List of rhinolophids

Last updated

Rufous horseshoe bat (Rhinolophus rouxii) Rhinolophus rouxii.jpg
Rufous horseshoe bat (Rhinolophus rouxii)

Rhinolophidae is one of the twenty families of bats in the mammalian order Chiroptera and part of the microbat suborder. A member of this family is called a rhinolophid or a horseshoe bat. They are named for the shape of their nose-leaf. They are found in Europe, Africa, Asia, and Australia, primarily in forests, savannas, grasslands, and caves, though some species can also be found in deserts or wetlands. They range in size from the Thai horseshoe bat and little Nepalese horseshoe bat, at 3 cm (1 in) plus a 1 cm (0.4 in) tail, to the Maclaud's horseshoe bat, at 10 cm (4 in) plus a 5 cm (2 in) tail. Like all bats, rhinolophids are capable of true and sustained flight, and have wing lengths ranging from 3 cm (1 in) in the Formosan lesser horseshoe bat and little Nepalese horseshoe bat, to 8 cm (3 in) in the great woolly horseshoe bat. They are all insectivorous and eat a variety of insects and spiders. [1] Most rhinolophids do not have population estimates, but the ones that do range from 900 adult individuals to 20,000. Twelve species—the Andaman horseshoe bat, Bornean woolly horseshoe bat, Guinean horseshoe bat, King horseshoe bat, Maclaud's horseshoe bat, Mount Mabu horseshoe bat, Poso horseshoe bat, Ruwenzori horseshoe bat, Timorese horseshoe bat, Willard's horseshoe bat, Yaeyama little horseshoe bat, and Ziama horseshoe bat—are categorized as endangered species, and Hill's horseshoe bat is categorized as critically endangered. These collectively make up almost fifteen percent of the species in the family.

Contents

The 92 extant species of Rhinolophidae are all included in a single genus, Rhinolophus. A few extinct prehistoric rhinolophid species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed. [2]

Conventions

IUCN Red List categories
Conservation status
 EX  Extinct (0 species)
 EW  Extinct in the wild (0 species)
 CR  Critically Endangered (1 species)
 EN  Endangered (12 species)
 VU  Vulnerable (5 species)
 NT  Near threatened (9 species)
 LC  Least concern (50 species)
Other categories
 DD  Data deficient (13 species)
 NE  Not evaluated (2 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the rhinolophid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

The family Rhinolophidae consists of a single genus, Rhinolophus, containing 92 species.

Rhinolophids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. [3]

Genus Rhinolophus Lacépède, 1799 – 92 species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Acuminate horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.15051.b ven - Rhinolophus acuminatus acuminatus - skin.jpeg

R. acuminatus
Peters, 1871
Southeastern Asia
Acuminate Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–4 cm (0.4–1.6 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Forest and caves [5]
 LC 


Unknown Blue question mark (italic).svg [5]

Adam's horseshoe bat


R. adami
Aellen & Brosset, 1968
Republic of the Congo
Adam's Horseshoe Bat area.png
Size: Unknown length, plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest and caves [7]
 DD 


Unknown Blue question mark (italic).svg [7]

Andaman horseshoe bat

Rhinolophus affinis 86903750.jpg

R. cognatus
K. Andersen, 1906
Andaman Islands
Andaman Horseshoe Bat area.png
Size: About 4 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [8]
 EN 


Unknown Decrease2.svg [8]

Arcuate horseshoe bat

Kelawar ladam2.jpg

R. arcuatus
Peters, 1871

Two subspecies
  • R. a. arcuatus
  • R. a. exiguus
Southeastern Asia
Arcuate Horseshoe Bat area.png
Size: 4–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [10]
 DD 


Unknown Steady2.svg [10]

Beddome's horseshoe bat

Lesser Woolly Horseshoe Bat (Rhinolophus beddomei).jpg

R. beddomei
K. Andersen, 1905
Southern India and Sri Lanka
Lesser Woolly Horseshoe Bat area.png
Size: 6–8 cm (2–3 in), plus 3–5 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length [11]

Habitat: Forest and caves [12]
 LC 


Unknown Steady2.svg [12]

Big-eared horseshoe bat


R. macrotis
Blyth, 1844

Five subspecies
  • R. m. caldwelli
  • R. m. episcopus
  • R. m. huananus
  • R. m. macrotis
  • R. m. topali
Eastern and southeastern Asia
Big-eared Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–4 cm (0.4–1.6 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Forest and caves [14]
 LC 


Unknown Steady2.svg [14]

Blasius's horseshoe bat

Rhinolopus blasii.png

R. blasii
Peters, 1866
Scattered Africa, southern Europe, and western Asia
Blasius's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest, savanna, shrubland, caves, and desert [15]
 LC 


Unknown Decrease2.svg [15]

Blyth's horseshoe bat

Rhinolophus lepidus by Davidraju.jpg

R. lepidus
Blyth, 1844

Two subspecies
  • R. l. lepidus
  • R. l. refulgens
Southern and southeastern Asia
Blyth's Horseshoe Bat area.png
Size: 3–6 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [16]
 LC 


