Catnip | |
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Catnip flowers | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Asterids |
Order: | Lamiales |
Family: | Lamiaceae |
Genus: | Nepeta |
Species: | N. cataria |
Binomial name | |
Nepeta cataria | |
Synonyms [1] | |
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Nepeta cataria, commonly known as catnip and catmint, is a species of the genus Nepeta in the mint family, native to southern and eastern Europe, the Middle East, and Central Asia. It is widely naturalized in northern Europe, New Zealand, and North America. The common name catmint can also refer to the genus as a whole.
The names catnip and catmint are derived from the intense attraction about two-thirds of cats have toward the plant. Catnip is also an ingredient in some herbal teas, and is valued for its sedative and relaxant properties.
Nepeta cataria is a short-lived perennial that grows 30 to 100 cm (12 to 39 in) tall, usually with several stems. [2] Each of its stems is square in cross section, as typical of the mint family, and somewhat gray in color. [3] It is a herbaceous plant that regrows from a taproot. [2] However, it does not deeply root. [4] Older plants tend to have more branches with particularly healthy plants becoming mound shaped. [5]
The leaves are canescent in appearance, white in color due to being covered in fine hairs, especially so on the lower side of the leaves. [2] They are attached in pairs to opposite sides of the stems. [6] Leaf shapes vary from cordate, deltoid, to ovate; shaped like a heart, triangle, or like an egg. [5] They are attached by leaf stems and have a length of 2 to 9 cm (0.8 to 3.5 in) and 0.6 to 6 cm (0.2 to 2.4 in) wide. [7] The edges of the leaves are coarsely crenate to serrate, having a wavy, rounded edge to have asymmetrical teeth like those of a saw that point forward. [2]
The flowers are in loose groups in an inflorescence. The lowest flowers more widely spaced and the end more tightly packed into a spike. [5] The inflorescences are at the end of branches and may be 2 to 8 cm (0.8 to 3.1 in) long and have inconspicuous bracts. [2] A single plant may produce several thousand flowers, but at any time less than 10% of them will be in full bloom. [8] The flowers themselves are somewhat small and inconspicuous, [9] but quite fragrant. [3] They are bilaterally symetrical and measure 10–12 mm long. [8] The petals are off white to pink and usually dotted with purple-pink spots. [10] They are bilabiate with the upper lip having two lobes and the lower one much wider with a scalloped edge. [2]
The fruit is a nutlet that is nearly triquetrous, three sided with sharp edges and concave sides, and over all shaped like an egg. They are approximately 1.7 mm by 1 mm. [11] Each nutlet may contain between one and four seeds. [8] They are dark reddish-brown in color with two white spots near the base. [12]
Nepeta cataria was one of the many species described by Linnaeus in 1753 in his landmark work Species Plantarum . [13] He had previously described it in 1738 as Nepeta floribus interrupte spicatis pedunculatis (meaning "Nepeta with flowers in a stalked, interrupted spike"), before the commencement of Linnaean taxonomy. [14] Catnip is classified in part of Nepeta in the Lamiaceae, commonly known as the mint family. [15] It has no subspecies or varieties. [1]
Nepeta cataria has 19 botanical synonyms, 16 of which are species. Only three are exactly equivelent to the current description of the species. [1]
Name | Year | Rank | Notes |
---|---|---|---|
Calamintha albifloraVaniot | 1904 | species | = het. |
Cataria tomentosaGilib. | 1782 | species | = het. opus utique oppr. |
Cataria vulgarisGaterau | 1789 | species | ≡ hom. |
Glechoma cataria(L.) Kuntze | 1891 | species | ≡ hom. |
Glechoma macrura(Ledeb. ex Spreng.) Kuntze | 1891 | species | = het. |
Nepeta americanaVitman | 1789 | species | = het. nom. illeg. |
Nepeta bodinieriVaniot | 1904 | species | = het. |
Nepeta cataria var. canescensSennen | 1903 | variety | = het. nom. nud. |
Nepeta cataria var. citriodoraDumoulin ex Lej. | 1825 | variety | = het. |
Nepeta cataria f. laurentii(Sennen) Font Quer | 1951 | form | = het. |
Nepeta ceretanaSennen | 1931 | species | = het. |
Nepeta citriodora(Dumoulin ex Lej.) Dumort. | 1827 | species | = het. |
Nepeta laurentiiSennen | 1934 | species | = het. |
Nepeta macruraLedeb. ex Spreng. | 1825 | species | = het. |
Nepeta minorMill. | 1768 | species | = het. |
