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Froghopper Temporal range: | |
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Prosapia bicincta | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hemiptera |
Suborder: | Auchenorrhyncha |
Superfamily: | Cercopoidea Leach, 1815 |
Families | |
The superfamily Cercopoidea, some members of which are called froghoppers and still others known as spittlebugs, are a group of hemipteran insects in the suborder Auchenorrhyncha. Adults are capable of jumping many times their height and length, giving the group their common name, but many species are best known for their plant-sucking nymphs which produce foam shelters, and are referred to as "spittlebugs".
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Relationships with other groups based on Cryan (2005). [1] |
Traditionally, most of this superfamily was considered a single family, the Cercopidae, but this family has been split into three families for many years now: the Aphrophoridae, Cercopidae, and Clastopteridae. The family Epipygidae has been removed from the Aphrophoridae by some authors (e.g. [2] ), but retained within Aphrophoridae by subsequent authors (e.g. [3] ).
These families are best known for the nymphal stage, which produces a cover of foamed-up plant sap visually resembling saliva; the nymphs are therefore commonly known as spittlebugs and their foam as cuckoo spit, frog spit, or snake spit. This characteristic spittle production is associated with the unusual trait of xylem feeding. Whereas most insects that feed on sap feed on the nutrient-rich fluid from the phloem, Cercopidae utilize the much more dilute sap flowing upward from the roots via the xylem. The large amount of excess water that must be excreted and the evolution of special breathing tubes allow the young spittlebug nymphs to grow in the relatively protective environment of the spittle. [4] Normally an animal should not be able to survive on a diet so low in nutrients, but the insects' digestive system contains symbiotic bacteria that provide them with the essential amino acids. [5]
The foam serves a number of purposes. It hides the nymph from the view of predators and parasites, and it insulates against heat and cold, thus providing thermal control and also moisture control; without the foam, the insect would risk dehydration. The nymphs pierce plants and suck sap causing very little damage; much of the filtered fluids go into the production of the foam, which has an acrid taste, deterring predators. A few species are serious agricultural pests.
A small family in the group, the Machaerotidae, known as the tube spittlebugs, is an outlier among the Cercopoidea because the nymphs live in calcareous tubes rather than producing foam as in the other families.
Adult froghoppers jump from plant to plant; some species can jump up to 70 cm (28 in) vertically: a more impressive performance relative to body weight than fleas. The froghopper can accelerate at 4,000 m/s2 (13,000 ft/s2) over 2 mm (0.079 in) as it jumps (experiencing over 400 gs of acceleration). [6] Spittlebugs can jump 100 times their own length. [7]
Many species of froghopper resemble leafhoppers, but can be distinguished by the possession of only a few stout spines on the hind tibiae, where leafhoppers have a series of small spines. Members of the family Machaerotidae greatly resemble treehoppers, due to a large thoracic spine, but the spine in machaerotids is an enlargement of the scutellum, where treehoppers have the pronotum enlarged. Members of the family Clastopteridae have their wings modified to form false heads at the tail end, an antipredator adaptation. Many adult Cercopidae can bleed reflexively from their tarsi, and the hemolymph appears to be distasteful; they are often aposematically colored.
The oldest froghoppers are known from the Early Jurassic. Mesozoic froghoppers are divided into two main families, Procercopidae known from the Early Jurassic to early Late Cretaceous of Asia, which are thought to be ancestral to living froghoppers, and Sinoalidae, which is known from the late Middle Jurassic and early Late Cretaceous of Asia. [8] The genus Qibinius the Middle Jurassic Yanan Formation of China mixes characters of both families and cannot be assigned to either. [9] The genus Cercopion from the Aptian aged Crato Formation of Brazil appears to be derived from the Procerocopidae and closely related to the crown group. [8]
Hemiptera is an order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from 1 mm (0.04 in) to around 15 cm (6 in), and share a common arrangement of piercing-sucking mouthparts. The name "true bugs" is often limited to the suborder Heteroptera.
Leafhopper is the common name for any species from the family Cicadellidae. These minute insects, colloquially known as hoppers, are plant feeders that suck plant sap from grass, shrubs, or trees. Their hind legs are modified for jumping, and are covered with hairs that facilitate the spreading of a secretion over their bodies that acts as a water repellent and carrier of pheromones. They undergo a partial metamorphosis, and have various host associations, varying from very generalized to very specific. Some species have a cosmopolitan distribution, or occur throughout the temperate and tropical regions. Some are pests or vectors of plant viruses and phytoplasmas. The family is distributed all over the world, and constitutes the second-largest hemipteran family, with at least 20,000 described species.
Treehoppers and thorn bugs are members of the family Membracidae, a group of insects related to the cicadas and the leafhoppers. About 3,200 species of treehoppers in over 400 genera are known. They are found on all continents except Antarctica; only five species are known from Europe. Individual treehoppers usually live for only a few months.
The Auchenorrhyncha suborder of the Hemiptera contains most of the familiar members of what was called the "Homoptera" – groups such as cicadas, leafhoppers, treehoppers, planthoppers, and spittlebugs. The aphids and scale insects are the other well-known "Homoptera", and they are in the suborder Sternorrhyncha.