Unknown Blue question mark (italic).svg [16]

Bokhara horseshoe bat

Rhinolophus bocharicus.jpg

R. bocharicus
Kastschenko & Akimov, 1917
Central Asia
Bokhara Horseshoe Bat area.png
Size: 3–5 cm (1–2 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Shrubland, grassland, rocky areas, caves, and desert [17]
 LC 


Unknown Steady2.svg [17]

Bornean horseshoe bat

Bornean Horseshoe Bat (Rhinolophus borneensis) maybe%3F (7113337169).jpg

R. borneensis
Peters, 1861
Southeastern Asia
Bornean Horseshoe Bat area.png
Size: Unknown length, plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Forest, rocky areas, and caves [18]
 LC 


Unknown Blue question mark (italic).svg [18]

Bornean woolly horseshoe bat


R. proconsulis
Hill, 1959
Borneo Size: Unknown length
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest [19]
 EN 


Unknown Blue question mark (italic).svg [19]

Bourret's horseshoe bat


R. paradoxolophus
Bourret, 1951
Southeastern Asia
Bourret's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–4 cm (0.4–1.6 in) tail
5–7 cm (2–3 in) arm/wing length [13]

Habitat: Forest and caves [20]
 LC 


Unknown Blue question mark (italic).svg [20]

Broad-eared horseshoe bat

Rhinolophus euryotis illustration 2.png

R. euryotis
Temminck, 1835
Southeastern Asia
Broad-eared Horseshoe Bat area.png
Size: 5–8 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Caves and forest [21]
 LC 


Unknown Blue question mark (italic).svg [21]

Bushveld horseshoe bat


R. simulator
K. Andersen, 1904
Eastern and western Africa
Bushveld Horseshoe Bat area.png
Size: 3–7 cm (1–3 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [6]

Habitat: Savanna and caves [22]
 LC 


Unknown Decrease2.svg [22]

Canut's horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.32724.b reg - Rhinolophus canuti - skin.jpeg

R. canuti
Thomas & Wroughton, 1909
Indonesia
Canut's Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Forest and caves [23]
 VU 


7,500–10,000 Decrease2.svg [23]

Cape horseshoe bat

Rhinolophus capensis.jpg

R. capensis
Lichtenstein, 1823
Southern Africa
Cape Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Shrubland and caves [24]
 LC 


Unknown Steady2.svg [24]

Chiewkwee's horseshoe bat


R. chiewkweeae
Yoshiyuki & Liat, 2005
Southeastern AsiaSize: 5–7 cm (2–3 in), plus 1–2 cm (0.4–0.8 in) tail
4–6 cm (2 in) arm/wing length [11]

Habitat: Forest [25]
 DD 


Unknown Blue question mark (italic).svg [25]

Chinese rufous horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.33160.b ven - Rhinolophus sinicus - skin.jpeg

R. sinicus
K. Andersen, 1905
Eastern Asia
Chinese Rufous Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [26]
 LC 


Unknown Blue question mark (italic).svg [26]

Cohen's horseshoe bat


R. cohenae
Taylor, Stoffberg, Monadjem, Schoeman, Bayliss, & Cotterill, 2012
South AfricaSize: 7–9 cm (3–4 in), plus 3–5 cm (1–2 in) tail
6–7 cm (2–3 in) arm/wing length [6]

Habitat: Savanna, grassland, rocky areas, and caves [27]
 VU 


900–1,000 Decrease2.svg [27]

Convex horseshoe bat


R. convexus
Csorba, 1997
Southeastern Asia
Convex Horseshoe Bat area.png
Size: Unknown length, plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Forest [28]
 DD 


Unknown Blue question mark (italic).svg [28]

Creagh's horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.33449 lat - Rhinolophus creaghi - skull.jpeg

R. creaghi
Thomas, 1896
Southeastern Asia
Creagh's Horseshoe Bat area.png
Size: About 5 cm (2 in), plus 1–2 cm (0.4–0.8 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Forest and caves [29]
 LC 


Unknown Decrease2.svg [29]

Croslet horseshoe bat


R. coelophyllus
Peters, 1867
Southeastern Asia
Croslet Horseshoe Bat area.png
Size: 3–5 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [30]
 LC 


Unknown Steady2.svg [30]

Damara horseshoe bat


R. damarensis
Roberts, 1946
Southern AfricaSize: 6–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Savanna, shrubland, and desert [31]
 LC 


20,000 Blue question mark (italic).svg [31]

Darling's horseshoe bat


R. darlingi
K. Andersen, 1905
Southern Africa
Darling's Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Savanna and caves [32]
 LC 


Unknown Blue question mark (italic).svg [32]

Decken's horseshoe bat

Naturalis Biodiversity Center - ZMA.MAM.4175.b dor - Rhinolophus deckenii - skin.jpeg

R. deckenii
Peters, 1867
Eastern Africa
Decken's Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest and caves [33]
 NT 


Unknown Decrease2.svg [33]

Dent's horseshoe bat


R. denti
Thomas, 1904
Southern and western Africa
Dent's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [6]

Habitat: Savanna and caves [34]
 LC 


Unknown Blue question mark (italic).svg [34]

