| Butternut | |
|---|---|
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| A mature butternut tree | |
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Clade: | Angiosperms |
| Clade: | Eudicots |
| Clade: | Rosids |
| Order: | Fagales |
| Family: | Juglandaceae |
| Genus: | Juglans |
| Section: | Juglans sect. Trachycaryon |
| Species: | J. cinerea |
| Binomial name | |
| Juglans cinerea L. 1759 | |
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| Natural range | |
| Synonyms [3] | |
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Juglans cinerea, commonly known as butternut or white walnut, [4] is a species of walnut native to the eastern United States and southeast Canada.
The distribution range of J. cinerea extends east to New Brunswick, and from southern Quebec west to Minnesota, south to northern Alabama and southwest to northern Arkansas. [5] It is absent from most of the Southern United States. [6] The species also proliferates at middle elevations (about 2,000 ft or 610 m above sea level) in the Columbia River basin, Pacific Northwest; as an off-site species. Trees with 7 ft or 2.1 m (over mature) class range diameter at breast height were noted in the Imnaha River drainage as late as January 26, 2015.[ citation needed ] Butternut favors a cooler climate than black walnut and its range does not extend into the Deep South. Its northern range extends into Wisconsin and Minnesota where the growing season is too short for black walnut.
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J. cinerea is a deciduous tree growing to 20 m (66 ft) tall, rarely 40 m (130 ft). Butternut is a slow-growing species, and rarely lives longer than 75 years. It has a 40–80 cm (16–31 in) stem diameter, with light gray bark.
The leaves are alternate and pinnate, 40–70 cm (16–28 in) long, with 11–17 leaflets, each leaflet 5–10 cm (2–4 in) long and 3–5 cm (1+1⁄4–2 in) broad. Leaves have a terminal leaflet at the end of the leafstalk and have an odd number of leaflets. The whole leaf is downy-pubescent, and a somewhat brighter, yellower green than many other tree leaves.
Like other members of the family Juglandaceae, butternut's leafout in spring is tied to photoperiod rather than air temperature and occurs when daylight length reaches 14 hours. This can vary by up to a month in the northern and southernmost extents of its range. Leaf drop in fall is early and is initiated when daylight drops to 11 hours. The species is monoecious. Male (staminate) flowers are inconspicuous, yellow-green slender catkins that develop from axillary buds and female (pistillate) flowers are short terminal spikes on current year's shoots. Each female flower has a light pink stigma. Flowers of both sexes do not usually mature simultaneously on any individual tree.
The fruit is a lemon-shaped nut, produced in bunches of two to six together; the nut is oblong-ovoid, 3–6 cm (1+1⁄4–2+1⁄4 in) long and 2–4 cm (3⁄4–1+1⁄2 in) broad, surrounded by a green husk before maturity in midautumn.
Butternut grows best on stream banks and on well-drained soils. It is seldom found on dry, compact, or infertile soils. It grows better than black walnut, however, on dry, rocky soils, especially those of limestone origin. Butternut's range includes the rocky soils of New England where black walnut is largely absent.
Butternut is found most frequently in coves, on stream benches and terraces, on slopes, in the talus of rock ledges, and on other sites with good drainage. It is found up to an elevation of 1,500 metres (4,900 ft) in the Virginias – much higher altitudes than black walnut. The nuts are eaten by wildlife. [7]
Butternut is found with many other tree species in several hardwood types in the mixed mesophytic forest. It is an associated species in the following four northern and central forest cover types: sugar maple–basswood, yellow poplar–white oak–northern red oak, beech–sugar maple, and river birch–sycamore. Commonly associated trees include basswood (Tilia spp.), black cherry (Prunus serotina), beech (Fagus grandifolia), black walnut (Juglans nigra), elm (Ulmus spp.), hemlock (Tsuga canadensis), hickory (Carya spp.), oak (Quercus spp.), red maple (Acer rubrum), sugar maple (Acer saccharum), yellow poplar (Liriodendron tulipifera), white ash (Fraxinus americana), and yellow birch (Betula alleghaniensis). In the northeast part of its range, it is often found with sweet birch (Betula lenta) and in the northern part of its range it is occasionally found with white pine (Pinus strobus). Forest stands seldom contain more than an occasional butternut tree, although in local areas, it may be abundant. In the past, West Virginia, Wisconsin, Indiana, and Tennessee have been the leading producers of butternut timber.[ citation needed ]
Although young trees may withstand competition from the side, butternut does not survive under shade from above. It must be in the overstory to thrive. Therefore, it is classed as intolerant of shade and competition.[ citation needed ]
The most serious disease of J. cinerea is butternut decline or butternut canker. [8] In the past, the causal organism of this disease was thought to be a fungus, Melanconis juglandis. Now this fungus has been associated with secondary infections and the primary causal organism of the disease has been identified as another species of fungus, Ophiognomonia clavigignenti-juglandacearum . The fungus is spread by wide-ranging vectors,[ citation needed ] so isolation of a tree offers no protection.
