Night vision is the ability to see in low-light conditions, either naturally with scotopic vision or through a night-vision device. Night vision requires both sufficient spectral range and sufficient intensity range. Humans have poor night vision compared to many animals such as cats, dogs, foxes and rabbits, in part because the human eye lacks a tapetum lucidum, [1] tissue behind the retina that reflects light back through the retina thus increasing the light available to the photoreceptors.
Night-useful spectral range techniques can sense radiation that is invisible to a human observer. Human vision is confined to a small portion of the electromagnetic spectrum called visible light. Enhanced spectral range allows the viewer to take advantage of non-visible sources of electromagnetic radiation (such as near-infrared or ultraviolet radiation). Some animals such as the mantis shrimp and trout can see using much more of the infrared and/or ultraviolet spectrum than humans. [2]
Sufficient intensity range is simply the ability to see with very small quantities of light. [3]
Many animals have better night vision than humans do, the result of one or more differences in the morphology and anatomy of their eyes. These include having a larger eyeball, a larger lens, a larger optical aperture (the pupils may expand to the physical limit of the eyelids), more rods than cones (or rods exclusively) in the retina, and a tapetum lucidum.
Enhanced intensity range is achieved via technological means through the use of an image intensifier, gain multiplication CCD, or other very low-noise and high-sensitivity arrays of photodetectors.
All photoreceptor cells in the vertebrate eye contain molecules of photoreceptor protein which is a combination of the protein photopsin in color vision cells, rhodopsin in night vision cells, and retinal (a small photoreceptor molecule). Retinal undergoes an irreversible change in shape when it absorbs light; this change causes an alteration in the shape of the protein which surrounds the retinal, and that alteration then induces the physiological process which results in vision.
The retinal must diffuse from the vision cell, out of the eye, and circulate via the blood to the liver where it is regenerated. In bright light conditions, most of the retinal is not in the photoreceptors, but is outside of the eye. It takes about 45 minutes of dark for all of the photoreceptor proteins to be recharged with active retinal, but most of the night vision adaptation occurs within the first five minutes in the dark. [4] Adaptation results in maximum sensitivity to light. In dark conditions only the rod cells have enough sensitivity to respond and to trigger vision.
Rhodopsin in the human rods is insensitive to the longer red wavelengths, so traditionally many people use red light to help preserve night vision. Red light only slowly depletes the rhodopsin stores in the rods, and instead is viewed by the red sensitive cone cells [ citation needed ].
Another theory posits that since stars typically emit light with shorter wavelengths, the light from stars will be in the blue-green color spectrum. Therefore, using red light to navigate would not desensitize the receptors used to detect star light. [6] [7]
Many animals have a tissue layer called the tapetum lucidum in the back of the eye that reflects light back through the retina, increasing the amount of light available for it to capture, but reducing the sharpness of the focus of the image. This is found in many nocturnal animals and some deep sea animals, and is the cause of eyeshine. Humans, and monkeys, lack a tapetum lucidum. [8] [9]
Nocturnal mammals have rods with unique properties that make enhanced night vision possible. The nuclear pattern of their rods changes shortly after birth to become inverted. In contrast to conventional rods, inverted rods have heterochromatin in the center of their nuclei and euchromatin and other transcription factors along the border. In addition, the outer layer of cells in the retina (the outer nuclear layer) in nocturnal mammals is thick due to the millions of rods present to process the lower light intensities. The anatomy of this layer in nocturnal mammals is such that the rod nuclei, from individual cells, are physically stacked such that light will pass through eight to ten nuclei before reaching the photoreceptor portion of the cells. Rather than being scattered, the light is passed to each nucleus individually, by a strong lensing effect due to the nuclear inversion, passing out of the stack of nuclei, and into the stack of ten photorecepting outer segments. The net effect of this anatomical change is to multiply the light sensitivity of the retina by a factor of eight to ten with no loss of focus. [10]
Night vision technologies can be broadly divided into three main categories: image intensification, active illumination, and thermal imaging.
