List of non-avian theropod type specimens

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This list of non-avian theropod type specimens is a list of fossils that are the official standard-bearers for inclusion in the Mesozoic species and genera of the dinosaur clade Theropoda, which includes the carnivorous dinosaurs like Tyrannosaurus and Velociraptor , their herbivorous relatives like the therizinosaurs, and birds. Type specimens are those that are definitionally members of biological taxa, and additional specimens can only be "referred" to these taxa if an expert deems them sufficiently similar to the type. Theropods have been found all over the globe, and are alive today as birds

Contents

The list

SpeciesGenusNicknameCatalogue numberInstitutionAgeUnitCountryNotesImages
Abelisaurus comahuensis Abelisaurus MC 11098 [1] Museo de Cipolleti Maastrichtian [1] Allen Formation [1] Argentina [1]
Reconstructed skull of Abelisaurus comahuensis, with bones from the type specimen Abelisaurus skull.jpg
Reconstructed skull of Abelisaurus comahuensis, with bones from the type specimen
Acheroraptor temertyorum Acheroraptor ROM 63777 [2] Royal Ontario Museum Upper Maastrichtian [2] Hell Creek Formation [2] United States [2]
Cast of the type specimen of Acheroraptor temertyorum Acheroraptor NMNH.jpg
Cast of the type specimen of Acheroraptor temertyorum
Achillesaurus manazzonei Achillesaurus MACN-PV-RN 1116 [3] Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” Santonian [3] Bajo de la Carpa Formation [3] Argentina [3]
Achillobator giganticus Achillobator MNUFR-15 [4] Mongolian National University Santonian-Campanian [4] Bayan Shireh Formation [4] Mongolia [4]
Skeletal reconstruction of Achillobator giganticus illustrating holotype material Achillobator Skeleton Reconstruction.jpg
Skeletal reconstruction of Achillobator giganticus illustrating holotype material
Acrocanthosaurus atokensis Acrocanthosaurus Holotype: OMNH 10146 [5]

Paratype: OMNH 10147 [5]