Dobson's horseshoe bat


R. yunanensis
Dobson, 1872
Southeastern Asia
Dobson's Horseshoe Bat area.png
Size: 5–8 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [11]

Habitat: Forest and caves [35]
 LC 


Unknown Blue question mark (italic).svg [35]

Eloquent horseshoe bat

Naturalis Biodiversity Center - ZMA.MAM.20842.a pal - Rhinolophus eloquens - skull.jpeg

R. eloquens
K. Andersen, 1905
Eastern Africa
Eloquent Horseshoe Bat area.png
Size: 6–8 cm (2–3 in), plus 2–5 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length [6]

Habitat: Forest, savanna, and caves [36]
 LC 


Unknown Blue question mark (italic).svg [36]

Forest horseshoe bat


R. silvestris
Aellen, 1959
West-central Africa
Forest Horseshoe Bat area.png
Size: Unknown length, plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Forest and caves [37]
 DD 


Unknown Decrease2.svg [37]

Formosan lesser horseshoe bat

Formosan Woolly Horseshoe Bat imported from iNaturalist photo 174918399 on 16 August 2024.jpg

R. monoceros
Temminck, 1835
Japan and ChinaSize: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–4 cm (1–2 in) arm/wing length [4]

Habitat: Unrecorded
 NE 


Unknown Blue question mark (italic).svg

Formosan woolly horseshoe bat

Rhinolophus formosae.jpg

R. formosae
Sanborn, 1939
Taiwan
Rhinolophus formosae area.png
Size: 6–9 cm (2–4 in), plus 2–5 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length [11]

Habitat: Forest and caves [38]
 LC 


Unknown Decrease2.svg [38]

Geoffroy's horseshoe bat

Rhinolophus clivosus.jpg

R. clivosus
Cretzschmar, 1828

Two subspecies
  • R. c. brachygnathus
  • R. c. clivosus
Africa and Arabian peninsula
Geoffroy's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Forest, savanna, shrubland, caves, and desert [39]
 LC 


Unknown Blue question mark (italic).svg [39]

Great woolly horseshoe bat

Woolly Horseshoe Bat imported from iNaturalist photo 246196123 on 16 August 2024.jpg

R. luctus
Temminck, 1834
Southern and southeastern Asia
Woolly Horseshoe Bat area.png
Size: 7–9 cm (3–4 in), plus 3–6 cm (1–2 in) tail
5–8 cm (2–3 in) arm/wing length [11]

Habitat: Caves, rocky areas, and forest [40]
 LC 


Unknown Blue question mark (italic).svg [40]

Greater horseshoe bat

Grand Rhinolophe.jpg

R. ferrumequinum
(Schreber, 1774)

Seven subspecies
  • R. f. creticum
  • R. f. ferrumequinum
  • R. f. irani
  • R. f. korai
  • R. f. nippon
  • R. f. proximus
  • R. f. tragatus
Europe, Asia, and northern Africa
Greater Horseshoe Bat area.png
Size: 5–8 cm (2–3 in), plus 3–5 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length [13]

Habitat: Forest, shrubland, grassland, and caves [41]
 LC 


Unknown Decrease2.svg [41]

Guinean horseshoe bat


R. guineensis
Eisentraut, 1960
Western Africa
Guinean Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest, savanna, grassland, inland wetlands, and caves [42]
 EN 


Unknown Decrease2.svg [42]

Halcyon horseshoe bat


R. alcyone
Temminck, 1852
Western and central Africa
Halcyon Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 1–4 cm (0.4–1.6 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest, savanna, and caves [43]
 LC 


Unknown Blue question mark (italic).svg [43]

Hildebrandt's horseshoe bat


R. hildebrandtii
Peters, 1878
Eastern Africa
Hildebrandt's Horseshoe Bat area.png
Size: 7–9 cm (3–4 in), plus 3–5 cm (1–2 in) tail
6–7 cm (2–3 in) arm/wing length [6]

Habitat: Forest, savanna, and caves [44]
 LC 


Unknown Blue question mark (italic).svg [44]

Hill's horseshoe bat


R. hilli
Aellen, 1973
Rwanda
Rhinolophus hilli area.png
Size: About 6 cm (2 in), plus about 3 cm (1 in) tail
About 5 cm (2 in) arm/wing length [13]

Habitat: Forest and caves [45]
 CR 


Unknown Decrease2.svg [45]

Hills' horseshoe bat


R. hillorum
Koopman, 1989
Scattered Equatorial Africa
Hill's Horseshoe Bat area.png
Size: 6–8 cm (2–3 in), plus 3–5 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Forest, grassland, and caves [46]
 VU 


Unknown Decrease2.svg [46]

Imaizumi's horseshoe bat


R. imaizumii
Hill & Yoshiyuki, 1980
JapanSize: 3–5 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Unrecorded
 NE 


Unknown Blue question mark (italic).svg

Indo-Chinese lesser brown horseshoe bat


R. microglobosus
Csorba and Jenkins, 1998
Southeastern Asia
Range Rhinolophus microglobosus.png
Size: 3–6 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Caves and forest [47]
 LC 


