Tofieldiaceae | |
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2 species of Tofieldia from: Thomé (1885) | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Order: | Alismatales |
Family: | Tofieldiaceae Takht. [1] |
Tofieldiaceae is a family of flowering plants in the monocot order Alismatales. [2] The family is divided into four genera, which together comprise 28 known species (Christenhusz & Byng 2016 [3] ). They are small, herbaceous plants, mostly of arctic and subarctic regions, but a few extend further south, and one genus is endemic to northern South America and Florida. Tofieldia pusilla is sometimes grown as an ornamental. [4]
William Hudson (1730-1793) named Tofieldia for the British botanist Thomas Tofield (1730–1779). [5] The family Tofieldiaceae was erected by Armen Takhtajan in 1995. [6] Molecular phylogenetic studies of DNA sequences have shown it to be the second diverging clade in Alismatales, after the most basal clade, the family Araceae. [7] Before the segregation of Tofieldiaceae was confirmed by cladistic methods, its genera had usually been assigned to Nartheciaceae, [8] Liliaceae, [9] or Melanthiaceae. Tofieldiaceae is recognized by the Angiosperm Phylogeny Group in their APG III system of plant classification.
The circumscription of genera in Tofieldiaceae has been controversial. Many authors do not recognize Triantha as a separate genus from Tofieldia. [8] [10] [11] A few authors have sunk other genera into Tofieldia as well. [12] A phylogenetic analysis, based on two nuclear genes and ten chloroplast genes, has confirmed that all five of the genera are monophyletic and that Triantha and Tofieldia are close sister taxa. [13]
Small herbaceous perennials. Leaves equitant (distichous and overlapping), isobifacial. Inflorescence a raceme. Calyculus present. Tepals free. Fruit a capsule, usually septicidal.
The following phylogenetic tree is based on a DNA paper that was published in 2011, [13] modified to accommodate the transfer of the species in Isidrogalvia to Harperocallis.
Tofieldiaceae | |||||||||||||||||||
The Polygonaceae are a family of flowering plants known informally as the knotweed family or smartweed—buckwheat family in the United States. The name is based on the genus Polygonum, and was first used by Antoine Laurent de Jussieu in 1789 in his book, Genera Plantarum. The name may refer to the many swollen nodes the stems of some species have, being derived from Greek, poly meaning 'many' and gony meaning 'knee' or 'joint'. Alternatively, it may have a different derivation, meaning 'many seeds'.
Cornaceae is a cosmopolitan family of flowering plants in the order Cornales. It contains approximately 85 species in two genera, Alangium and Cornus. They are mostly trees and shrubs, which may be deciduous or evergreen, although a few species are perennial herbs. Members of the family usually have opposite or alternate simple leaves, four- or five-parted flowers clustered in inflorescences or pseudanthia, and drupaceous fruits. The family is primarily distributed in northern temperate regions and tropical Asia. In northern temperate areas, Cornaceae are well known from the dogwoods Cornus.
Dioscoreaceae is a family of monocotyledonous flowering plants, with about 715 known species in nine genera. The best-known member of the family is the yam.
Melanthiaceae, also called the bunchflower family, is a family of flowering herbaceous perennial plants native to the Northern Hemisphere. Along with many other lilioid monocots, early authors considered members of this family to belong to the family Liliaceae, in part because both their sepals and petals closely resemble each other and are often large and showy like those of lilies, while some more recent taxonomists have placed them in a family Trilliaceae. The most authoritative modern treatment, however, the APG III system of 2009, places the family in the order Liliales, in the clade monocots. Circumscribed in this way, the family includes up to 17 genera.
Petrosaviaceae is a family of flowering plants belonging to a monotypic order, Petrosaviales. Petrosaviales are monocots, and are grouped within the lilioid monocots. Petrosaviales are a very small order of photosynthetic (Japonolirion) and rare leafless achlorophyllous, mycoheterotrophic plants (Petrosavia) found in dark montane rainforests in Japan, China, Southeast Asia and Borneo. They are characterised by having bracteate racemes, pedicellate flowers, six persistent tepals, septal nectaries, three almost distinct carpels, simultaneous microsporogenesis, monosulcate pollen, and follicular fruit.
Burmanniaceae is a family of flowering plants, consisting of 99 species of herbaceous plants in eight genera.
Brodiaeoideae are a monocot subfamily of flowering plants in the family Asparagaceae, order Asparagales. They have been treated as a separate family, Themidaceae. They are native to Central America and western North America, from British Columbia to Guatemala. The name of the subfamily is based on the type genus Brodiaea.