Oklahoma Museum of Natural History Albian-Aptian [5] Antlers Formation [5] United States [6]
Skeletal diagrams of Acrocanthosaurus atokensis specimens. The holotype is the topmost image, while the second image from the top is the paratype Acrocanthosaurus skeletons.tif
Skeletal diagrams of Acrocanthosaurus atokensis specimens. The holotype is the topmost image, while the second image from the top is the paratype
Adasaurus mongoliensis Adasaurus IGM 100/20 [7] Mongolian Institute of Geology Late Cretaceous [7] Nemegt Formation [7] Mongolia [7]
Reconstructed skull of Adasaurus mongoliensis. Material in white indicates known material from the holotype Adasaurus Skull.png
Reconstructed skull of Adasaurus mongoliensis. Material in white indicates known material from the holotype
Aepyornithomimus tugrikinensis Aepyornithomimus MPC-D 100/130 [8] Institute of Paleontology and Geology, Mongolian Academy of Sciences Campanian [8] Djadokhta Formation [8] Mongolia [8]
Holotype metatarsals of Aepyornithomimus tugrikinensis Aepyornithomimus tugrikinensis.jpg
Holotype metatarsals of Aepyornithomimus tugrikinensis
Aerosteon riocoloradensis Aerosteon MCNA-PV-3137 [9] Museo de Ciencias Naturales y Antropológicas (J. C. Moyano) de Mendoza Santonian [9] Anacleto Formation [9] Argentina [9]
Cast of the holotype left ilium of Aerosteon riocoloradensis Aerosteon ilium photos.png
Cast of the holotype left ilium of Aerosteon riocoloradensis
Afromimus tenerensis Afromimus MNBH GAD112 [10] Musée National Boubou Hama Aptian-Albian [10] Elrhaz Formation [10] Niger [10]
Afrovenator abakensis Afrovenator UC OBA 1 [11] University of Chicago Middle Jurassic [12] Tiouaren Formation [12] Niger [11]
Reconstruction of the skull of Afrovenator abakensis based on the holotype Afrovenator reconstructed skull.png
Reconstruction of the skull of Afrovenator abakensis based on the holotype
Albertavenator curriei Albertavenator TMP 1993.105.0001 [13] Royal Tyrell Museum of Paleontology Earliest Maastrichtian [13] Horsethief Member, Horseshoe Canyon Formation [13] Canada [13]
Illustration of the holotype frontal of Albertavenator curriei Albertavenator.png
Illustration of the holotype frontal of Albertavenator curriei
Albertonykus borealis Albertonykus TMP 2001.45.91 [14] Royal Tyrell Museum of Paleontology Maastrichtian [14] Horseshoe Canyon Formation [14] Canada [14]
Albertosaurus arctunguis Albertosaurus ROM 807 [15] Royal Ontario Museum Campanian-Maastrichtian [16] Horseshoe Canyon Formation [16] Canada [16] Synonymous with Albertosaurus sarcophagus [15]
Albertosaurus sarcophagus Albertosaurus CMN 5600 [16] Canadian Museum of Nature Campanian-Maastrichtian [16] Horseshoe Canyon Formation [16] Canada [16]
CMN 5600 Tyrrell - Crane Albertosaure.jpg
CMN 5600
Albinykus baatar Albinykus IGM 100/3004 [17] Mongolian Institute of Geology Santonian-Campanian [17] Javkhlant Formation [17] Mongolia [17]
Alectrosaurus olseni Alectrosaurus AMNH 6368 [18] American Museum of Natural History Upper Cretaceous [18] Iren Dabasu Formation [18] Mongolia [18]
Holotype right foot of Alectrosaurus olseni Alectrosaurm olseni.jpg
Holotype right foot of Alectrosaurus olseni
Alioramus altai Alioramus IGM 100/1844 [19] Mongolian Institute of Geology Maastrichtian [19] Nemegt Formation [19] Mongolia [19]
Holotype skull bones of Alioramus altai Alioramus altai skull.png
Holotype skull bones of Alioramus altai
Alioramus remotus Alioramus PIN 3141/1 [20] Paleontological Institute, Russian Academy of Sciences Late Cretaceous [19] Mongolia [19]
Allosaurus europaeus Allosaurus ML415 [21] Museu da Lourinhã Kimmeridgian [21] Porto Novo Member, Lourinhã Formation [21] Portugal [21]
Holotype skull (C) of Allosaurus europaeus Skulls of Allosaurus species.png
Holotype skull (C) of Allosaurus europaeus
Allosaurus fragilis Allosaurus YPM 1930 [22] Peabody Museum of Natural History Upper Jurassic [22] Lower Morrison Formation [22] Colorado [22]
Allosaurus jimmadseni Allosaurus DINO 11541 [23] Dinosaur National Monument Kimmeridgian [23] Morrison Formation [23] United States [23]
Holotype skull (B) of Allosaurus jimmadseni Skulls of Allosaurus species.png
Holotype skull (B) of Allosaurus jimmadseni
Almas ukhaa Almas IGM 100/1323 [24] Mongolian Institute of Geology Campanian [24] Djadokhta Formation [24] Mongolia [24]
Alnashetri cerropoliciensis Alnashetri MPCA-477 [25] Museo Provincial Carlos Ameghino Cenomanian-Turonian [25] Candeleros Formation [25] Argentina [25]