Unknown Steady2.svg [47]

Insular horseshoe bat

Rhinolophus keyensis 247104551.jpg

R. keyensis
Peters, 1871
Indonesia
Insular Horseshoe Bat area.png
Size: 3–5 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Unknown [48]
 DD 


Unknown Blue question mark (italic).svg [48]

Intermediate horseshoe bat

Rhinolophus affinis.jpg

R. affinis
Horsfield, 1823

Nine subspecies
  • R. a. affinis
  • R. a. andamanensis
  • R. a. hainanus
  • R. a. himalayanus
  • R. a. macrurus
  • R. a. nesites
  • R. a. princes
  • R. a. superans
  • R. a. tener
Southeastern Asia
Rhinolophus affinis area.png
Size: 4–7 cm (2–3 in), plus 1–4 cm (0.4–1.6 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [49]
 LC 


Unknown Steady2.svg [49]

King horseshoe bat


R. rex
Allen, 1923
Eastern China
King Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–4 cm (0.4–1.6 in) tail
5–7 cm (2–3 in) arm/wing length [13]

Habitat: Forest and caves [50]
 EN 


1,500–2,000 Decrease2.svg [50]

Lander's horseshoe bat

Rhinolophus landeri.jpg

R. landeri
Martin, 1837
Sub-Saharan Africa
Lander's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus about 3 cm (1 in) tail
3–5 cm (1–2 in) arm/wing length [6]

Habitat: Forest, savanna, shrubland, and caves [51]
 LC 


Unknown Blue question mark (italic).svg [51]

Large rufous horseshoe bat


R. rufus
Eydoux & Gervais, 1836
Philippines
Large Rufous Horseshoe Bat area.png
Size: 8–10 cm (3–4 in), plus 3–4 cm (1–2 in) tail
6–8 cm (2–3 in) arm/wing length [9]

Habitat: Forest and caves [52]
 NT 


Unknown Decrease2.svg [52]

Large-eared horseshoe bat

Rhinolophus philippinensis.jpg

R. philippinensis
Waterhouse, 1843
Southeastern Asia and northeastern Australia
Large-eared Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Caves and forest [53]
 LC 


Unknown Blue question mark (italic).svg [53]

Least horseshoe bat

Rhinolophus pusillus.jpg

R. pusillus
Temminck, 1834
Eastern and southeastern Asia
Least Horseshoe Bat area.png
Size: 3–5 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [54]
 LC 


Unknown Steady2.svg [54]

Lesser brown horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.32722.b ven - Rhinolophus stheno - skin.jpeg

R. stheno
K. Andersen, 1905
Southeastern Asia
Lesser Brown Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest [55]
 LC 


Unknown Steady2.svg [55]

Lesser horseshoe bat

Bat(20070605).jpg

R. hipposideros
(Bechstein, 1800)
Europe, western Asia, and northern Africa
Lesser Horseshoe Bat area.png
Size: 3–5 cm (1–2 in), plus 2–4 cm (1–2 in) tail
3–5 cm (1–2 in) arm/wing length [9]

Habitat: Forest, shrubland, grassland, and caves [56]
 LC 


Unknown Decrease2.svg [56]

Lesser woolly horseshoe bat

Lesser Woolly Horseshoe Bat (Rhinolophus sedulus) (8688716960).jpg

R. sedulus
K. Andersen, 1905
Southeastern Asia
Rhinolophus sedulus area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [9]

Habitat: Forest and caves [57]
 NT 


Unknown Decrease2.svg [57]

Little Japanese horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.20927.b ven - Rhinolophus cornutus - skin.jpeg

R. cornutus
Temminck, 1834

Three subspecies
  • R. c. cornutus
  • R. c. orii
  • R. c. pumilus
Japan and ChinaSize: 3–6 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest [58]
 NT 


Unknown Blue question mark (italic).svg [58]

Little Nepalese horseshoe bat


R. subbadius
Blyth, 1844
Eastern Asia
Little Nepalese Horseshoe Bat area.png
Size: 3–4 cm (1–2 in), plus 1–2 cm (0.4–0.8 in) tail
3–4 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [59]
 LC 


Unknown Decrease2.svg [59]

Maclaud's horseshoe bat


R. maclaudi
Pousargues, 1897
Western Africa
Maclaud's Horseshoe Bat area.png
Size: 7–10 cm (3–4 in), plus 3–5 cm (1–2 in) tail
6–7 cm (2–3 in) arm/wing length [13]

Habitat: Savanna and caves [60]
 EN 


Unknown Decrease2.svg [60]

Madura horseshoe bat


R. madurensis
K. Andersen, 1918
Indonesia
Rhinolophus madurensis area.png
Size: Unknown length
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [61]
 VU 


Unknown Decrease2.svg [61]

Maendeleo horseshoe bat


R. maendeleo
Kock, Csorba, & Howell, 2000
Eastern Africa
Maendeleo Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest and caves [62]
 DD 


Unknown Blue question mark (italic).svg [62]

