Juglandaceae

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Juglandaceae
Juglans regia Broadview.jpg
Juglans regia
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Fagales
Family: Juglandaceae
Augustin Pyramus Perleb [1] fermium
Type genus
Juglans
L.
Subfamilies

See text

Engelhardioideae Distribution.svg
The range of subfamily Engelhardioideae
Juglandoideae Distribution.svg
The range of subfamily Juglandoideae
Synonyms [2]
  • Platycaryaceae Nakai ex Doweld
  • Pterocaryaceae Nakai, nom. inval.
  • Rhoipteleaceae Hand.-Mazz. 1932, nom. cons.

The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia.

Contents

The nine or ten genera in the family have a total of around 50 species, [3] and include the commercially important nut-producing trees walnut (Juglans), pecan (Carya illinoinensis), and hickory (Carya). The Persian walnut, Juglans regia , is one of the major nut crops of the world. Walnut, hickory, and gaulin are also valuable timber trees while pecan wood is also valued as cooking fuel.

Description

Members of the walnut family have large, aromatic leaves that are usually alternate, but opposite in Alfaroa and Oreomunnea . The leaves are pinnately compound or ternate, and usually 20–100 cm long. The trees are wind-pollinated, and the flowers are usually arranged in catkins.

The fruits of the Juglandaceae are often confused with drupes but are accessory fruit because the outer covering of the fruit is technically an involucre and thus not morphologically part of the carpel; this means it cannot be a drupe but is instead a drupe-like nut. [4]

Taxonomy

The known living genera are grouped into subfamilies, tribes, and subtribes as follows: [5]

Systematics

Modern molecular phylogenetics suggest the following relationships: [7]

Myricaceae  (outgroup)

Juglandaceae
Rhoipteleoideae

Rhoiptelea

Engelhardioideae

Engelhardia

Alfaroa

Oreomunnea

Juglandoideae

Carya

Annamocarya

Platycarya

Cyclocarya

Pterocarya

Juglans

Related Research Articles

<span class="mw-page-title-main">Fagales</span> Order of flowering plants

The Fagales are an order of flowering plants, including some of the best-known trees. The order name is derived from genus Fagus, beeches. They belong among the rosid group of dicotyledons. The families and genera currently included are as follows:

<span class="mw-page-title-main">Drupe</span> Fleshy fruit with hard inner layer (endocarp or stone) surrounding the seed

In botany, a drupe is an indehiscent type of fruit in which an outer fleshy part surrounds a single shell of hardened endocarp with a seed (kernel) inside. These fruits usually develop from a single carpel, and mostly from flowers with superior ovaries.

<span class="mw-page-title-main">Scrophulariaceae</span> Figwort family of flowering plants

The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral (zygomorphic) or rarely radial (actinomorphic) symmetry. The Scrophulariaceae have a cosmopolitan distribution, with the majority found in temperate areas, including tropical mountains. The family name is based on the name of the included genus Scrophularia L.

<i>Juglans</i> Genus of trees

Walnut trees are any species of tree in the plant genus Juglans, the type genus of the family Juglandaceae, the seeds of which are referred to as walnuts. All species are deciduous trees, 10–40 metres (33–131 ft) tall, with pinnate leaves 200–900 millimetres (7.9–35.4 in), with 5–25 leaflets; the shoots have chambered pith, a character shared with the wingnuts (Pterocarya), but not the hickories (Carya) in the same family.

<span class="mw-page-title-main">Betulaceae</span> Family of flowering plants comprising hazel and birch trees

Betulaceae, the birch family, includes six genera of deciduous nut-bearing trees and shrubs, including the birches, alders, hazels, hornbeams, hazel-hornbeam, and hop-hornbeams numbering a total of 167 species. They are mostly natives of the temperate Northern Hemisphere, with a few species reaching the Southern Hemisphere in the Andes in South America. Their typical flowers are catkins and often appear before leaves.

<span class="mw-page-title-main">Pecan</span> Species of hickory native to the southern USA and northern Mexico

The pecan is a species of hickory native to the southern United States and northern Mexico in the region of the Mississippi River.

<span class="mw-page-title-main">Nut (fruit)</span> In botany, type of dry indehiscent fruit

A nut is a fruit consisting of a hard or tough nutshell protecting a kernel which is usually edible. In general usage and in a culinary sense, a wide variety of dry seeds are called nuts, but in a botanical context "nut" implies that the shell does not open to release the seed (indehiscent).