Altispinax dunkeri Altispinax NHMUK R1828 [26] Natural History Museum, London Berriasian-Valanginian [27] Hasting Beds Group [27] United Kingdom [27]
Illustration of the holotype specimen of Altispinax dunkeri Becklespinax.jpg
Illustration of the holotype specimen of Altispinax dunkeri
Alvarezsaurus calvoi Alvarezsaurus MUCPv 54 [28] Museo de Ciencias Naturales, Universidad Nacional del Comahue Coniacian-Santonian [28] Bajo de la Carpa Member, Río Colorado Formation [28] Argentina [28]
Skeletal diagram of Alvarezsaurus calvoi indicating holotype elements Alvarezsaurus Jaime Headden.jpg
Skeletal diagram of Alvarezsaurus calvoi indicating holotype elements
Alxasaurus elesitaiensis Alxasaurus IVPP 88402 [29] Institute of Vertebrate Paleontology and Paleoanthropology Albian [29] Bayin Gobi Formation [29] China [29]
Holotype dentary of Alxasaurus elesitaiensis Therizinosaurian mandibular morphology (Alxasaurus jaw).png
Holotype dentary of Alxasaurus elesitaiensis
Ambopteryx longibrachium Ambopteryx IVPP V24192 [30] Institute of Vertebrate Paleontology and Paleoanthropology Oxfordian [30] China [30]
Anchiornis huxleyi Anchiornis IVPP V14378 [31] Institute of Vertebrate Paleontology and Paleoanthropology Oxfordian [32] Tiaojishan Formation [32] China [31]
Aniksosaurus darwini Aniksosaurus MDT-PV 1/48 [33] Museo Desiderio Torres Late Cretaceous [33] Bajo Barreal Formation [33] Argentina [33]
Holotype tibia (C and D) of Aniksosaurus darwini Aniksosaurus.png
Holotype tibia (C and D) of Aniksosaurus darwini
Anomalipes zhaoi Anomalipes ZCDM V0020 [34] Zhucheng Dinosaur Museum Upper Cretaceous [34] Wangshi Group [34] China [34]
Holotype pedal elements of Anomalipes zhaoi Anomalipes pes.jpg
Holotype pedal elements of Anomalipes zhaoi
Anserimimus planinychus Anserimimus IGM 100/300 [35] Mongolian Institute of Geology Maastrichtian [36] Nemegt Formation [36] Mongolia [36]
Antrodemus valens Antrodemus USNM 218 [4] National Museum of Natural History Upper Jurassic Morrison Formation [4] United States [4] Nomen dubium [22]
Anzu wyliei Anzu CM 78000 [37] Carnegie Museum of Natural History Upper Maastrichtian [37] Hell Creek Formation [37] United States [37]
Holotype elements (Left: C-F, I-L. Right: F-G) of Anzu wyliei Anzu wyliei.png
Holotype elements (Left: C-F, I-L. Right: F-G) of Anzu wyliei
Aoniraptor libertatem Aoniraptor MPCA-Pv 804/1 to 804/25 [38] Museo Provincial “Carlos Ameghino” Cenomanian-Turonian [38] Huincul Formation [38] Argentina [38]
Aorun zhaoi Aorun IVPP V15709 [39] Institute of Vertebrate Paleontology and Paleoanthropology Oxfordian-Callovian [39] Shishugou Formation [39] China [39]
Apatoraptor pennatus Apatoraptor TMP 1993.051.0001 [40] Royal Tyrell Museum of Paleontology Campanian-Maastrichtian [40] Horsethief Member, Horseshoe Canyon Formation [40] Canada [40]
Holotype skeleton of Apatoraptor pennatus Dinosaur fossil (1).jpg
Holotype skeleton of Apatoraptor pennatus
Appalachiosaurus montgomeriensis Appalachiosaurus RMM 6670 [41] McWane Science Center Campanian [41] Demopolis Formation [41] United States [41]
Aratasaurus museunacionali Aratasaurus MPSC R 2089 [42] Museu de Paleontologia, Universidade Regional do Cariri Aptian [42] Romualdo Formation [42] Brazil [42]
Holotype leg bones of Aratasaurus museunacionali Aratasaurus museunacionali holotype.jpg
Holotype leg bones of Aratasaurus museunacionali
Archaeopteryx albersdoerferi Archaeopteryx The Daiting specimenSNSB BSPG VN-2010/1 [43] Bayerische Staatssammlung für Paläontologie und Geologie Lower Tithonian [43] Mörnsheim Formation [43] Germany [43]
The Daiting specimen, the holotype of Archaeopteryx albersdoerferi Archaeopteryx (Daiting Specimen).jpg
The Daiting specimen, the holotype of Archaeopteryx albersdoerferi
Archaeopteryx lithographica Archaeopteryx The London specimenNHMUK 37001 [44] Natural History Museum, London Lower Tithonian [43] Solnhofen Limestone [44] Germany [45]
Replica of the London specimen, the neotype of Archaeopteryx lithographica Archaeopteryx lithographica, replica of London specimen, Staatliches Museum fur Naturkunde Karlsruhe, Germany - 20100925.jpg
Replica of the London specimen, the neotype of Archaeopteryx lithographica
Archaeopteryx siemensiiArchaeopteryxThe Berlin specimenHMN 1880/81 [46] Museum für Naturkunde Lower Tithonian [43] Solnhofen Limeston [46] Germany [46]