Cicadomorpha is an infraorder of the insect order Hemiptera which contains the cicadas, leafhoppers, treehoppers, and spittlebugs. There are approximately 35,000 described species worldwide. Distributed worldwide, all members of this group are plant-feeders, and many produce either audible sounds or substrate vibrations as a form of communication. The earliest fossils of cicadomorphs first appear during the Late Permian. Notable extinct members include the "giant cicadas" belonging to Palaeontinidae.
A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment and that they often "hop" for quick transportation in a similar way to that of grasshoppers. However, planthoppers generally walk very slowly. Distributed worldwide, all members of this group are plant-feeders, though few are considered pests. Fulgoromorphs are most reliably distinguished from the other Auchenorrhyncha by two features; the bifurcate (Y-shaped) anal vein in the forewing, and the thickened, three-segmented antennae, with a generally round or egg-shaped second segment (pedicel) that bears a fine filamentous arista.
The superfamily Membracoidea of sap-sucking true-bugs includes two of the largest families within what used to be called the "Homoptera": the leafhoppers (Cicadellidae) and the treehoppers (Membracidae). The other families in this group are quite small, and have, at various points, generally been included as members within other families, though they are all presently considered to be valid, monophyletic groups. The relict family Myerslopiidae is restricted to New Zealand and South America while the Melizoderidae consist of two genera restricted to South America. The great diversity of Neotropical taxa suggests that the group originated in that region.
Cercopis vulnerata is a species of froghopper in the family Cercopidae.
Machaerotidae are a family of bugs in the superfamily Cercopoidea which were formerly placed within Cercopidae. They are sometimes called tube-forming spittle-bugs as the nymphs form a calcareous tube within which they live. These bugs are mainly found in the Old World tropics. The adults of many genera have a long, free and spine-like process originating from the scutellum and thus superficially similar to the tree-hoppers, Membracidae. The tegmen or forewing, like typical bugs of the suborder Heteroptera, always has a distinct, membranous apical area.
Cercopidae are the largest family of Cercopoidea, a xylem-feeding insect group, commonly called froghoppers. They belong to the hemipteran suborder Auchenorrhyncha. A 2023 phylogenetic study of the family suggested the elevation of subfamily Ischnorhininae to full family status as Ischnorhinidae, leaving a monophyletic Cercopinae.
Aphrophora alni, the European alder spittle bug, is a species belonging to the family Aphrophoridae.
Philaenus spumarius, the meadow froghopper or meadow spittlebug, is a species of insect belonging to the spittlebug family Aphrophoridae. In Italy and America, it is economically important as one of the vectors of Pierce's disease.
Cercopis sanguinolenta is a species of froghopper in the family Cercopidae.
Prosapia bicincta, the two-lined spittlebug, is a species of insect in the family Cercopidae. Adults are black with two red or orange lines crossing the wings. It reaches a length of 8–10 mm. It is widespread in the eastern half of the United States. A similar species, Prosapia simulans, can be found throughout Central America where it is considered an agricultural pest.
Aetalionidae are a family of treehoppers in the superfamily Membracoidea. Aetalionidae are somewhat like Membracidae in that they have one to three rows of short spines on the hind tibia but differ in having the front femur fused to the trochanter and the scutellum is completely exposed. The females have finger-like protrusions on the genital capsule. The family is mostly Neotropical. The subfamily Biturritiinae is Neotropical while the subfamily Aetalioninae has a Neotropical genus Aetalion and the sole Old World representative genus Darthula with a single species Darthula hardwickii.
Carystoterpa fingens, commonly known as the variegated spittlebug, is a spittlebug of the family Aphrophoridae. It is endemic to New Zealand.
Clastopteridae is a family of spittlebugs in the order Hemiptera. There are at least 10 genera and 100 described species in Clastopteridae.
Prosapia ignipectus, known generally as the red-legged spittlebug or black spittlebug, is a species of froghopper in the family Cercopidae. It is found in North America. It measures about 8 mm (0.3 in) in length.
Epipygidae is a lineage of froghoppers in the insect superfamily Cercopoidea. There are at least three genera and about five described species in Epipygidae, found in the American tropics. In addition, there are more than 20 undescribed species in the family. Molecular analyses indicate that the group is monophyletic, but is clearly nested within the family Aphrophoridae and is probably best relegated to the status of a subfamily or tribe, rather than retained as a separate family.
Sinoalidae is an extinct family of froghoppers known from the late Middle Jurassic to the early Late Cretaceous of Asia. They are one of two main Mesozoic families of froghoppers, alongside Procercopidae, unlike Procercopidae, Sinoalidae is thought to be an extinct side branch and not ancestral to modern froghoppers. Sinoalids have a temporally disjunct distribution being only known from the late Middle Jurassic (Callovian) Yanliao Biota of Inner Mongolia and the early Late Cretaceous (Cenomanian) aged Burmese amber of Myanmar, separated by over 60 million years. The family is "recognized by its tegmen with the costal area and clavus commonly more sclerotized and punctate than the remaining part, and its hind tibia with two rows of lateral spines"