Butternut canker first entered the United States around the beginning of the 20th century, when it arrived on imported nursery stock of Japanese walnut.
Symptoms of the disease include dying branches and stems. Initially, cankers develop on branches in the lower crown. Spores developing on these dying branches are spread by rainwater to tree stems. Stem cankers develop 1 to 3 years after branches die. Tree tops killed by stem-girdling cankers do not resprout. Diseased trees usually die within several years. Completely free-standing trees seem better able to withstand the fungus than those growing in dense stands or forest. In some areas, 90% of the butternut trees have been killed. The disease is reported to be spreading rapidly in Wisconsin. By contrast, black walnut seems to be resistant to the disease.
Butternut hybridizes readily with Japanese walnut. The hybrid between butternut and the Japanese walnut is commonly known as the 'buartnut' and inherits Japanese walnut's resistance to the disease. Researchers are back-crossing butternut to buartnut, creating 'butter-buarts" which should have more butternut traits than buartnuts. They are selecting for resistance to the disease. Most butternuts found as landscaping trees are buartnuts rather than the pure species.[ citation needed ]
Bunch disease also attacks butternut. Currently, the causal agent is thought to be a mycoplasma-like organism. Symptoms include a yellow witches' broom resulting from sprouting and growth of auxiliary buds that would normally remain dormant. Infected branches fail to become dormant in the fall and are killed by frost; highly susceptible trees may eventually be killed. Butternut seems to be more susceptible to this disease than black walnut.
The common grackle has been reported to destroy immature fruit and may be considered a butternut pest [9] when populations are high.
Butternut is very susceptible to fire damage, and although the species is generally wind firm, it is subject to frequent storm damage. [9]
The species is not listed as threatened federally in the US, but is listed as "Special Concern" in Kentucky, "Exploitably Vulnerable" in New York State, and "Threatened" in Tennessee. [10] This species is also listed as "Endangered" in Illinois. [11]
The Committee on the Status of Endangered Wildlife in Canada placed the butternut on the endangered species list in Canada in 2005. [12]
Approximately 60 grafted butternut trees were planted in a seed orchard in Huntingburg, Indiana, in 2012 as part of a larger effort by the USDA Forest Service to conserve the species and to breed resistance to butternut canker disease. Forest Service staff from the Hoosier National Forest, the Eastern Region National Forest genetics program, the Northern Research Station, and the Hardwood Tree Improvement and Regeneration Center at Purdue University are involved in the project. [13]
The American Forest National Champion is located in Oneida, New York. In 2016 its circumference at breast height was 288 in (7,300 mm), the height was 67 ft (20 m), and the spread was 88 ft (27 m). [14]
The Bush butternut tree was planted by settler George Bush (1845) in current Tumwater, Washington, brought from Missouri. It was seriously damaged in a windstorm in 2015, and collapsed on May 1, 2021, at the age of 176 years. [15]
Butternut wood has been found at the pre-Columbian Viking settlement at L'Anse aux Meadows at the northernmost tip of Newfoundland, an island where butternut does not grow and to which it therefore is presumed to have been transported from more southerly locations. [16] The butternut log specimens found there were cut with European tools. [17]
| Nutritional value per 100 g (3.5 oz) | |||||||||||||||||||||||||||||||||||||||||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Energy | 2,560 kJ (610 kcal) | ||||||||||||||||||||||||||||||||||||||||||||||||
12 g | |||||||||||||||||||||||||||||||||||||||||||||||||
| Dietary fiber | 4.7 g | ||||||||||||||||||||||||||||||||||||||||||||||||
57 g | |||||||||||||||||||||||||||||||||||||||||||||||||
| Saturated | 1.31 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Monounsaturated | 10.4 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Polyunsaturated | 42.7 g 8.72 g 33.7 g | ||||||||||||||||||||||||||||||||||||||||||||||||
24.9 g | |||||||||||||||||||||||||||||||||||||||||||||||||
| Tryptophan | 0.366 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Threonine | 0.94 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Isoleucine | 1.18 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Leucine | 2.2 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Lysine | 0.77 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Methionine | 0.611 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Cystine | 0.484 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Phenylalanine | 1.44 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Tyrosine | 0.977 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Valine | 1.54 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Arginine | 4.86 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Histidine | 0.808 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Alanine | 1.37 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Aspartic acid | 3.1 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Glutamic acid | 6.08 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Glycine | 1.51 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Proline | 1.24 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| Serine | 1.64 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| |||||||||||||||||||||||||||||||||||||||||||||||||
| Other constituents | Quantity | ||||||||||||||||||||||||||||||||||||||||||||||||
| Water | 3.34 g | ||||||||||||||||||||||||||||||||||||||||||||||||
| †Percentages estimated using US recommendations for adults, [18] except for potassium, which is estimated based on expert recommendation from the National Academies. [19] | |||||||||||||||||||||||||||||||||||||||||||||||||
The butternuts are edible [20] and were made into a butter-like oil by Native Americans [21] for various purposes. The young green nuts, while still soft, can be pickled; survivalist Bradford Angier recommends this be done with a change of salt water every other day for a week, and a subsequent seasoning of at least two weeks. [21] The sap can be used to make syrup. [21]
Butternut wood is light in weight and takes polish well, and is highly rot resistant, but is much softer than black walnut wood. Oiled, the grain of the wood usually shows much light. It is often used to make furniture, and is a favorite of woodcarvers.