This magnifies the amount of received photons from various natural sources such as starlight or moonlight. Examples of such technologies include night glasses and low light cameras. In the military context, Image Intensifiers are often called "Low Light TV" since the video signal is often transmitted to a display within a control center. These are usually integrated into a sensor containing both visible and IR detectors and the streams are used independently or in fused mode, depending on the mission at hand's requirements. [11]
The image intensifier is a vacuum-tube based device (photomultiplier tube) that can generate an image from a very small number of photons (such as the light from stars in the sky) so that a dimly lit scene can be viewed in real-time by the naked eye via visual output, or stored as data for later analysis. While many believe the light is "amplified," it is not. When light strikes a charged photocathode plate, electrons are emitted through a vacuum tube and strike the microchannel plate. This causes the image screen to illuminate with a picture in the same pattern as the light that strikes the photocathode and on a wavelength the human eye can see. This is much like a CRT television, but instead of color guns the photocathode does the emitting.
The image is said to become "intensified" because the output visible light is brighter than the incoming light, and this effect directly relates to the difference in passive and active night vision goggles. Currently, the most popular image intensifier is the drop-in ANVIS module, though many other models and sizes are available at the market. Recently, the US Navy announced intentions to procure a dual-color variant of the ANVIS for use in the cockpit of airborne platforms. [12]
Active illumination couples imaging intensification technology with an active source of illumination in the near infrared (NIR) or shortwave infrared (SWIR) band. Examples of such technologies include low light cameras.
Active infrared night-vision combines infrared illumination of spectral range 700–1,000 nm (just over the visible spectrum of the human eye) with CCD cameras sensitive to this light. The resulting scene, which is apparently dark to a human observer, appears as a monochrome image on a normal display device. [13] Because active infrared night-vision systems can incorporate illuminators that produce high levels of infrared light, the resulting images are typically higher resolution than other night-vision technologies. [14] [15] Active infrared night vision is now commonly found in commercial, residential and government security applications, where it enables effective night time imaging under low-light conditions. However, since active infrared light can be detected by night-vision goggles, there can be a risk of giving away position in tactical military operations.
Laser range gated imaging is another form of active night vision which utilizes a high powered pulsed light source for illumination and imaging. Range gating is a technique which controls the laser pulses in conjunction with the shutter speed of the camera's detectors. [16] Gated imaging technology can be divided into single shot, where the detector captures the image from a single light pulse, and multi-shot, where the detector integrates the light pulses from multiple shots to form an image. One of the key advantages of this technique is the ability to perform target recognition rather than mere detection, as is the case with thermal imaging.
Thermal imaging detects the temperature difference between background and foreground objects. Some organisms are able to sense a crude thermal image by means of special organs that function as bolometers. This allows thermal infrared sensing in snakes, which functions by detecting thermal radiation.
Thermal imaging cameras are excellent tools for night vision. They detect thermal radiation and do not need a source of illumination. They produce an image in the darkest of nights and can see through light fog, rain, and smoke (to a certain extent). Thermal imaging cameras make small temperature differences visible. They are widely used to complement new or existing security networks, and for night vision on aircraft, where they are commonly referred to as "FLIR" (for "forward-looking infrared"). When coupled with additional cameras (for example, a visible spectrum camera or SWIR) multispectral sensors are possible, which take advantage of the benefits of each detection band's capabilities. Contrary to misconceptions portrayed in the media, thermal imagers cannot "see" through solid objects (walls, for example), nor can they see through glass or acrylic, as both these materials have their own thermal signature and are opaque to long wave infrared radiation.
Before the introduction of image intensifiers, night glasses were the only method of night vision, and thus were widely utilized, especially at sea. Second World War era night glasses usually had a lens diameter of 56 mm or more with magnification of seven or eight. Major drawbacks of night glasses are their large size and weight.
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A night vision device (NVD) is a device comprising an image intensifier tube in a rigid casing, commonly used by military forces. Lately, night vision technology has become more widely available for civilian use. For example, enhanced vision systems (EVS) have become available for aircraft, to augment the situational awareness of pilots to prevent accidents. These systems are included in the latest avionics packages from manufacturers such as Cirrus and Cessna. The US Navy has begun procurement of a variant integrated into a helmet-mounted display, produced by Elbit Systems.
A specific type of NVD, the night vision goggle (NVG) is a night vision device with dual eyepieces. The device can utilize either one intensifier tube with the same image sent to both eyes, or a separate image intensifier tube for each eye. Night vision goggles combined with magnification lenses constitutes night vision binoculars. Other types include monocular night vision devices with only one eyepiece which may be mounted to firearms as night sights. NVG and EVS technologies are becoming more popular with helicopter operations, to improve safety. The NTSB is considering EVS as recommended equipment for safety features.