Nepeta mollisSalisb. | 1796 | species | = het. nom. illeg. |
Nepeta ruderalisBoiss. | 1879 | species | = het. nom. illeg. |
Nepeta tomentosaVitman | 1789 | species | = het. |
Nepeta vulgarisLam. | 1779 | species | ≡ hom. nom. superfl. |
Notes: ≡ homotypic synonym ; = heterotypic synonym |
The species name cataria means "of cats". It derives from the medival Latin herba catti or herba cattaria used by medieval herbalists. [16] The English common name catnip is first recorded in 1775 in the colony of Pennsylvania, but now has worldwide usage. [17] The variant catnep was also coined in the United States around 1806, but never became common elsewhere and is now very rarely used. [18]
The first usage of catmint was in about 1300 in the form kattesminte. It continues to be used for Nepeta cataria, though it is also used for other species in the genus and the Nepeta as a genus. [19] In medieval English it was also called cat-wort, but it is no longer used in English dying out by about 1500. [20]
Another name with a medieval origin was nep, neps, or nepe. Originating about 1475, it was more common but has become a regional name for catnip used in East Anglia. [21] [22]
In medieval England it was known by various names in botanical manuscripts. It was called calamentum minus and nasturcium mureligi. [23] It was also called nepeta or variants, but other species or genuses like the dead-nettles ( Lamium ) were also sometimes called this. [24] It was also sometimes called collocasia, but this was more often applied to horse-mints especially Mentha longifolia . [25]
According to Plants of the World Online, the native range of catnip includes a large part of Eurasia. [1] In Europe it is certainly native to the south around the Mediterranean and in the east, but sources disagree on its native status in the north in countries like the Baltic Countries, Germany, the Netherlands, and England. Around the Mediterranean it is idetified as native in Portugal, Spain, France, Corsica, Italy, Switzerland, the former Yugoslavia, Albania, and Greece. In the East it is native to Bulgaria, Romania, Ukraine, Belarus, European Russia, and the Caucasus. It is generally agreed to be an introduced species in Scandinavia, Poland, and may also grow in Ireland. [1] [26]
In Asia its range extends from Turkey into Syria, Lebanon, and Iraq. Eastward it continues to Iran and Pakistan and the western Himalayas, but no further into India. It is native to all of Central Asia including Afghanistan, Kazakhstan, Kirghistan, Tajikistan, Turkmenistan, and Uzbekistan and also extends to western Siberia. Its native status in China is disputed as it also is in the Russian Far East, Nepal, Korea, and Japan. [1] [26]
In Africa it may grow in Morrocco, but this report is doubtful. [1] It also grows as introduced species on the island of Java. [1] In Australia it has been reported in the states of South Australia, New South Wales, Victoria, Queensland, and Tasmania. [27] It grows on both the north and south islands of New Zealand and was introduced there in 1870. [28]
In North America it grows in Canada from the island of Newfoundland to British Columbia, but not in Labrador or the three northern Canadian territories. [29] In the United States it is present in 48 states, only absent from Florida and Hawaii. [30]
In South America it grows in many parts of Argentina as well as in Colombia. [1]
It grows in a variety of soils from clay to sandy or even shallow and rocky. It requires good drainage and not to be water logged. [31]
The plant terpenoid nepetalactone is the main chemical constituent of the essential oil of Nepeta cataria. Nepetalactone can be extracted from catnip by steam distillation. [32]
Nepeta cataria is cultivated as an ornamental plant for use in gardens. It is also grown for its attractant qualities to house cats and butterflies. [3]
The plant is drought-tolerant and deer-resistant. It can be a repellent for certain insects, including aphids and squash bugs. [3] Catnip is best grown in full sunlight and grows as a loosely branching, low perennial. [33]
The cultivar Nepeta cataria 'Citriodora', also known as lemon catmint, is known for the strong lemon-scent of its leaves. [34]
The iridoid that is deposited on cats who have rubbed themselves against the plants and scratched the surfaces of catnip and silver vine (Actinidia polygama) leaves repels mosquitoes. [35] The compound iridodial, an iridoid extracted from catnip oil, has been found to attract lacewings that eat aphids and mites. [36]
