List of hipposiderids

Commerson's leaf-nosed bat (Hipposideros commersoni)

Hipposideridae is one of the twenty families of bats in the mammal order Chiroptera, and part of the microbat suborder grouping. A member of this family is called a hipposiderid, or an Old World leaf-nosed bat. They are named for the shape of their nose-leaf. They are found in Africa, Asia, and Australia, primarily in forests, savannas, rocky areas, and caves, though some species can also be found in grasslands or wetlands. They range in size from the Malayan tailless leaf-nosed bat, at 3 cm (1 in) and no tail, to the striped leaf-nosed bat, at 13 cm (5 in) plus a 4 cm (2 in) tail. Like all bats, hipposiderids are capable of true and sustained flight, and have wing lengths ranging from multiple species with 3 cm (1 in), to the giant roundleaf bat at 13 cm (5 in). They are all insectivorous and primarily eat cicadas, cockroaches, termites, and beetles, though some species may eat trace amounts of fruit while consuming insects within. ^[1] Most hipposiderids do not have population estimates, but the ones that do range from 150 adult individuals to 10,000. The lesser great leaf-nosed bat, Makira roundleaf bat, Nicobar leaf-nosed bat, Pomona roundleaf bat, short-tailed roundleaf bat, Cox's roundleaf bat, and Sorensen's leaf-nosed bat are categorized as endangered species, and the Kolar leaf-nosed bat and Lamotte's roundleaf bat are categorized as critically endangered.

The 86 extant species of Hipposideridae are divided into seven genera; 70 of the species are in the Hipposideros genus of roundleaf bats. The other six genera are Anthops , or the flower-faced bat; Asellia , containing four trident bat species; Aselliscus , containing three trident bat species; Coelops , containing two tailless leaf-nosed bat species; Doryrhina , containing two roundleaf bat species; and Macronycteris , containing four leaf-nosed and roundleaf bat species. A few extinct prehistoric hipposiderid species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed. ^[2]

Conventions

IUCN Red List categories

Conservation status
EX	Extinct (0 species)
EW	Extinct in the wild (0 species)
CR	Critically Endangered (2 species)
EN	Endangered (7 species)
VU	Vulnerable (15 species)
NT	Near threatened (10 species)
LC	Least concern (41 species)
Other categories
DD	Data deficient (11 species)
NE	Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the hipposiderid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

The family Hipposideridae consists of seven genera: Anthops , with one species; Asellia , containing four species; Aselliscus , containing three species; Coelops , containing two species; Doryrhina , containing two species; Hipposideros , containing 70 species; and Macronycteris , containing four species.

Family Hipposideridae

• Genus Anthops (flower-faced bat): one species
• Genus Asellia (trident bats): four species
• Genus Aselliscus (trident bats): three species
• Genus Coelops (tailless leaf-nosed bats): two species
• Genus Doryrhina (roundleaf bats): two species
• Genus Hipposideros (roundleaf bats): 70 species
• Genus Macronycteris (leaf-nosed bats): four species

Hipposideridae [3]    Doryrhina Macronycteris Asellia Coelops Aselliscus Hipposideros Anthops

Hipposiderids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. ^[4]

Genus Anthops – Thomas, 1888 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Flower-faced bat	A. ornatus Thomas, 1888	Papua New Guinea and the Solomon Islands	Size: 4–7 cm (2–3 in) long, plus 0.3–1 cm (0.1–0.4 in) tail 4–6 cm (2 in) arm/wing length [5] Habitat: Forest and caves [6]	VU  Unknown [6]

Genus Asellia – Gray, 1838 – four species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Arabian trident bat	A. arabica Benda, Vallo, & Reiter, 2011	Oman and Yemen	Size: 4–6 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 4–5 cm (2 in) arm/wing length [7] Habitat: Savanna [8]	DD  Unknown [8]
Patrizi's trident leaf-nosed bat	A. patrizii Beaux, 1931	Eritrea, Ethiopia, and Saudi Arabia	Size: 4–5 cm (2 in) long, plus 1–2 cm (0.4–0.8 in) tail 3–5 cm (1–2 in) arm/wing length [7] Habitat: Shrubland, grassland, and caves [9]	LC  Unknown [9]
Somalian trident bat	A. italosomalica Beaux, 1936	Horn of Africa	Size: 4–6 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 4–5 cm (2 in) arm/wing length [7] Habitat: Savanna, shrubland, grassland, caves, and desert [10]	DD  Unknown [10]
Trident bat	A. tridens (Geoffroy, 1813)	Northern Africa and Western Asia	Size: 4–6 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 4–6 cm (2 in) arm/wing length [7] Habitat: Forest, savanna, shrubland, grassland, caves, and desert [11]	LC  Unknown [11]

Genus Aselliscus – Tate, 1941 – three species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Dong Bac's trident bat	A. dongbacanus Tu et al., 2015	Northern Vietnam	Size: 3–5 cm (1–2 in) long, plus 3–5 cm (1–2 in) tail about 4 cm (2 in) arm/wing length [5] Habitat: Forest and caves [12]	NT  Unknown [12]
Stoliczka's trident bat	A. stoliczkanus (Dobson, 1871)	Southeastern Asia	Size: 4–5 cm (2 in) long, plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [13]	LC  Unknown [13]
Temminck's trident bat	A. tricuspidatus (Temminck, 1835)	Northern Oceania	Size: 3–5 cm (1–2 in) long, plus 1–3 cm (0.4–1.2 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [14]	LC  Unknown [14]