Night glasses are single or binocular with a large diameter objective. Large lenses can gather and concentrate light, thus intensifying light with purely optical means and enabling the user to see better in the dark than with the naked eye alone. Often night glasses also have a fairly large exit pupil of 7 mm or more to let all gathered light into the user's eye. However, many people cannot take advantage of this because of the limited dilation of the human pupil. To overcome this, soldiers were sometimes issued atropine eye drops to dilate pupils.[ when? ]
Currently, the PVS-14 monocular is the most widely used and preferred night vision device across NATO forces. It is used by the United States army, and is known for its low cost and wide range of uses and modification ability. Some higher end devices including the PVS-31 binocular and GPNVG-18 quad-tube night vision are used by special forces groups, but are costly. Monoculars are generally preferred by developed forces.
Night vision systems can also be installed in vehicles. An automotive night vision system is used to improve a vehicle driver's perception and seeing distance in darkness or poor weather. Such systems typically use infrared cameras, sometimes combined with active illumination techniques, to collect information that is then displayed to the driver. Such systems are currently offered as optional equipment on certain premium vehicles.
The red-eye effect in photography is the common appearance of red pupils in color photographs of the eyes of humans and several other animals. It occurs when using a photographic flash that is very close to the camera lens in ambient low light.
The retina is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then processes that image within the retina and sends nerve impulses along the optic nerve to the visual cortex to create visual perception. The retina serves a function which is in many ways analogous to that of the film or image sensor in a camera.
An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system.
The tapetum lucidum is a layer of tissue in the eye of many vertebrates and some other animals. Lying immediately behind the retina, it is a retroreflector. It reflects visible light back through the retina, increasing the light available to the photoreceptors. The tapetum lucidum contributes to the superior night vision of some animals. Many of these animals are nocturnal, especially carnivores, while others are deep sea animals.
The visual system is the physiological basis of visual perception. The system detects, transduces and interprets information concerning light within the visible range to construct an image and build a mental model of the surrounding environment. The visual system is associated with the eye and functionally divided into the optical system and the neural system.
A night-vision device (NVD), also known as a night optical/observation device (NOD) or night-vision goggle (NVG), is an optoelectronic device that allows visualization of images in low levels of light, improving the user's night vision. The device enhances ambient visible light and converts near-infrared light into visible light which can be seen by the user; this is known as I2 (image intensification). By comparison, viewing of infrared thermal radiation is referred to as thermal imaging and operates in a different section of the infrared spectrum. A night vision device usually consists of an image intensifier tube, a protective housing, and may have some type of mounting system. Many NVDs also include a protective sacrificial lens, mounted over the front lens (ie. objective lens) on NVDs to protect the latter from damage by environmental hazards, and some can incorporate telescopic lenses. The image produced by an NVD is typically monochrome green, as green was considered to be the easiest color to look at for prolonged periods in the dark. Night vision devices may be passive, relying solely on ambient light, or may be active, using an IR (infrared) illuminator to visualize the environment better.
A photoreceptor cell is a specialized type of neuroepithelial cell found in the retina that is capable of visual phototransduction. The great biological importance of photoreceptors is that they convert light into signals that can stimulate biological processes. To be more specific, photoreceptor proteins in the cell absorb photons, triggering a change in the cell's membrane potential.
Rod cells are photoreceptor cells in the retina of the eye that can function in lower light better than the other type of visual photoreceptor, cone cells. Rods are usually found concentrated at the outer edges of the retina and are used in peripheral vision. On average, there are approximately 92 million rod cells in the human retina. Rod cells are more sensitive than cone cells and are almost entirely responsible for night vision. However, rods have little role in color vision, which is the main reason why colors are much less apparent in dim light.
The choroid, also known as the choroidea or choroid coat, is a part of the uvea, the vascular layer of the eye. It contains connective tissues, and lies between the retina and the sclera. The human choroid is thickest at the far extreme rear of the eye, while in the outlying areas it narrows to 0.1 mm. The choroid provides oxygen and nourishment to the outer layers of the retina. Along with the ciliary body and iris, the choroid forms the uveal tract.