The Berlin specimen, the holotype specimen of Archaeopteryx siemensii Archaeopteryx lithographica (Berlin specimen).jpg
The Berlin specimen, the holotype specimen of Archaeopteryx siemensii
Archaeornithoides deinosauriscus Archaeornithoides ZPAL MgD-II/29 [5] Institute of Paleobiology, Polish Academy of Sciences Late Santonian-Early Campanian [5] Djadokhta Formation [5] Mongolia [5]
Reconstruction of the skull of Archaeornithoides deinosauriscus. Holotype material is shown in brown Archaeornithoides skull.png
Reconstruction of the skull of Archaeornithoides deinosauriscus. Holotype material is shown in brown
Archaeornithomimus asiaticus Archaeornithomimus AMNH 6565 [47] American Museum of Natural History Cenomanian? [47] Iren Dabasu Formation [47] China [47]
Arcovenator escotae Arcovenator MHNA-PV-2011.12.1, MHNA-PV-2011.12.2, MHNA.PV.2011.12.15, MHNA.PV.2011.12.5, MHNA.PV.2011.12.4, MHNA.PV.2011.12.3 [48] Muséum d’Histoire Naturelle d’Aix-en-Provence Late Campanian [48] Argiles Rutilantes Formation [48] France [48]
Aristosuchus pusilius Aristosuchus NHMUK R178 [49] Natural History Museum, London Barremian [49] Wessex Formation [49] United Kingdom [49]
Arkansaurus fridayi Arkansaurus UAM 74-16 [50] University of Arkansas Museum Albian-Aptian [50] Trinity Group [50] United States [50]
Asfaltovenator vialidadi Asfaltovenator MPEF PV 3440 [51] Museo Paleontológico Egidio Feruglio [51] Late Toarcian-Bajocian [51] Cañadón Asfalto Formation [51] Argentina [51]
Holotype skull of Asfaltovenator vialidadi Asfaltovenatorskull.png
Holotype skull of Asfaltovenator vialidadi
Atrociraptor marshalli Atrociraptor TMP 95.166.1 [52] Royal Tyrell Museum of Paleontology Upper Campanian-Lower Maastrichtian [52] Horseshoe Canyon Formation [52] Canada [52]
Holotype skull of Atrociraptor marshalli Atrociraptor royal tyrrell.jpg
Holotype skull of Atrociraptor marshalli
Aublysodon mirandus Aublysodon ANSP 9535 [53] Academy of Natural Sciences of Philadelphia Judith River Formation [54] United States [54] Type specimen now lost, might be a nomen dubium [53]
Holotype tooth (41-45) of Aublysodon mirandus Deinodon.JPG
Holotype tooth (41-45) of Aublysodon mirandus
Aucasaurus garridoi Aucasaurus MCF-PVPH-236 [55] Museo Municipal Carmen Funes Campanian [55] Anacleto Member, Río Colorado Formation [55] Argentina [55]
Holotype braincase of Aucasaurus garridoi Aucasaurus braincase.jpg
Holotype braincase of Aucasaurus garridoi
Aurornis xui Aurornis YFGP-T5198 [7] Yizhou Fossil and Geology Park Middle-Late Jurassic [7] Tiaojishan Formation [7] China [7]
Australovenator wintonensis Australovenator AODF 604 [56] Australian Age of Dinosaurs Albian [56] Winton Formation [56] Australia [56]
Holotype dentary of Australovenator wintonensis Australovenator dentary.png
Holotype dentary of Australovenator wintonensis
Austrocheirus isasii Austrocheirus MPM-PV 10003 [57] Museo Padre Molina Campanian [58] Pari Aike Formation [57] Argentina [57]
Austroraptor cabazai Austroraptor MML-195 [59] Museo Municipal de Lamarque Campanian-Maastrichtian [59] Allen Formation [59] Argentina [59]
Skeletal reconstruction of Austroraptor cabazai illustrating the holotype material Austroraptor.jpg
Skeletal reconstruction of Austroraptor cabazai illustrating the holotype material
Aviatyrannis jurassica Aviatyrannis IPFUB Gui Th 1 [60] Institut für Geologische Wissenschaften, Freie Universtät Berlin Kimmeridgian [60] Alcobaça Formation [60] Portugal [60]
Avimimus nemegtensis Avimimus MPC-D 102/81 [61] Institute of Paleontology and Geology, Mongolian Academy of Sciences Late Cretaceous [61] Nemegt Formation [61] Mongolia [61]
Avimimus portentosus Avimimus PIN 3907-1 [61] Paleontological Institute, Russian Academy of Sciences Upper Cretaceous [62] Nemegt Formation [62] Mongolia [62]
Bagaraatan ostromi Bagaraatan ZPAL MgD-I/108 [63] Institute of Paleobiology, Polish Academy of Sciences Middle Maastrichtian? [63] Nemegt Formation [63] Mongolia [63]
Holotype left mandible of Bagaraatan ostromi Bagaraatan.jpg
Holotype left mandible of Bagaraatan ostromi
Bahariasaurus ingens Bahariasaurus BSP 1922 X 47 [64] Bayerische Staatssammlung für Paläontologie und historische Geologie, München Cenomanian [65] Bahariya Formation [65] Egypt [65] Type specimen was destroyed in World War II
Balaur bondoc Balaur EME PV.313 [66] Transylvanian Museum Society, Cluj-Napoca Maastrichtian [66] Sebeş Formation [66] Romania [66]
Holotype specimen of Balaur bondoc Balaur bondoc 1.jpg