Nepetalactone is a mosquito and fly repellent. [37] [38] Oil isolated from catnip by steam distillation is a repellent against insects, in particular mosquitoes, cockroaches, and termites. [39] [40] Research suggests that, while a more effective spatial repellant than DEET, [41] it is not as effective of a repellent when used on the skin of humans as SS220 or DEET. [42]
Catnip has a history of use in traditional medicine for a variety of ailments such as stomach cramps, indigestion, fevers, hives, and nervous conditions. The plant has been consumed as a tisane, juice, tincture, infusion, or poultice, and has also been smoked. [43] However, its medicinal use has fallen out of favor with the development of modern medicine. [44]
Catnip contains the feline attractant nepetalactone. N. cataria (and some other species within the genus Nepeta) are known for their behavioral effects on the cat family, not only on domestic cats, but also other species. [44] Several tests showed that leopards, cougars, servals, and lynxes often reacted strongly to catnip in a manner similar to domestic cats. Lions and tigers may react strongly as well, but they do not react consistently in the same fashion. [45] [46] [47] [48]
With domestic cats, N. cataria is used as a recreational substance for the enjoyment of pet cats, and catnip and catnip-laced products designed for use with domesticated cats are available to consumers. Common behaviors cats display when they sense the bruised leaves or stems of catnip are rubbing on the plant, rolling on the ground, pawing at it, licking it, and chewing it. Consuming much of the plant is followed by drooling, sleepiness, anxiety, leaping about, and purring. [49] Some growl, meow, scratch, or bite at the hand holding it. [50] [51] The main response period after exposure is generally between 5 and 15 minutes, after which olfactory fatigue usually sets in. [52] However, about one-third of cats are not affected by catnip. [53] [44] The behavior is hereditary. [54]
Cats detect nepetalactone through their olfactory epithelium, not through their vomeronasal organ. [55] At the olfactory epithelium, the nepetalactone binds to one or more olfactory receptors.
A 1962 pedigree analysis of 26 cats in a Siamese breeding colony suggested that the catnip response was caused by a Mendelian-dominant gene. A 2011 pedigree analysis of 210 cats in two breeding colonies (taking into account measurement error by repeated testing) showed no evidence for Mendelian patterns of inheritance but demonstrated heritabilities of h2 = 0.51–0.89 for catnip response behavior, indicating a polygenic liability threshold model. [56] [57]
A study published in January 2021 suggests that felines are specifically attracted to the iridoids nepetalactone and nepetalactol, present in catnip and silver vine, respectively. [58]
Cats younger than six months might not exhibit behavioral change to catnip. [59] Up to a third of cats are genetically immune to catnip effects but may respond in a similar way to other plants such as valerian (Valeriana officinalis) root and leaves, silver vine or matatabi ( Actinidia polygama ), and Tatarian honeysuckle ( Lonicera tatarica ) wood. [60]
Nepeta is a genus of flowering plants in the family Lamiaceae. The genus name, from Latin nepeta (“catnip”), is reportedly in reference to Nepete, an ancient Etruscan city. There are 295 accepted species.
Valerian is a perennial flowering plant native to Europe and Asia. In the summer when the mature plant may have a height of 1.5 metres, it bears sweetly scented pink or white flowers that attract many fly species, especially hoverflies of the genus Eristalis. It is consumed as food by the larvae of some Lepidoptera species, including the grey pug.
Nepetalactone is a name for multiple iridoid analog stereoisomers. Nepetalactones are produced by Nepeta cataria (catnip) and many other plants belonging to the genus Nepeta, in which they protect these plants from herbivorous insects by functioning as insect repellents. They are also produced by many aphids, in which they are sex pheromones. Nepetalactones are cat attractants, and cause the behavioral effects that catnip induces in domestic cats. However, they affect visibly only about two thirds of adult cats. They produce similar behavioral effects in many other felids, especially in lions and jaguars. In 1941, the research group of Samuel M. McElvain was the first to determine the structures of nepetalactones and several related compounds.