Saxifragaceae is a family of herbaceous perennial flowering plants, within the core eudicot order Saxifragales. The taxonomy of the family has been greatly revised and the scope much reduced in the era of molecular phylogenetic analysis. The family is divided into ten clades, with about 640 known species in about 35 accepted genera. About half of these consist of a single species, but about 400 of the species are in the type genus Saxifraga. The family is predominantly distributed in the northern hemisphere, but also in the Andes in South America.
The Potamogetonaceae, commonly referred to as the pondweed family, is an aquatic family of monocotyledonous flowering plants. The roughly 110 known species are divided over six genera. The largest genus in the family by far is Potamogeton, which contains about 100 species.
Zosteraceae is a family of marine perennial flowering plants found in temperate and subtropical coastal waters, with the highest diversity located around Korea and Japan. Most seagrasses complete their entire life cycle under water, having filamentous pollen especially adapted to dispersion in an aquatic environment and ribbon-like leaves that lack stomata. Seagrasses are herbaceous and have prominent creeping rhizomes. A distinctive characteristic of the family is the presence of characteristic retinacules, which are present in all species except members of Zostera subgenus Zostera.
Asparagaceae, known as the asparagus family, is a family of flowering plants, placed in the order Asparagales of the monocots. Its best known member is Asparagus officinalis, garden asparagus.
Myrothamnus is a genus of flowering plants, consisting of two species of small xerophytic shrubs, in the southern parts of tropical Africa and in Madagascar. Myrothamnus is recognized as the only genus in the family Myrothamnaceae.
Triuridaceae are a family of tropical and subtropical flowering plants, including nine genera with a total of ca 55 known species. All members lack chlorophyll and are mycoheterotrophic. The heterotrophic lifestyle of these plants has resulted in a loss of xylem vessels and stomata, and a reduction of leaves to scales.
Cyclanthaceae is a family of flowering plants.
Campynemataceae (Campynemaceae) is a family of flowering plants. The family consists of two genera and four species of perennial herbaceous plants endemic to New Caledonia and Tasmania.
Boryaceae is a family of highly drought-tolerant flowering plants native to Australia, placed in the order Asparagales of the monocots. The family includes two genera, with twelve species in total in Australia.
Tecophilaeaceae is a family of flowering plants, placed in the order Asparagales of the monocots. It consists of nine genera with a total of 27 species.
Molluginaceae are a family of flowering plants recognized by several taxonomists. It was previously included in the larger family Aizoaceae. The APG III system of 2009 made no change in the status of the family as compared to the APG II system of 2003 and the APG system of 1998, apart from a reassignment of several genera, such as the placement of Corrigiola and Telephium into Caryophyllaceae, Corbichonia in Lophiocarpaceae, Microtea into Microteaceae and Limeum in Limeaceae, because the family was found to be widely polyphyletic in Caryophyllales. In addition Macarthuria was found not to be related to Limeum as previously thought and thus it was placed in Macarthuriaceae, and similarly species formerly placed in Hypertelis, apart from type species Hypertelis spergulacea, a true Molluginaceae, were found to belong elsewhere and were described as Kewa in the family Kewaceae, named for the Royal Botanic Gardens Kew. Molluginaceae is still assigned to the order Caryophyllales in the clade core eudicots, although the generic circumscription is difficult because Mollugo is not monophyletic.
Haloragaceae is a eudicot flowering plant family in the order Saxifragales, based on the phylogenetic APG system. In the Cronquist system, it was included in the order Haloragales.
The taxonomy of Liliaceae has had a complex history since the first description of this flowering plant family in the mid-eighteenth century. Originally, the Liliaceae or Lily family were defined as having a "calix" (perianth) of six equal-coloured parts, six stamens, a single style, and a superior, three-chambered (trilocular) ovary turning into a capsule fruit at maturity. The taxonomic circumscription of the family Liliaceae progressively expanded until it became the largest plant family and also extremely diverse, being somewhat arbitrarily defined as all species of plants with six tepals and a superior ovary. It eventually came to encompass about 300 genera and 4,500 species, and was thus a "catch-all" and hence paraphyletic taxon. Only since the more modern taxonomic systems developed by the Angiosperm Phylogeny Group (APG) and based on phylogenetic principles, has it been possible to identify the many separate taxonomic groupings within the original family and redistribute them, leaving a relatively small core as the modern family Liliaceae, with fifteen genera and 600 species.
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