Holotype specimen of Balaur bondoc
Bambiraptor feinbergi Bambiraptor Bambi [67] AMNH FR 30556 [68] American Museum of Natural History Upper Cretaceous [68] Two Medicine Formation [68] Montana [68]
Skeletal reconstruction of Bambiraptor feinbergi based on the holotype Bambiraptor feinbergi.jpg
Skeletal reconstruction of Bambiraptor feinbergi based on the holotype
Banji long Banji IVPP V 16896 [69] Institute of Vertebrate Paleontology and Paleoanthropology Upper Cretaceous [69] Nanxiong Formation? [69] China [69]
Holotype skull of Banji long Banji skull.png
Holotype skull of Banji long
Bannykus wulatensis Bannykus IVPP V25026 [70] Institute of Vertebrate Paleontology and Paleoanthropology Aptian [70] Bayingobi Formation [70] China [70]
Holotype of Bannykus wulatensis Wulatesaurus.jpg
Holotype of Bannykus wulatensis
Baryonyx walkeri Baryonyx Claws [71] NHMUK VP R9951 [72] Natural History Museum, London Barremian [71] Weald Clay [71] United Kingdom [71]
Holotype elements of Baryonyx walkeri Baryonyx fossil elements.jpg
Holotype elements of Baryonyx walkeri
Beibeilong sinensis Beibeilong Baby Louie [73] HGM 41HIII1219 [73] Henan Geological Museum Cenomanian-Turonian [73] Gaogou Formation [73] China [73]
Holotype skeleton of Beibeilong sinensis Skeleton of Beibeilong sinensis (HGM 41HIII1219).jpg
Holotype skeleton of Beibeilong sinensis
Beipiaosaurus inexpectus Beipiaosaurus IVPP V11559 [74] Institute of Vertebrate Paleontology and Paleoanthropology Lower Cretaceous [74] Yixian Formation [74] China [74]
Holotype elements of Beipiaosaurus inexpectus Beipiaosaurus holotype fossil.png
Holotype elements of Beipiaosaurus inexpectus
Beishanlong grandis Beishanlong FRDC-GS GJ (06) 01-18 [75] Fossil Research and Development Center, Third Geology and Mineral Resources Exploration Academy, Gansu Provincial Bureau of Geo-Exploration and Mineral Development Aptian-Albian [75] Xinminpu Group [75] China [75]
Berberosaurus liassicus Berberosaurus MHNM-Pt9, Pt23, Pt22, Pt19, Pt21, Pt16, Pt20 [76] Muséum d’Histoire Naturelle de Marrakech Pliensbachian-Toarcian [76] Toundoute Continental Series [76] Morocco [76]
Berthasaura leopoldinae Berthasaura MN 7821-V Museu Nacional, Universidade Federal do Rio de Janeiro Aptian-Albian Goio Erê Formation Brazil
Skeletal reconstruction of Berthasaura leopoldinae displaying several bones from the holotype specimen Berthasaura.webp
Skeletal reconstruction of Berthasaura leopoldinae displaying several bones from the holotype specimen
Betasuchus bredai Betasuchus NHMUK R 42997 [77] Natural History Museum, London Maastrichtian [78] Maastricht Formation? [78] Netherlands [78]
Cast of the holotype of Betasuchus bredai Betasuchus bredai.jpg
Cast of the holotype of Betasuchus bredai
Bicentenaria argentina Bicentenaria MPCA 865 [79] Museo Provincial "Carlos Ameghino" Cenomanian [79] Candeleros Formation [79] Argentina [79]
Holotype skull elements of Bicentenaria argentina Bicentenaria skull elements.png
Holotype skull elements of Bicentenaria argentina
Bistahieversor sealeyi Bistahieversor NMMNH P-27469 [80] New Mexico Museum of Natural History and Science Campanian [80] Hunter Wash Member, Kirtland Formation [80] United States [80]
Holotype skull of Bistahieversor sealeyi Bistahieversor.jpg
Holotype skull of Bistahieversor sealeyi
Bonapartenykus ultimus Bonapartenykus MPCA 1290 [81] Museo Provincial "Carlos Ameghino" Campanian-Maastrichtian [81] Allen Formation [81] Argentina [81]
Boreonykus certekorum Boreonykus TMP 1989.055.0047 [82] Royal Tyrell Museum of Paleontology Campanian [82] Wapiti Formation [82] Canada [82]
Borogovia gracilicrus Borogovia ZPAL MgD-I/174 [83] Institute of Paleobiology, Polish Academy of Sciences Maastrichtian [83] Nemegt Formation [83] Mongolia [83]
Holotype tibiotarsus of Borogovia gracilicrus Borogovia ZPAL MgD-I 174 right tibiotarsus.png
Holotype tibiotarsus of Borogovia gracilicrus
Bradycneme draculae Bradycneme BMNH A1588 [84] Natural History Museum, London Maastrichtian [85] Hațeg Basin [86] Romania [85]
Buitreraptor gonzalezorum Buitreraptor MPCA 245 [87] Museo Provincial "Carlos Ameghino" Cenomanian-Turonian [87] Candeleros Formation [87] Argentina [87]
Reconstruction of Buitreraptor gonzalezorum based on the holotype skeleton Reconstruccion Buitreraptor gonzalezorum.jpg
Reconstruction of Buitreraptor gonzalezorum based on the holotype skeleton