Butternut bark and nut rinds were once often used to dye cloth to colors between light yellow [4] and dark brown. [22] The husks contain a natural yellow-orange dye. [7] To produce the darker colors, the bark is boiled to concentrate the color. This appears to never have been used as a commercial dye, but rather was used to color homespun cloth.
In the mid-19th century, inhabitants of areas such as southern Illinois and southern Indiana – many of whom had moved there from the Southern United States – were known as "butternuts" from the butternut-dyed homespun cloth that some of them wore. Later, during the American Civil War, the term "butternut" was sometimes applied to Confederate soldiers. Some Confederate uniforms apparently faded from gray to a tan or light brown. It is also possible that butternut was used to color the cloth worn by a small number of Confederate soldiers. [22] The resemblance of these uniforms to butternut-dyed clothing, and the association of butternut dye with home-made clothing, resulted in the derisive nickname.
Crushed fruits can be used to poison fish, though the practice is illegal in most jurisdictions. Bruised fruit husks of the closely related black walnut can be used to stun fish. [23]
Quercus muehlenbergii, the chinquapinoak, is a deciduous species of tree in the white oak group. The species was often called Quercus acuminata in older literature. Quercus muehlenbergii is native to eastern and central North America. It ranges from Vermont to Minnesota, south to the Florida panhandle, and west to New Mexico in the United States. In Canada it is only found in southern Ontario, and in Mexico it ranges from Coahuila south to Hidalgo.
Walnut trees are any species of tree in the plant genus Juglans, the type genus of the family Juglandaceae, the seeds of which are referred to as walnuts. All species are deciduous trees, 10–40 metres (33–131 ft) tall, with pinnate leaves 200–900 millimetres (7.9–35.4 in), with 5–25 leaflets; the shoots have chambered pith, a character shared with the wingnuts (Pterocarya), but not the hickories (Carya) in the same family.
Quercus velutina, the black oak, is a species of oak in the red oak group, native and widespread in eastern and central North America. It is sometimes called the eastern black oak.
The pecan is a species of hickory native to the southern United States and northern Mexico in the region of the Mississippi River.
Juglans nigra, the eastern American black walnut, is a species of deciduous tree in the walnut family, Juglandaceae, native to North America. It grows mostly in riparian zones, from southern Ontario, west to southeast South Dakota, south to Georgia, northern Florida and southwest to central Texas. Wild trees in the upper Ottawa Valley may be an isolated native population or may have derived from planted trees.
A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticulture. Their causes include a wide range of organisms as fungi, bacteria, mycoplasmas and viruses. The majority of canker-causing organisms are bound to a unique host species or genus, but a few will attack other plants. Weather and animal damage can also cause stress to the plant resulting in cankers. Other causes of cankers is pruning when the bark is wet or using un-sterilized tools.
Juglans regia, the Persian walnut, English walnut, Carpathian walnut, Madeira walnut, or, especially in Great Britain, common walnut, is an Old World walnut tree species native to the region stretching from the Caucasus eastward to the Kashmir region. It is widely cultivated across Asia, Northern America and Europe.