Malayan horseshoe bat


R. malayanus
Bonhote, 1903
Southeastern Asia
Malayan Horseshoe Bat area.png
Size: 3–5 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [63]
 LC 


Unknown Steady2.svg [63]

Marshall's horseshoe bat


R. marshalli
Thonglongya, 1973
Southeastern Asia
Marshall's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [13]

Habitat: Forest, rocky areas, and caves [64]
 LC 


Unknown Blue question mark (italic).svg [64]

McIntyre's horseshoe bat


R. mcintyrei
Hill & Schlitter, 1982
Papua New Guinea Size: 4–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest [65]
 DD 


Unknown Blue question mark (italic).svg [65]

Mediterranean horseshoe bat

Rhinolophus euryale-cropped.jpg

R. euryale
Blasius, 1853
Europe, western Asia, and northern Africa
Mediterranean Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest, shrubland, and caves [66]
 NT 


Unknown Decrease2.svg [66]

Mehely's horseshoe bat

Rhinolophus mehelyi-cropped.jpg

R. mehelyi
Matschie, 1901
Southern Europe, western Asia, and northern Africa
Mehely's Horseshoe Bat area.png
Size: 4–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
2–4 cm (1–2 in) arm/wing length [6]

Habitat: Shrubland and caves [67]
 VU 


Unknown Decrease2.svg [67]

Mitred horseshoe bat

EB1911 Chiroptera Fig. 6.jpg

R. mitratus
Blyth, 1844
Eastern India
Mitred Horseshoe Bat area.png
Size: About 7 cm (3 in), plus about 3 cm (1 in) tail
About 6 cm (2 in) arm/wing length [11]

Habitat: Unknown [68]
 DD 


Unknown Blue question mark (italic).svg [68]

Mount Mabu horseshoe bat


R. mabuensis
Taylor, Stoffberg, Monadjem, Schoeman, Bayliss, & Cotterill, 2012
Mozambique Size: 7–9 cm (3–4 in), plus 3–5 cm (1–2 in) tail
6–7 cm (2–3 in) arm/wing length [6]

Habitat: Caves and forest [69]
 EN 


Unknown Blue question mark (italic).svg [69]

Mozambican horseshoe bat

Rhinolophus mossambicus.jpg

R. mossambicus
Taylor, Stoffberg, Monadjem, Schoeman, Bayliss, & Cotterill, 2012
Southeastern AfricaSize: Unknown length
6–7 cm (2–3 in) arm/wing length [6]

Habitat: Forest, savanna, and caves [70]
 LC 


Unknown Decrease2.svg [70]

Neriad horseshoe bat


R. nereis
K. Andersen, 1905
Siantan Island in Indonesia
Neriad Horseshoe Bat area.png
Size: Unknown length, plus about 2 cm (1 in) tail
About 4 cm (2 in) arm/wing length [4]

Habitat: Unknown [71]
 DD 


Unknown Blue question mark (italic).svg [71]

Osgood's horseshoe bat


R. osgoodi
Sanborn, 1939
Southern China
Osgood's Horseshoe Bat area.png
Size: Unknown length, plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Caves [72]
 LC 


Unknown Steady2.svg [72]

Pearson's horseshoe bat

Rhinolophus pearsonii.jpg

R. pearsonii
Horsfield, 1851
Eastern and southeastern Asia
Pearson's Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [11]

Habitat: Forest and caves [73]
 LC 


Unknown Blue question mark (italic).svg [73]

Peninsular horseshoe bat


R. robinsoni
K. Andersen, 1918
Malaysia and Thailand
Rhinolophus robinsoni area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Rocky areas and forest [74]
 NT 


Unknown Decrease2.svg [74]

Philippine forest horseshoe bat


R. inops
K. Andersen, 1905
Philippines
Philippine Forest Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest [75]
 LC 


Unknown Steady2.svg [75]

Poso horseshoe bat


R. belligerator
Patrick, McCulloch, & Ruedas, 2013
Sulawesi Island in IndonesiaSize: 5–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Unknown [76]
 EN 


Unknown Blue question mark (italic).svg [76]

Rufous horseshoe bat

Rhinolophus rouxii.jpg

R. rouxii
Temminck, 1835
Southern Asia
Rufous Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [77]
 LC 


Unknown Blue question mark (italic).svg [77]

Ruwenzori horseshoe bat


R. ruwenzorii
Hill, 1942
Central Africa
Ruwenzori Horseshoe Bat area.png
Size: 5–8 cm (2–3 in), plus 2–4 cm (1–2 in) tail
5–7 cm (2–3 in) arm/wing length [13]

Habitat: Forest, shrubland, grassland, and caves [78]
 EN 


Unknown Decrease2.svg [78]

Rüppell's horseshoe bat

Rhinolophus fumigatus, Limpopo.jpg

R. fumigatus
Rüppell, 1842
Sub-Saharan Africa
Ruppell's Horseshoe Bat area.png
Size: 4–8 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest, savanna, and caves [79]
 LC 


Unknown Blue question mark (italic).svg [79]