Byronosaurus jaffei Byronosaurus IGM 100/983 [88] Mongolian Institute of Geology Late Cretaceous [88] Djadokhta Formation [88] Mongolia [88]
Skeletal reconstruction of Byronosaurus jaffei based on the holotype material Byronosaurus jaffei headden.jpg
Skeletal reconstruction of Byronosaurus jaffei based on the holotype material
Caenagnathasia martinsoni Caenagnathasia CCMGE 401/12457 [89] Chernyshev's Central Museum of Geological Exploration Turonian [90] Bissekty Formation [90] Uzbekistan [90]
Holotype and paratype mandibles of Caenagnathasia martinsoni Caenagnathasia martinsoni.jpg
Holotype and paratype mandibles of Caenagnathasia martinsoni
Caenagnathus collinsi Caenagnathus CMN 8776 [91] Canadian Museum of Nature Campanian [91] Dinosaur Park Formation [91] Canada [91]
Holotype mandible of Caenagnathus collinsi Caenagnathus mandible.jpg
Holotype mandible of Caenagnathus collinsi
Caihong juji Caihong PMoL-B00175 [92] Paleontological Museum of Liaoning Oxfordian [92] Tiaojishan Formation [92] China [92]
Holotype skeleton of Caihong juji Caihong skeleton.jpg
Holotype skeleton of Caihong juji
Calamosaurus foxi Calamosaurus BMNH R901 [26] Natural History Museum, London Barremian [26] Wealden Group [93] United Kingdom [26]
Holotype vertebrae of Calamosaurus foxi Calamosaurus.jpg
Holotype vertebrae of Calamosaurus foxi
Calamospondylus oweni Calamospondylus Upper Barremian [26] Vectis Formation [26] United Kingdom [93] Type specimen has been lost [93]
Camarillasaurus cirugedae Camarillasaurus MPG-KPC1-46 [94] Museo Paleontológico de Galve Lower Barremian [94] Camarillas Formation [94] Spain [94]
Holotype ribs and vertebrae of Camarillasaurus cirugedae Camarillasaurus.jpg
Holotype ribs and vertebrae of Camarillasaurus cirugedae
Camposaurus arizonensis Camposaurus UCMP 34498 [95] University of California Museum of Paleontology Lower-Middle Norian [95] Mesa Redondo Member, Chinle Formation [95] United States [95]
Holotype ankle of Camposaurus arizonensis Camposaurus holotype ankle.png
Holotype ankle of Camposaurus arizonensis
Carcharodontosaurus iguidensis Carcharodontosaurus MNN IGU2 [96] Musée National du Niger [96] Cenomanian [96] Echkar Formation [96] Niger [96]
Carcharodontosaurus saharicus Carcharodontosaurus SGM-Din 1 [96] Ministére de l’Energie et des Mines, Rabat [96] Cenomanian [96] Kem Kem Group [96] Morocco [96]
Carnotaurus sastrei Carnotaurus MACN-CH 894 [97] Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” Maastrichtian [98] La Colonia Formation [98] Argentina [97]
Holotype skull of Carnotaurus sastrei Carnotaurus skull.jpg
Holotype skull of Carnotaurus sastrei
Caudipteryx dongi Caudipteryx IVPP V12344 [99] Institute of Vertebrate Paleontology and Paleoanthropology Barremian [100] Yixian Formation [99] China [99]
Caudipteryx zoui Caudipteryx NGMC 97-4-A [101] National Geological Museum of China Barremian [100] Jiulongsong Member, Chaomidianzi Formation [101] China [101]
Holotype skeleton of Caudipteryx zoui Caudipteryx-Geological Museum of China.jpg
Holotype skeleton of Caudipteryx zoui
Ceratonykus oculatus Ceratonykus MPC 100/124 [102] Paleontological Center, Mongolian Academy of Sciences Santonian [102] Baruungoyot Formation [102] Mongolia [102]
Ceratosaurus dentisulcatus Ceratosaurus UMNH 5278 [103] Utah Museum of Natural History Late Jurassic [103] Brushy Basin Member, Morrison Formation [103] United States [103] Possible junior synonym of Ceratosaurus nasicornis [64]
Ceratosaurus magnicornis Ceratosaurus MWC 1 [103] Museum of Western Colorado Late Jurassic [103] Brushy Basin Member, Morrison Formation [103] United States [103] Possible junior synonym of Ceratosaurus nasicornis [64]
Ceratosaurus nasicornis Ceratosaurus USNM 4735 [64] National Museum of Natural History Late Jurassic [104] Morrison Formation [104] United States [104]
Holotype skeleton of Ceratosaurus nasicornis Stegosaurus and Ceratosaurus in Smithsonian National Museum.jpg
Holotype skeleton of Ceratosaurus nasicornis
Ceratosuchops inferodios Ceratosuchops IWCMS 2014.95.1-3, IWCMS 2014.95.5, IWCMS 2021.30 [105] Dinosaur Isle Museum Barremian [105] Wessex Formation [105] United Kingdom [105]
Holotype skull of Ceratosuchops inferodios Ceratosuchops holotype.webp
Holotype skull of Ceratosuchops inferodios
Changyuraptor yangi Changyuraptor HG B016 [106] Paleontological Center, Bohai University Early Cretaceous [106] Yixian Formation [106] China [106]