Juglans ailantifolia, the Japanese walnut, is a species of walnut native to Japan and Sakhalin. It is a deciduous tree growing to 20 m (66 ft) tall, rarely 30 m (98 ft), and 40–80 cm (16–31 in) stem diameter, with light grey bark. The leaves are pinnate, 50–90 cm (20–35 in) long, with 11–17 leaflets, each leaflet 7–16 cm (2.8–6.3 in) long and 3–5 cm (1.2–2.0 in) broad. The whole leaf is downy-pubescent, and a somewhat brighter, yellower green than many other tree leaves. The male flowers are inconspicuous yellow-green catkins produced in spring at the same time as the new leaves appear. The female flowers have pink/red pistils. The fruit is a nut, produced in bunches of 4–10 together; the nut is spherical, 3–5 cm long and broad, surrounded by a green husk before maturity in mid autumn.
Juglone, also called 5-hydroxy-1,4-naphthalenedione (IUPAC) is a phenolic organic compound with the molecular formula C10H6O3. In the food industry, juglone is also known as C.I. Natural Brown 7 and C.I. 75500. It is insoluble in benzene but soluble in dioxane, from which it crystallizes as yellow needles. It is an isomer of lawsone, which is the active dye compound in the henna leaf.
Castanea pumila, commonly known as the Allegheny chinquapin, American chinquapin or dwarf chestnut, is a species of chestnut native to the southeastern United States. The native range is from Massachusetts and New York to Maryland and extreme southern New Jersey and southeast Pennsylvania south to central Florida, west to eastern Texas, and north to southern Missouri and Kentucky. The plant's habitat is dry sandy and rocky uplands and ridges mixed with oak and hickory to 1000 m elevation. It grows best on well-drained soils in full sun or partial shade.
Juglans californica, the California black walnut, also called the California walnut, or the Southern California black walnut, is a large shrub or small tree of the walnut family, Juglandaceae, endemic to the Central Valley and the Coast Range valleys from Northern to Southern California.
Ophiognomonia clavigignenti-juglandacearum is a mitosporic fungus that causes the butternut canker, a lethal disease of butternut trees. It is also known to parasitize other members of the genus Juglans on occasion, and very rarely other related trees including hickories. The fungus is found throughout North America, occurring on up to 91% of butternut trees, and may be threatening the viability of butternut as a species.
Carya glabra, the pignut hickory, is a common, but not abundant species of hickory in the oak-hickory forest association in the Eastern United States and Canada. Other common names are pignut, sweet pignut, coast pignut hickory, smoothbark hickory, swamp hickory, and broom hickory. The pear-shaped nut ripens in September and October, has a sweet maple like smell, and is an important part of the diet of many wild animals. The wood is used for a variety of products, including fuel for home heating. Its leaves turn yellow in the Fall.
Juglans neotropica is a species of plant in the Juglandaceae family. It is found in Colombia, Ecuador, and Peru. It is threatened by habitat loss. Common names include Colombian walnut, Ecuadorian walnut, Andean walnut, nogal, cedro negro, cedro nogal, and nogal Bogotano.
Juglans mandshurica, also known as monkey nuts, or tigernut, is a species of deciduous tree in the walnut family Juglandacea, native to the Eastern Asiatic Region. It grows to about 25 m.
Juglans hindsii, commonly called the Northern California black walnut and Hinds's black walnut, is a species of walnut tree native to the western United States. It is commonly called claro walnut by the lumber industry and woodworkers, and is the subject of some confusion over its being the root stock for English walnut orchard stock.
Thousand cankers disease (TCD) is a recently recognized disease of certain walnuts. The disease results from the combined activity of the walnut twig beetle and a canker producing fungus, Geosmithia morbida. Until July 2010 the disease was only known to the western United States where over the past decade it has been involved in several large scale die-offs of walnut, particularly black walnut, Juglans nigra. However, in late July 2010 a well-established outbreak of the disease was found in the Knoxville, Tennessee, area. This new finding is the first locating it within the native range of its susceptible host, black walnut. In 2013, an outbreak was found in the Veneto region of Italy, where the disease has been found on both black walnut and English walnut.
A walnut is the edible seed of any tree of the genus Juglans, particularly the Persian or English walnut, Juglans regia. They are accessory fruit because the outer covering of the fruit is technically an involucre and thus not morphologically part of the carpel; this means it cannot be a drupe but is instead a drupe-like nut.
Husk in botany is the outer shell or coating of a seed. In the United States, the term husk often refers to the leafy outer covering of an ear of maize (corn) as it grows on the plant. Literally, a husk or hull includes the protective outer covering of a seed, fruit, or vegetable.
