argentina. The type species D. pozzi was formerly known as Onactornis pozzi. It stood about 2.5 metres (8.2 ft) tall, making it one of the largest Phorusrhacids and carnivorous birds known.
Palaeochenoides is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Eurofluvioviridavis is a genus of extinct primitive birds from the Middle Eocene Messel Pit, Germany. It contains a single species, Eurofluvioviridavis robustipes. It is related to Avolatavis and Vastanavis, other members of the family Vastanavidae.
Bohaiornithidae is a group of early predatory enantiornitheans from the early Cretaceous Period of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125–120 million years ago. Bohaiornithidae was first coined as a family of enantiornithean birds by Wang and colleagues in 2014. They defined it as the natural group formed by all descendants of the common ancestor of the type species, Bohaiornis guoi, and Shenqiornis mengi.
Bathornis is an extinct lineage of birds related to modern day seriemas, that lived in North America about 37–20 million years ago. Like the closely related and also extinct phorusrhacids, it was a flightless predator, occupying predatory niches in environments classically considered to be dominated by mammals. It was a highly diverse and successful genus, spanning a large number of species that occurred from the Priabonian Eocene to the Burdigalian Miocene epochs.
Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.
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