Chenanisaurus barbaricus Chenanisaurus OCP DEK-GE 772 [107] Office Cherifien des Phosphates Maastrichtian [107] Ouled Abdoun Basin [107] Morocco [107]
Holotype dentary of Chenanisaurus barbaricus Chenanisaurus barbaricus jaw by Nick Longrich.jpg
Holotype dentary of Chenanisaurus barbaricus
Chienkosaurus ceratosauroides Chienkosaurus IVPP V237 [108] Institute of Vertebrate Paleontology and Paleoanthropology Tithonian [64] Kuangyuan Series [64] China [64] Dubious [108]
Chilantaisaurus tashuikouensis Chilantaisaurus IVPP V2884.1 [109] [110] Institute of Vertebrate Paleontology and Paleoanthropology Late Cretaceous [109] Ulansuhai Formation [110] China [109]
Chingkankousaurus fragilis Chingkankousaurus IVPP V836 [111] Institute of Vertebrate Paleontology and Paleoanthropology Campanian - Maastrichtian [111] Wangshi Series [111] China [111]
Chirostenotes pergracilis Chirostenotes CMN 2367 [112] Canadian Museum of Nature Upper Campanian [112] Dinosaur Park Formation [112] Canada [112]
Holotype hands of Chirostenotes pergracilis Chirostenotes pergracilis hand.jpg
Holotype hands of Chirostenotes pergracilis
Chuandongocoelurus primitivus Chuandongocoelurus CCG 20010 [110] Chengdu University of Geology Middle Jurassic [110] Xiashaximiao Formation [110] China [110]
Citipati osmolskae Citipati IGM 100/978 [113] Mongolian Institute of Geology Upper Cretaceous [113] Djadokhta Formation [113] Mongolia [113]
Citipes elegans Citipes ROM 781 [114] Royal Ontario Museum Campanian [114] Dinosaur Park Formation [114] Canada [114]
Holotype foot of Citipes elegans Caenagnathid tarsometatarsi.jpg
Holotype foot of Citipes elegans
Coelophysis bauri Coelophysis AMNH 7224 [115] American Museum of Natural History Upper Triassic [116] Chinle Formation [116] United States [115] Neotype [115]
Two skeletons of Coelophysis bauri, including the neotype specimen Coelophysis bauri.jpg
Two skeletons of Coelophysis bauri, including the neotype specimen
Coeluroides largus Coeluroides GSI K27/562, K27/574, K27/595 [117] Geological Survey of India Maastrichtian [117] Lameta Formation [117] India [117]
Coelurus fragilis Coelurus YPM 1991, 1993 [118] Yale Peabody Museum Upper Jurassic [118] Morrison Formation [118] United States [118] Syntypes [118]
Compsognathus longipes Compsognathus SNSB-BSPG AS I 563 [119] Bavarian State Collection for Paleontology and Geology Tithonian [119] Solnhofen Limestone [119] Germany [119]
Cast of the holotype specimen of Compsognathus longipes Compsognathus longipes 01.JPG
Cast of the holotype specimen of Compsognathus longipes
Compsosuchus solus Compsosuchus GSI K27/578 [117] Geological Survey of India Maastrichtian [117] Lameta Formation [117] India [117]
Concavenator corcovatus Concavenator MCCM-LH 6666 [120] Museo de las Ciencias de Castilla-La Mancha Upper Barremian [120] Calizas de La Huérguina Formation [120] Spain [120]
Holotype of Concavenator corcovatus Concavenator.jpg
Holotype of Concavenator corcovatus
Conchoraptor gracilis Conchoraptor MPC-D 100/20 [61] Institute of Paleontology and Geology, Mongolian Academy of Sciences Late Cretaceous [61] Barun Goyot Formation [61] Mongolia [61]
Condorraptor currumilli Condorraptor MPEF-PV 1672 [121] Museo Paleontológico Egidio Feruglio Callovian [121] Cañadón Asfalto Formation [121] Argentina [121]
Corythoraptor jacobsi Corythoraptor JPM-2015-001 [122] Jinzhou Paleontological Museum Campanian - Maastrichtian [122] Nanxiong Formation [122] China [122]
Holotype of Corythoraptor jacobsi Corythoraptor.jpg
Holotype of Corythoraptor jacobsi
Cristatusaurus lapparenti Cristatusaurus MNHN GDF 366 [123] Muséum National d'Histoire Naturelle Aptian [123] Elrhaz Formation [124] Niger [123]
Holotype specimen of Cristatusaurus lapparenti Cristatusaurus.jpg
Holotype specimen of Cristatusaurus lapparenti
Cruxicheiros newmanorum Cruxicheiros WARMS G15770 [125] Warwickshire Museum Service Lower Bathonian [125] Chipping Norton Limestone [125] United Kingdom [125]
Holotype femur (C-E) of Cruxicheiros newmanorum Cruxicheiros newmanorum.jpg
Holotype femur (C-E) of Cruxicheiros newmanorum
Cryolophosaurus ellioti Cryolophosaurus FMNH PR1821 [126] Field Museum of Natural History Early Jurassic [127] Hanson Formation [127] Antarctica [126]
Holotype skull of Cryolophosaurus ellioti Cryolophosaurus ellioti holotype.jpg
Holotype skull of Cryolophosaurus ellioti
Tyrannosaurus rex Tyrannosaurus CM 9380 Carnegie Museum of Natural History United States
CM 9380 Tyrannosaurus Rex Holotype.jpg
CM 9380