Sakeji horseshoe bat


R. sakejiensis
Cotterill, 2002
Zambia
Sakeji Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Savanna [80]
 DD 


Unknown Blue question mark (italic).svg [80]

Shamel's horseshoe bat


R. shameli
Tate, 1943
Southeastern Asia
Shamel's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [81]
 LC 


Unknown Steady2.svg [81]

Shortridge's horseshoe bat


R. shortridgei
K. Andersen, 1918
Eastern Asia
Rhinolophus shortridgei area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest [82]
 DD 


Unknown Blue question mark (italic).svg [82]

Small rufous horseshoe bat


R. subrufus
K. Andersen, 1905
Philippines
Small Rufous Horseshoe Bat area.png
Size: 6–7 cm (2–3 in), plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest and caves [83]
 DD 


Unknown Blue question mark (italic).svg [83]

Smaller horseshoe bat

Rhinolophus megaphyllus.jpg

R. megaphyllus
Gray, 1834
Eastern Australia and Papua New Guinea
Smaller Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Caves and forest [84]
 LC 


Unknown Blue question mark (italic).svg [84]

Smithers's horseshoe bat

Rhinolophus smithersi.jpg

R. smithersi
Taylor, Stoffberg, Monadjem, Schoeman, Bayliss, & Cotterill, 2012
Southern AfricaSize: 7–9 cm (3–4 in), plus 3–5 cm (1–2 in) tail
6–7 cm (2–3 in) arm/wing length [6]

Habitat: Savanna, shrubland, inland wetlands, and caves [85]
 NT 


Unknown Steady2.svg [85]

Sulawesi broad-eared horseshoe bat


R. tatar
Bergmans & Rozendaal, 1982
Sulawesi Island in IndonesiaSize: 5–7 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest [86]
 LC 


Unknown Blue question mark (italic).svg [86]

Sulawesi horseshoe bat

Naturalis Biodiversity Center - RMNH.MAM.15058.b ven - Rhinolophus celebensis javanicus - skin.jpeg

R. celebensis
K. Andersen, 1905
Indonesia
Sulawesi Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Forest and caves [87]
 LC 


Unknown Blue question mark (italic).svg [87]

Swinny's horseshoe bat


R. swinnyi
Gough, 1908
Southern Africa
Swinny's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [6]

Habitat: Forest, savanna, and caves [88]
 LC 


Unknown Blue question mark (italic).svg [88]

Thai horseshoe bat


R. siamensis
Gyldenstolpe, 1917
Southeastern Asia
Rhinolophus siamensis area.png
Size: 3–4 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [13]

Habitat: Forest and caves [89]
 LC 


Unknown Blue question mark (italic).svg [89]

Thailand horseshoe bat


R. thailandensis
Wu, Harada, & Motokawa, 2009
Northern Thailand
Range Rhinolophus thailandensis.png
Size: 6–8 cm (2–3 in), plus 1–3 cm (0.4–1.2 in) tail
5–7 cm (2–3 in) arm/wing length [11]

Habitat: Forest and caves [90]
 LC 


Unknown Steady2.svg [90]

Thomas's horseshoe bat

Thomas's horseshoe bat imported from iNaturalist photo 233070250 on 16 August 2024.jpg

R. thomasi
K. Andersen, 1905
Southeastern Asia
Thomas's Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Caves [91]
 LC 


Unknown Blue question mark (italic).svg [91]

Timorese horseshoe bat


R. montanus
Goodwin, 1979
East Timor
Rhinolophus montanus area.png
Size: 4–6 cm (2 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Caves and unknown [92]
 EN 


Unknown Blue question mark (italic).svg [92]

Trefoil horseshoe bat

Rhinolophus trifoliatus.jpg

R. trifoliatus
Temminck, 1834
Southeastern Asia
Trefoil Horseshoe Bat area.png
Size: 5–7 cm (2–3 in), plus 2–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [9]

Habitat: Forest [93]
 NT 


Unknown Decrease2.svg [93]

Wedge-sellaed horseshoe bat


R. xinanzhongguoensis
Zhou, Guillén-Servent, Lim, Eger, Wang, & Jiang, 2009
ChinaSize: 5–7 cm (2–3 in), plus 3–4 cm (1–2 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Forest [94]
 NT 


Unknown Decrease2.svg [94]

Willard's horseshoe bat


R. willardi
Peterhans & Fahr, 2013
Democratic Republic of the CongoSize: 6–8 cm (2–3 in), plus 2–3 cm (1 in) tail
4–6 cm (2 in) arm/wing length [13]

Habitat: Forest and caves [95]
 EN 


Unknown Decrease2.svg [95]

Yaeyama little horseshoe bat


R. perditus
K. Andersen, 1918
Yaeyama Islands in JapanSize: 3–5 cm (1–2 in), plus 1–3 cm (0.4–1.2 in) tail
4–6 cm (2 in) arm/wing length [4]

Habitat: Caves, inland wetlands, and forest [96]
 EN 


Unknown Decrease2.svg [96]