In visual physiology, adaptation is the ability of the retina of the eye to adjust to various levels of light. Natural night vision, or scotopic vision, is the ability to see under low-light conditions. In humans, rod cells are exclusively responsible for night vision as cone cells are only able to function at higher illumination levels. Night vision is of lower quality than day vision because it is limited in resolution and colors cannot be discerned; only shades of gray are seen. In order for humans to transition from day to night vision they must undergo a dark adaptation period of up to two hours in which each eye adjusts from a high to a low luminescence "setting", increasing sensitivity hugely, by many orders of magnitude. This adaptation period is different between rod and cone cells and results from the regeneration of photopigments to increase retinal sensitivity. Light adaptation, in contrast, works very quickly, within seconds.
Scanning laser ophthalmoscopy (SLO) is a method of examination of the eye. It uses the technique of confocal laser scanning microscopy for diagnostic imaging of the retina or cornea of the human eye.
Melanopsin is a type of photopigment belonging to a larger family of light-sensitive retinal proteins called opsins and encoded by the gene Opn4. In the mammalian retina, there are two additional categories of opsins, both involved in the formation of visual images: rhodopsin and photopsin in the rod and cone photoreceptor cells, respectively.
In the study of visual perception, scotopic vision is the vision of the eye under low-light conditions. The term comes from Greek skotos, meaning "darkness", and -opia, meaning "a condition of sight". In the human eye, cone cells are nonfunctional in low visible light. Scotopic vision is produced exclusively through rod cells, which are most sensitive to wavelengths of around 498 nm (blue-green) and are insensitive to wavelengths longer than about 640 nm (red-orange). This condition is called the Purkinje effect.
Intrinsically photosensitive retinal ganglion cells (ipRGCs), also called photosensitive retinal ganglion cells (pRGC), or melanopsin-containing retinal ganglion cells (mRGCs), are a type of neuron in the retina of the mammalian eye. The presence of ipRGCs was first suspected in 1927 when rodless, coneless mice still responded to a light stimulus through pupil constriction, This implied that rods and cones are not the only light-sensitive neurons in the retina. Yet research on these cells did not advance until the 1980s. Recent research has shown that these retinal ganglion cells, unlike other retinal ganglion cells, are intrinsically photosensitive due to the presence of melanopsin, a light-sensitive protein. Therefore, they constitute a third class of photoreceptors, in addition to rod and cone cells.
A retinal implant is a visual prosthesis for restoration of sight to patients blinded by retinal degeneration. The system is meant to partially restore useful vision to those who have lost their photoreceptors due to retinal diseases such as retinitis pigmentosa (RP) or age-related macular degeneration (AMD). Retinal implants are being developed by a number of private companies and research institutions, and three types are in clinical trials: epiretinal, subretinal, and suprachoroidal. The implants introduce visual information into the retina by electrically stimulating the surviving retinal neurons. So far, elicited percepts had rather low resolution, and may be suitable for light perception and recognition of simple objects.
The pigmented layer of retina or retinal pigment epithelium (RPE) is the pigmented cell layer just outside the neurosensory retina that nourishes retinal visual cells, and is firmly attached to the underlying choroid and overlying retinal visual cells.
A visual prosthesis, often referred to as a bionic eye, is an experimental visual device intended to restore functional vision in those with partial or total blindness. Many devices have been developed, usually modeled on the cochlear implant or bionic ear devices, a type of neural prosthesis in use since the mid-1980s. The idea of using electrical current to provide sight dates back to the 18th century, discussed by Benjamin Franklin, Tiberius Cavallo, and Charles LeRoy.
Mammals normally have a pair of eyes. Although mammalian vision is not so excellent as bird vision, it is at least dichromatic for most of mammalian species, with certain families possessing a trichromatic color perception.
Vision is an important sensory system for most species of fish. Fish eyes are similar to the eyes of terrestrial vertebrates like birds and mammals, but have a more spherical lens. Birds and mammals normally adjust focus by changing the shape of their lens, but fish normally adjust focus by moving the lens closer to or further from the retina. Fish retinas generally have both rod cells and cone cells, and most species have colour vision. Some fish can see ultraviolet and some are sensitive to polarised light.
Retinal regeneration refers to the restoration of vision in vertebrates that have suffered retinal lesions or retinal degeneration.
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