Neotrirachodon Temporal range: Anisian ~ | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Suborder: | † Therocephalia |
Superfamily: | † Baurioidea |
Family: | † Bauriidae (?) |
Genus: | † Neotrirachodon Tatarinov, 2002 |
Species: | †N. expectatus |
Binomial name | |
†Neotrirachodon expectatus Tatarinov, 2002 [1] | |
Synonyms [2] | |
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Neotrirachodon is an extinct genus of therapsids which existed in Russia during the Middle Triassic period. Its type and only species is Neotrirachodon expectatus. [3]
It is known from a jaw fossil consisting of the left dentary with postcanines. It was recovered from the Donguz Formation, Orenburg Oblast of Russia. [4] [5] The fossil was first described as belonging to the genus Antecosuchus (as Antecosuchus ochevi), [4] but in 2002 it was placed in its own monotypic genus. [3]
It has tentatively been classified as a trirachodontid gomphodont pending further fossil discoveries. Gomphodonts were a group of herbivorous cynodonts which lived during the Triassic. [6] If correct, it would be the first trirachodontid described from Russia. [4]
Its classification remains uncertain, however. Other studies treat it as an incertae sedis gomphodont, while others propose that it is more likely to be a bauriid therocephalian rather than a trirachodontid. [7] [8]
The cynodonts are a clade of eutheriodont therapsids that first appeared in the Late Permian, and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety of lifestyles, including carnivory and herbivory. Mammals are cynodonts, as are their extinct ancestors and close relatives, having evolved from advanced probainognathian cynodonts during the Late Triassic. All other cynodont lines went extinct, with the last known non-mammalian cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous.
Cynognathia is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus Cynognathus and the herbivorous traversodontids. Cynognathians can be identified by several synapomorphies including a very deep zygomatic arch that extends above the middle of the orbit.
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil.
Trirachodon is an extinct genus of cynodonts. Fossils have been found in the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Omingonde Formation of Namibia, dating back to the Early and Middle Triassic.
Lumkuia is an extinct genus of cynodonts, fossils of which have been found in the Cynognathus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic. It contains a single species, Lumkuia fuzzi, which was named in 2001 on the basis of the holotype specimen BP/1/2669, which can now be found at the Bernard Price Institute in Johannesburg, South Africa. The genus has been placed in its own family, Lumkuiidae. Lumkuia is not as common as other cynodonts from the same locality such as Diademodon and Trirachodon.
Langbergia is an extinct genus of trirachodontid cynodont from the Early Triassic of South Africa. The type and only species L. modisei was named in 2006. Langbergia was found in the Burgersdorp Formation in the Beaufort Group, a part of the Cynognathus Assemblage Zone. The closely related trirachodontids Trirachodon and Cricodon were found in the same area.
Menadon is an extinct genus of traversodontid cynodonts. The type and only species is Menadon besairiei.
Malasaurus is an extinct genus of therocephalian therapsids which existed in Russia. The type species is Malasaurus germanus.
Trirachodontidae is an extinct, possibly paraphyletic family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quickly spread over a wide geographic area in a comparatively brief amount of time from 250 to 237 million years ago.
Gornogomphodon is an extinct genus of cynodonts from the Gorno Formation of Bergamo, Italy. It existed during the Middle Carnian age of the Late Triassic. It contains only one species: Gornogomphodon caffi.
Andescynodon is a genus of traversodontid cynodonts from the Middle Triassic of Argentina. Fossils are known from the Cerro de las Cabras and Cacheutá Formations. Andescynodon is one of the most basal traversodontids. Another traversodontid called Rusconiodon has also been identified from the Cerro de las Cabras Formation but is now considered a junior synonym of Andescynodon.
Scalenodon is an extinct genus of traversodontid cynodonts from the Middle Triassic of Africa and possibly Russia. The type species S. angustifrons was named in 1946 and several other species were named in the following years. Most of the species from Africa are now thought to belong to different genera than Scalenodon.
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.
Diademodontidae is an extinct family of Triassic gomphodonts. The best-known genus is Diademodon from South Africa. Titanogomphodon from Namibia may also be a member of Diademodontidae. The Chinese genera Hazhenia and Ordosiodon have also been included in the family, but were more recently identified as baurioid therocephalians. Remains of a diademodontid were reported in the Early-Middle Triassic Fremouw Formation in Antarctica, but that specimen was later referred to the trirachodontid Impidens
Titanogomphodon is an extinct genus of diademodontid cynodonts from the Middle Triassic Omingonde Formation of Namibia. It is known from a single partial skull that was described in 1973 from the Omingonde Formation. The type and only species is Titanogomphodon crassus. At about 40 centimetres (16 in), the skull of Titanogomphodon was significantly larger than that of its closest relative, Diademodon. Its teeth are similar to those of another group of cynodonts called Traversodontidae, but the similarities are likely the result of convergent evolution. Aside from its larger size, Titanogomphodon differs from Diademodon in having a bony projection on the postorbital bar behind the eye socket.
Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.
Aleodon is an extinct genus of cynodonts that lived from the Middle to the Late Triassic. Relatively few analyses have been conducted to identify the phylogenetic placement of Aleodon, however those that have place Aleodon as a sister taxon to Chiniquodon. Two species of Aleodon are recognized: A. brachyramphus which was discovered in Tanzania, and A. cromptoni which was discovered most recently in Brazil.
Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.
Etjoia is an extinct genus of traversodontid cynodonts that lived during the Middle Triassic or Late Triassic period in southern Africa. This medium-sized omnivorous cynognathian provides important information on the dental evolution of early diverging gomphodonts and traversodontids.
Arctotraversodontinae is a subfamily of Late Triassic cynodonts belonging to the family Traversodontidae. Members of the subfamily include Arctotraversodon, Boreogomphodon and Plinthogomphodon from North America, and Habayia, Maubeugia, Microscalenodon and Rosieria from Europe.