Yellow-faced horseshoe bat


R. virgo
K. Andersen, 1905
Philippines
Yellow-faced Horseshoe Bat area.png
Size: 4–6 cm (2 in), plus 1–3 cm (0.4–1.2 in) tail
3–5 cm (1–2 in) arm/wing length [4]

Habitat: Caves and forest [97]
 LC 


Unknown Steady2.svg [97]

Ziama horseshoe bat


R. ziama
Fahr, Vierhaus, Hütterer, & Kock, 2002
Western Africa
Rhinolophus ziama area.png
Size: 7–8 cm (3 in), plus 3–4 cm (1–2 in) tail
About 6 cm (2 in) arm/wing length [13]

Habitat: Caves and forest [98]
 EN 


Unknown Decrease2.svg [98]

Related Research Articles

<span class="mw-page-title-main">Croslet horseshoe bat</span> Species of bat

The croslet horseshoe bat is a species of bat in the family Rhinolophidae. It is found in Laos, Malaysia, Myanmar, and Thailand.

<span class="mw-page-title-main">Convex horseshoe bat</span> Species of bat

The convex horseshoe bat is a species of bat in the family Rhinolophidae. It is found in Malaysia and Laos.

<span class="mw-page-title-main">Eloquent horseshoe bat</span> Species of bat

The eloquent horseshoe bat is a species of bat in the family Rhinolophidae. It is found in Ethiopia, Kenya, Rwanda, Somalia, South Sudan, Tanzania, and Uganda. Its natural habitats are subtropical or tropical moist lowland forests, dry savanna, moist savanna, and caves.

<span class="mw-page-title-main">Blyth's horseshoe bat</span> Species of bat

Blyth's horseshoe bat is a species of bat in the family Rhinolophidae. It is found across southern Asia from Afghanistan to Vietnam. The species can be identified from its pointed, bifid sella.

<span class="mw-page-title-main">Neriad horseshoe bat</span> Species of bat

The neriad horseshoe bat is a species of bat in the family Rhinolophidae.

<span class="mw-page-title-main">Shamel's horseshoe bat</span> Species of bat

Shamel's horseshoe bat is a species of bat in the family Rhinolophidae. It is found in Cambodia, Laos, Myanmar, Thailand and Vietnam.

<span class="mw-page-title-main">Lesser brown horseshoe bat</span> Species of bat

The lesser brown horseshoe bat is a species of bat in the family Rhinolophidae. It is found in Indonesia, Laos, Malaysia, Thailand, and Vietnam.

<span class="mw-page-title-main">Dobson's horseshoe bat</span> Species of bat

Dobson's horseshoe bat is a species of bat in the family Rhinolophidae. It is found in China, India, Myanmar, and Thailand.

<span class="mw-page-title-main">Mozambican horseshoe bat</span> Species of bat

The Mozambican horseshoe bat is a species of horseshoe bat found in southern Africa.

The Mount Mabu horseshoe bat is a species of horseshoe bat. It is endemic to Mozambique.

