The Malpighiales comprise one of the largest orders of flowering plants, containing about 36 families and more than 16,000 species, about 7.8% of the eudicots. The order is very diverse, containing plants as different as the willow, violet, poinsettia, manchineel, rafflesia and coca plant, and are hard to recognize except with molecular phylogenetic evidence. It is not part of any of the classification systems based only on plant morphology. Molecular clock calculations estimate the origin of stem group Malpighiales at around 100 million years ago (Mya) and the origin of crown group Malpighiales at about 90 Mya.
Nymphaeaceae is a family of flowering plants, commonly called water lilies. They live as rhizomatous aquatic herbs in temperate and tropical climates around the world. The family contains five genera with about 70 known species. Water lilies are rooted in soil in bodies of water, with leaves and flowers floating on or emergent from the surface. Leaves are round, with a radial notch in Nymphaea and Nuphar, but fully circular in Victoria and Euryale. It is the national flower of Bangladesh.
Ceratophyllaceae is a cosmopolitan family of flowering plants including one living genus commonly found in ponds, marshes, and quiet streams in tropical and in temperate regions. It is the only extant family in the order Ceratophyllales. Species are commonly called coontails or hornworts, although hornwort is also used for unrelated plants of the division Anthocerotophyta.
The Clusiaceae or GuttiferaeJuss. (1789) are a family of plants including 13 genera and ca 750 species. Several former members of Clusiacae are now placed in Calophyllaceae and Hypericaceae. They are mostly trees and shrubs, with milky sap and fruits or capsules for seeds. The family is primarily tropical. More so than many plant families, it shows large variation in plant morphology. According to the APG III, this family belongs to the order Malpighiales.
Burmanniaceae is a family of flowering plants, consisting of 99 species of herbaceous plants in eight genera.
Hypericaceae is a plant family in the order Malpighiales, comprising six to nine genera and up to 700 species, and commonly known as the St. John’s wort family. Members are found throughout the world apart from extremely cold or dry habitats. Hypericum and Triadenum occur in temperate regions but other genera are mostly tropical.
Asphodelaceae is a family of flowering plants in the order Asparagales. Such a family has been recognized by most taxonomists, but the circumscription has varied widely. In its current circumscription in the APG IV system, it includes about 40 genera and 900 known species. The type genus is Asphodelus.
The family Pandaceae consists of three genera that were formerly recognized in the Euphorbiaceae. Those are:
Achariaceae is a family of flowering plants consisting of 32-33 genera with about 155 species of tropical herbs, shrubs, and trees. The APG IV system has greatly expanded the scope of the family by including many genera previously classified in Flacourtiaceae. Molecular data strongly support the inclusion of this family in the order Malpighiales.
Malpighiaceae is a family of flowering plants in the order Malpighiales. It comprises about 73 genera and 1315 species, all of which are native to the tropics and subtropics. About 80% of the genera and 90% of the species occur in the New World and the rest in the Old World.
Chrysobalanaceae is a family of flowering plants, consisting of trees and shrubs in 27 genera and about 700 species of pantropical distribution with a centre of diversity in the Amazon. Some of the species contain silica in their bodies for rigidity and so the mesophyll often has sclerenchymatous idioblasts. The widespread species Chrysobalanus icaco produces a plum-like fruit and the plant is commonly known as the coco plum.
Peridiscaceae is a family of flowering plants in the order Saxifragales. Four genera comprise this family: Medusandra, Soyauxia, Peridiscus, and Whittonia., with a total of 12 known species. It has a disjunct distribution, with Peridiscus occurring in Venezuela and northern Brazil, Whittonia in Guyana, Medusandra in Cameroon, and Soyauxia in tropical West Africa. Whittonia is possibly extinct, being known from only one specimen collected below Kaieteur Falls in Guyana. In 2006, archeologists attempted to rediscover it, however, it proved unsuccessful.
Euptelea is a genus of two species of flowering plants in the monogeneric family Eupteleaceae. The genus is found from Assam east through China to Japan, and consists of shrubs or small trees:
Myrothamnus is a genus of flowering plants, consisting of two species of small xerophytic shrubs, in the southern parts of tropical Africa and in Madagascar. Myrothamnus is recognized as the only genus in the family Myrothamnaceae.
Boryaceae is a family of highly drought-tolerant flowering plants native to Australia, placed in the order Asparagales of the monocots. The family includes two genera, with twelve species in total in Australia.
Schoepfiaceae is a family of flowering plants recognized in the APG III system of 2009. The family was previously only recognized by few taxonomists; the plants in question usually being assigned to family Olacaceae and Santalaceae.
Hydrostachys is a genus of about 22 species of flowering plants native to Madagascar and southern and central Africa. It is the only genus in the family Hydrostachyaceae. All species of Hydrostachys are aquatic, growing on rocks in fast-moving water. They have tuberous roots, usually pinnately compound leaves, and highly reduced flowers on dense spikes.
Huaceae is a family of plant in the rosids group, which has been classed in the orders Malpighiales, Malvales, and Violales or in its own order Huales. The APG II system placed it in the clade eurosids I, whereas the APG III system of 2009 and APG IV (2016) place it within the Oxalidales. The family is endemic to central Africa. It contains four species in the following two genera:
Calophyllaceae is a family of flowering plants in the order Malpighiales and is recognized by the APG III system of classification. Most of the 14 genera and 475 species included in this family were previously recognized in the tribe Calophylleae of the family Clusiaceae. The Angiosperm Phylogeny Group determined that splitting this clade of genera off into their own family was necessary.
Centroplacaceae is a family of flowering plants in the order Malpighiales and is recognized by the APG III system of classification. The family comprises two genera: Bhesa, which was formerly recognized in the Celastraceae, and Centroplacus, which was formerly recognized in the Euphorbiaceae, together comprising six species. The Angiosperm Phylogeny Group determined that based on previous phylogenetic analysis, these two genera formed an isolated clade and recognition of the family was "reasonable."