<span class="mw-page-title-main">Hickory</span> Genus of trees

Hickory is a common name for trees composing the genus Carya, which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in Mexico, and two to four are native to Canada. A number of hickory species are used for their edible nuts, lumber or other wood and woodcraft products.

<i>Pterocarya</i> Genus of trees

Pterocarya, often called wingnuts in English, are trees in the walnut family Juglandaceae. They are native to Asia. The botanic name is from Ancient Greek πτερον (pteron) "wing" + κάρυον (karyon) "nut".

<i>Carya ovata</i> Species of tree

Carya ovata, the shagbark hickory, is a common hickory in the Eastern United States and southeast Canada. It is a large, deciduous tree, growing well over 100 ft (30 m) tall, and can live more than 350 years. The tallest measured shagbark, located in Savage Gulf, Tennessee, is over 150 ft (46 m) tall. Mature shagbarks are easy to recognize because, as their name implies, they have shaggy bark. This characteristic is, however, only found on mature trees; young specimens have smooth bark.

<span class="mw-page-title-main">Juglandeae</span> Tribe of flowering plants

Juglandeae is a tribe of the Juglandoideae subfamily, in the Juglandaceae family.

Carya sinensis is a species of tree native to southwestern China and northern Vietnam, in the hickory genus Carya. It is sometimes called Chinese hickory or beaked hickory. It is closely related to Carya kweichowensis.

<i>Alfaroa costaricensis</i> Species of tree

Alfaroa costaricensis, also known as campano chile, chiciscua, gaulin, gavilán Colorado, or gavilancillo, is nut bearing timber tree in the Juglandaceae family. It is native to the Neotropics, from Mexico, through Central America to Colombia.

Alfaroa manningii, or gavilán colorado, is a valued lumber tree of the Walnut family endemic to the premontane Costa Rican rain forest. The specific epithet honors the American botanist Wayne Eyer Manning (1899–2004).

<span class="mw-page-title-main">Juglandoideae</span> Subfamily of flowering plants

Juglandoideae is a subfamily of the walnut family Juglandaceae.

<i>Curculio caryae</i> Species of beetle

The pecan weevil, Curculio caryae is an obligate feeder on the nuts of North American hickories and pecans, most widely recognized as an economically important pest of the pecan, Carya illinoinensis. It has also been observed to infest one Juglans species, the Persian walnut, Juglans regia.

<span class="mw-page-title-main">Juglandinae</span> Subtribe of flowering plants

Juglandinae is a subtribe of the Juglandeae tribe, of the Juglandoideae subfamily, in the Juglandaceae family. Walnut tree species make up the genus Juglans, which belongs to the subtribe Juglandinae.

<i>Carya pallida</i> Species of flowering plant

Carya pallida, sand hickory, or pale hickory is a species of hickory native to the southeastern United States. It is a perennial, dicotyledonous plant which prefers rocky or sandy habitats. The sand hickory can reach heights of up to 30m, but its typical height is between 9-24m. In an open area, Carya crowns are usually towering and slim. The sand hickory nut is edible and consumed by various organisms.

References

  1. Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III" (PDF). Botanical Journal of the Linnean Society. 161 (2): 105–121. doi: 10.1111/j.1095-8339.2009.00996.x . Retrieved 2013-07-06.
  2. "Family: Juglandaceae DC. ex Perleb, nom. cons". Germplasm Resources Information Network. United States Department of Agriculture. 2003-01-17. Archived from the original on 2015-10-03. Retrieved 2011-11-17.
  3. Christenhusz, M. J. M.; Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. 261 (3): 201–217. doi: 10.11646/phytotaxa.261.3.1 .
  4. John Derek Bewley, Michael Black, Peter Halmer (2006) The Encyclopedia of Seeds: Science, Technology And Uses
  5. Manos, P. S.; D. E. Stone (2001). "Evolution, phylogeny and systematics of the Juglandaceae" (PDF). Annals of the Missouri Botanical Garden. 88 (2): 231–269. doi:10.2307/2666226. JSTOR   2666226.
  6. "GRIN Genera of Juglandaceae". Germplasm Resources Information Network. United States Department of Agriculture. Archived from the original on 2015-09-24. Retrieved 2011-11-17.
  7. Xiang XG, Wang W, Li RQ, Lin L, Liu Y, Zhou ZK, Li ZY, Chen ZD (2014). "Large-scale phylogenetic analyses reveal fagalean diversification promoted by the interplay of diaspores and environments in the Paleogene". Perspectives in Plant Ecology, Evolution and Systematics. 16 (3): 101–110. doi:10.1016/j.ppees.2014.03.001.