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  52. 1 2 Alviola, P. A.; Sedlock, J.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Pedregosa, M.; Veluz, M. J.; Tanalgo, K.; Cielo, K. L. S.; Garcia, J. J. L.; Cacho, M. A.; Ong, P.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Heaney, L. (2019). "Rhinolophus rufus". IUCN Red List of Threatened Species . 2019: e.T19564A21995212. doi: 10.2305/IUCN.UK.2019-3.RLTS.T19564A21995212.en .
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  56. 1 2 Taylor, P. (2016). "Rhinolophus hipposideros". IUCN Red List of Threatened Species . 2016: e.T19518A21972794. doi: 10.2305/IUCN.UK.2016-2.RLTS.T19518A21972794.en .
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  60. 1 2 Shapiro, J.; Cooper-Bohannon, R.; Fahr, J. (2019). "Rhinolophus maclaudi". IUCN Red List of Threatened Species . 2019: e.T19549A21978925. doi: 10.2305/IUCN.UK.2019-3.RLTS.T19549A21978925.en .
  61. 1 2 Csorba, G.; Kingston, T.; Hutson, A. M. (2016). "Rhinolophus madurensis". IUCN Red List of Threatened Species . 2016: e.T136410A22016850. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136410A22016850.en .
  62. 1 2 Cooper-Bohannon, R. (2020). "Rhinolophus maendeleo". IUCN Red List of Threatened Species . 2020: e.T44783A22067758. doi: 10.2305/IUCN.UK.2020-2.RLTS.T44783A22067758.en .
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  65. 1 2 Patrick, L.; Ruedas, L. (2017). "Rhinolophus mcintyrei". IUCN Red List of Threatened Species . 2017: e.T84372245A84372277. doi: 10.2305/IUCN.UK.2017-2.RLTS.T84372245A84372277.en .
  66. 1 2 Juste, J.; Alcaldé, J. (2016). "Rhinolophus euryale". IUCN Red List of Threatened Species . 2016: e.T19516A21971185. doi: 10.2305/IUCN.UK.2016-2.RLTS.T19516A21971185.en .
  67. 1 2 Alcaldé, J.; Benda, P.; Juste, J. (2016). "Rhinolophus mehelyi". IUCN Red List of Threatened Species . 2016: e.T19519A21974380. doi: 10.2305/IUCN.UK.2016-2.RLTS.T19519A21974380.en .
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  70. 1 2 Shoeman, C. (2017). "Rhinolophus mossambicus". IUCN Red List of Threatened Species . 2017: e.T64589126A64589338. doi: 10.2305/IUCN.UK.2017-2.RLTS.T64589126A64589338.en .
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  72. 1 2 Sun, K. (2020). "Rhinolophus osgoodi". IUCN Red List of Threatened Species . 2020: e.T19557A21992735. doi: 10.2305/IUCN.UK.2020-3.RLTS.T19557A21992735.en .
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  75. 1 2 Duya, M. R.; Sedlock, J.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Pedregosa, M.; Veluz, M. J.; Jakosalem, G.; Tanalgo, K.; Cielo, K. L. S.; Garcia, J. J. L.; Cacho, M. A. (2019). "Rhinolophus inops". IUCN Red List of Threatened Species . 2019: e.T19545A21978063. doi: 10.2305/IUCN.UK.2019-3.RLTS.T19545A21978063.en .
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  79. 1 2 Monadjem, A.; Griffin, M.; Cotterill, F.; Jacobs, D.; Taylor, P. J. (2017). "Rhinolophus fumigatus". IUCN Red List of Threatened Species . 2017: e.T19541A21980197. doi: 10.2305/IUCN.UK.2017-2.RLTS.T19541A21980197.en .
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  82. 1 2 Sun, K. (2020). "Rhinolophus shortridgei". IUCN Red List of Threatened Species . 2020: e.T136631A21987430. doi: 10.2305/IUCN.UK.2020-2.RLTS.T136631A21987430.en .
  83. 1 2 Ong, P.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Heaney, D.; Balete, M.; Duya, K. (2016). "Rhinolophus subrufus". IUCN Red List of Threatened Species . 2016: e.T19571A21992005. doi: 10.2305/IUCN.UK.2016-2.RLTS.T19571A21992005.en .
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  86. 1 2 Patrick, L.; Ruedas, L. (2017). "Rhinolophus tatar". IUCN Red List of Threatened Species . 2017: e.T84372447A84372450. doi: 10.2305/IUCN.UK.2017-2.RLTS.T84372447A84372450.en .
  87. 1 2 Bouillard, N.; Hutson, A. M.; Suyanto, A.; Kingston, T. (2021). "Rhinolophus celebensis". IUCN Red List of Threatened Species . 2021: e.T19530A21980994. doi: 10.2305/IUCN.UK.2021-2.RLTS.T19530A21980994.en .
  88. 1 2 Monadjem, A.; Cotterill, F. (2017). "Rhinolophus swinnyi". IUCN Red List of Threatened Species . 2017: e.T19572A21992092. doi: 10.2305/IUCN.UK.2017-2.RLTS.T19572A21992092.en .
  89. 1 2 Tu, V.; Furey, N.; Görföl, T.; Csorba, G. (2019). "Rhinolophus siamensis". IUCN Red List of Threatened Species . 2019: e.T136651A21990143. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136651A21990143.en .
  90. 1 2 Bouillard, N. (2021). "Rhinolophus thailandensis". IUCN Red List of Threatened Species . 2021: e.T82348077A82348673. doi: 10.2305/IUCN.UK.2021-2.RLTS.T82348077A82348673.en .
  91. 1 2 Thong, V. D.; Thanh, H. T.; Soisook, P.; Bates, P. J. J.; Csorba, G. (2019). "Rhinolophus thomasi". IUCN Red List of Threatened Species . 2019: e.T19573A21990671. doi: 10.2305/IUCN.UK.2019-3.RLTS.T19573A21990671.en .
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  93. 1 2 Huang, J. C. -C. (2020). "Rhinolophus trifoliatus". IUCN Red List of Threatened Species . 2020: e.T19574A21990821. doi: 10.2305/IUCN.UK.2020-2.RLTS.T19574A21990821.en .
  94. 1 2 Zhou, Z.; Jiang, X. (2017). "Rhinolophus xinanzhongguoensis". IUCN Red List of Threatened Species . 2017: e.T82348701A82349975. doi: 10.2305/IUCN.UK.2017-2.RLTS.T82348701A82349975.en .
  95. 1 2 Monadjem, A. (2020). "Rhinolophus willardi". IUCN Red List of Threatened Species . 2020: e.T82346260A82347169. doi: 10.2305/IUCN.UK.2020-2.RLTS.T82346260A82347169.en .
  96. 1 2 Fukui, D.; Sano, A. (2020). "Rhinolophus perditus". IUCN Red List of Threatened Species . 2020: e.T85707170A85707174. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85707170A85707174.en .
  97. 1 2 Duya, M. R.; Alviola, P. A.; Sedlock, J.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Veluz, M. J.; Jakosalem, P. G. (2019). "Rhinolophus virgo". IUCN Red List of Threatened Species . 2019: e.T19575A21991148. doi: 10.2305/IUCN.UK.2019-3.RLTS.T19575A21991148.en .
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