See also

Related Research Articles

<span class="mw-page-title-main">Therizinosauria</span> Extinct clade of dinosaurs

Therizinosaurs are an extinct group of large herbivorous theropod dinosaurs whose fossils have been found across the Middle Jurassic to Late Cretaceous deposits in Europe, Asia and North America. Various features of the forelimbs, skull and pelvis unite these finds as both theropods and maniraptorans, making them relatives of birds. The name of the representative genus, Therizinosaurus, is derived from the Greek θερίζω and σαῦρος. The older representative, Segnosaurus, is derived from the Latin sēgnis ('slow') and the Greek σαῦρος.

<i>Byronosaurus</i> Extinct genus of dinosaurs

Byronosaurus is a genus of troodontid dinosaur from the Late Cretaceous Period of Mongolia.

<span class="mw-page-title-main">Philip J. Currie</span> Canadian paleontologist and curator

Philip John Currie is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the 1980s, he became the director of the Canada-China Dinosaur Project, the first cooperative palaeontological partnering between China and the West since the Central Asiatic Expeditions in the 1920s, and helped describe some of the first feathered dinosaurs. He is one of the primary editors of the influential Encyclopedia of Dinosaurs, and his areas of expertise include theropods, the origin of birds, and dinosaurian migration patterns and herding behavior. He was one of the models for palaeontologist Alan Grant in the film Jurassic Park.

<i>Fukuiraptor</i> Megaraptoran theropod dinosaur genus from the Early Cretaceous epoch

Fukuiraptor was a medium-sized megaraptoran theropod dinosaur of the Early Cretaceous epoch that lived in what is now Japan. Fukuiraptor is known from the Kitadani Formation and possibly also the Sebayashi Formation.

<i>Rugops</i> Genus of dinosaur

Rugops is a monospecific genus of basal abelisaurid theropod dinosaur from Niger that lived during the Late Cretaceous period in what is now the Echkar Formation. The type and only species, Rugops primus, is known only from a partial skull. It was named and described in 2004 by Paul Sereno, Jeffery Wilson and Jack Conrad. Rugops has an estimated length of 4.4–5.3 metres (14–17 ft) and weight of 410 kilograms (900 lb). The top of its skull bears several pits which correlates with overlaying scale and the front of the snout would have had an armour-like dermis.

<i>Archaeornithomimus</i> Extinct genus of dinosaurs

Archaeornithomimus is a genus of ornithomimosaurian theropod dinosaur that lived in Asia during the Late Cretaceous period, around 96 million years ago in the Iren Dabasu Formation.

<span class="mw-page-title-main">Barun Goyot Formation</span>

The Barun Goyot Formation is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia.

<span class="mw-page-title-main">Nemegt Formation</span> Geological formation in Mongolia

The Nemegt Formation is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds.

<span class="mw-page-title-main">Djadochta Formation</span> Geologic formation in Mongolia

The Djadochta Formation is a highly fossiliferous geological formation situated in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 million to 71 million years ago. The type locality is the Bayn Dzak locality, famously known as the Flaming Cliffs. Reptile and mammal remains are among the fossils recovered from the formation.

<span class="mw-page-title-main">Caenagnathidae</span> Extinct family of dinosaurs

Caenagnathidae is a family of derived caenagnathoid dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that Chirostenotes pergracilis, originally named on the basis of a pair of hands, and Citipes elegans, originally thought to be an ornithomimid, named from a foot, were caenagnathids as well.

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2000.

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2001.

The Iren Dabasu Formation is a Late Cretaceous geologic formation in the Iren Nor region of Inner Mongolia. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation. The formation was first described and defined by Henry Fairfield Osborn in 1922 and it is located in the Iren Nor region of China.

<span class="mw-page-title-main">Bayan Mandahu Formation</span>

The Bayan Mandahu Formation is a geological unit of "redbeds" located near the village of Bayan Mandahu in Inner Mongolia and China and dates from the late Cretaceous Period. Laid down in the Campanian, it is dated somewhat uncertainly to between 75 and 71 mya.

<span class="mw-page-title-main">Timeline of therizinosaur research</span>

The timeline of therizinosaur research is a chronological listing of events in the history of paleontology focused on therizinosaurs. They were unusually long-necked, pot-bellied, and large-clawed herbivorous theropods most closely related to birds. The early history of therizinosaur research occurred in three phases. The first phase was the discovery of scanty and puzzling fossils in Asia by the Central Asiatic Expeditions of the 1920s and Soviet-backed research in the 1950s. This phase resulted in the discovery of the Therizinosaurus cheloniformis type specimen. Soviet paleontologist Evgeny Maleev interpreted these unusual remains as belonging to some kind of gigantic turtle.

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