Primula sect. Dodecatheon is a section of herbaceous flowering plants in the family Primulaceae. Primula species in this section were formerly placed in a separate genus, Dodecatheon. The species have basal clumps of leaves and nodding flowers that are produced at the top of tall stems rising from where the leaves join the crown. The genus is largely confined to North America and part of northeastern Siberia. Common names include shooting star, American cowslip, mosquito bills, mad violets, and sailor caps. A few species are grown in gardens for their showy and unique flower display.
Species Plantarum is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the starting point for the naming of plants.
Cornus canadensis is a species of flowering plant in the dogwood family Cornaceae, native to eastern Asia and North America. Common names include Canadian dwarf cornel, Canadian bunchberry, quatre-temps, crackerberry, and creeping dogwood. Unlike its relatives, which are for the most part substantial trees and shrubs, C. canadensis is a creeping, rhizomatous perennial growing to about 20 centimetres tall.
Catnip, Nepeta cataria, is a species in the family Lamiaceae (mint).
Agastache is a genus of aromatic flowering herbaceous perennial plants in the family Lamiaceae. It contains 22 species, mainly native to North America, one species native to eastern Asia. The common names of the species are a variety of fairly ambiguous and confusing "hyssops" and "mints"; as a whole the genus is known as giant hyssops or hummingbird mints.
Nepeta × faassenii, a flowering plant also known as catmint and Faassen's catnip, is a primary hybrid of garden origin. The parent species are Nepeta racemosa and Nepeta nepetella.
Chromolaena odorata is a tropical and subtropical species of flowering shrub in the family Asteraceae. It is native to the Americas, from Florida and Texas in the United States south through Mexico and the Caribbean to South America. It has been introduced to tropical Asia, West Africa, and parts of Australia.
A cat pheromone is a chemical molecule, or compound, that is used by cats and other felids for communication. These pheromones are produced and detected specifically by the body systems of cats and evoke certain behavioural responses.
Actinidia polygama is a species of kiwifruit in the family Actinidiaceae. It grows in the mountainous areas of Korea, Japan and China at elevations between 500 and 1,900 metres.
Lonicera tatarica is a species of honeysuckle known by the common name Tatarian honeysuckle. Native to Eurasia, the plant is one of several exotic bush honeysuckles present in North America, being considered an invasive species there.
Iridomyrmecin is a defensive chemical, classified as an iridoid, isolated from ants of the genus Iridomyrmex. It has also evolved into a sex pheromone in wasps such as Leptopilina, with host species using the smell of iridomyrmecin as a way of detecting the presence of the parasitoid wasps. Iridomyrmecin is also found in a variety of plants including Actinidia polygama.
Nepeta racemosa, the dwarf catnip or raceme catnip, syn. N. mussiniii, is a species of flowering plant in the mint family Lamiaceae, native to the Caucasus, Turkey and northern Iran. Growing to 30 cm (12 in) tall by 45 cm (18 in) wide, it is a herbaceous perennial with aromatic leaves and violet or lilac-blue flowers in summer.
Nepeta nepetella, common name lesser cat-mint, is a low-growing species of catnip belonging to the family Lamiaceae. It is native to France, Spain, Italy, Algeria, and Morocco.
Nepeta grandiflora is a species of flowering plant in the mint family Lamiaceae, native to the Caucasus. Growing to 75 cm (30 in) tall by 30 cm (12 in), it is a clump-forming, erect deciduous herbaceous perennial with aromatic, slightly hairy, grey-green leaves, and spikes of purple/blue flowers in early summer. Species of Nepeta are called catnip or catmint, with reference to their reported effect on some domestic cats. The plants seem to induce a euphoria in the animals, causing them to roll in the foliage and exhibit signs of intoxication.
Nepeta sibirica, the Siberian catmint, is a species of flowering plant in the mint family Lamiaceae, native to Siberia.
Nepeta curviflora, commonly known as Syrian catnip, is a herbaceous flowering plant native to the Middle East. It was first described in 1844.
Several non-human animal species are said to engage in apparent recreational drug use, that is, the intentional ingestion of psychoactive substances in their environment for pleasure, though claims of such behavior in the wild are often controversial. This is distinct from zoopharmacognosy, in which animals ingest or topically apply non-food substances for their health benefits, as a form of self-medication.
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