Genus Coelops – Blyth, 1848 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
East Asian tailless leaf-nosed bat	C. frithii Blyth, 1848 Five subspecies C. f. bernsteini C. f. formosanus C. f. frithii C. f. inflatus C. f. sinicus	Southeastern Asia	Size: 3–5 cm (1–2 in) long, with no tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [15]	NT  Unknown [15]
Malayan tailless leaf-nosed bat	C. robinsoni Bonhote, 1908	Southeastern Asia	Size: 3–4 cm (1–2 in) long, with no tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Caves and forest [16]	VU  Unknown [16]

Genus Doryrhina – Peters, 1871 – two species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Greater roundleaf bat	D. camerunensis Eisentraut, 1956	Central Africa	Size: 9–10 cm (4 in) long, plus 2–5 cm (1–2 in) tail 7–8 cm (3–3 in) arm/wing length [5] Habitat: Forest [17]	DD  Unknown [17]
Cyclops roundleaf bat	D. cyclops Temminck, 1853	Central and western Africa	Size: 7–10 cm (3–4 in) long, plus 1–4 cm (0.4–1.6 in) tail 5–8 cm (2–3 in) arm/wing length [5] Habitat: Forest and savanna [18]	LC  Unknown [18]

Genus Hipposideros – Gray, 1831 – 70 species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Aba roundleaf bat	H. abae Allen, 1917	Central and western Africa	Size: 6–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail 5–7 cm (2–3 in) arm/wing length [19] Habitat: Savanna and rocky areas [20]	LC  Unknown [20]
Aellen's roundleaf bat	H. marisae Aellen, 1954	Western Africa	Size: 4–5 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 3–5 cm (1–2 in) arm/wing length [19] Habitat: Caves, rocky areas, and forest [21]	VU  Unknown [21]
Andersen's leaf-nosed bat	H. gentilis Andersen, 1918	Southern and southeastern Asia	Size: About 4 cm (2 in) long, plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [22]	LC  Unknown [22]
Arnhem leaf-nosed bat	H. inornatus McKean, 1970	Northern Australia	Size: 6–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail 6–8 cm (2–3 in) arm/wing length [19] Habitat: Forest, savanna, shrubland, and caves [23]	VU  Unknown [23]
Ashy roundleaf bat	H. cineraceus Blyth, 1853 Two subspecies H. c. cineraceus H. c. wrighti	Southern and southeastern Asia	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [24]	LC  Unknown [24]
Benito roundleaf bat	H. beatus K. Andersen, 1906 Two subspecies H. b. beatus H. b. maximus	Central and western Africa	Size: 4–5 cm (2 in) long, plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [19] Habitat: Forest and inland wetlands [25]	LC  Unknown [25]
Biak roundleaf bat	H. papua (Thomas & Doria, 1886)	Indonesia	Size: 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Forest and caves [26]	LC  Unknown [26]
Bicolored roundleaf bat	H. bicolor (Temminck, 1834)	Southeastern Asia	Size: Unknown length 4–5 cm (2 in) arm/wing length [19] Habitat: Forest and caves [27]	LC  Unknown [27]
Big-eared roundleaf bat	H. macrobullatus Tate, 1941	Indonesia	Size: Unknown length 4–5 cm (2 in) arm/wing length [19] Habitat: Forest and caves [28]	DD  Unknown [28]
Boeadi's roundleaf bat	H. boeadii Rossiter, Suyanto, Kingston, & Bates, 2007	Indonesia	Size: Unknown length, plus 1–2 cm (0.4–0.8 in) tail 4–5 cm (2 in) arm/wing length [19] Habitat: Forest [29]	DD  Unknown [29]
Borneo roundleaf bat	H. doriae (Peters, 1871)	Southeastern Asia	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [19] Habitat: Forest [30]	NT  Unknown [30]
Cantor's roundleaf bat	H. galeritus Cantor, 1846	Southern and southeastern Asia	Size: 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Rocky areas, caves, and forest [31]	LC  Unknown [31]
Cox's roundleaf bat	H. coxi Shelford, 1901	Western Borneo	Size: Unknown length 5–6 cm (2 in) arm/wing length [19] Habitat: Unknown [32]	EN  Unknown [32]
Crested roundleaf bat	H. inexpectatus Laurie & Hill, 1954	Indonesia	Size: Unknown length About 10 cm (4 in) arm/wing length [19] Habitat: Caves and unknown [33]	DD  Unknown [33]
Dayak roundleaf bat	H. dyacorum Thomas, 1902	Indonesia and Malaysia	Size: Unknown length 3–5 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [34]	LC  Unknown [34]
Diadem leaf-nosed bat	H. diadema (É. Geoffroy, 1813)	Southeastern Asia	Size: 6–10 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail 5–10 cm (2–4 in) arm/wing length [19] Habitat: Forest, savanna, and caves [35]	LC  Unknown [35]
Dusky leaf-nosed bat	H. ater Templeton, 1848 Six subspecies H. a. amboinensis H. a. aruensis H. a. ater H. a. gilberti H. a. nicobarulae H. a. saevus		Size: 3–5 cm (1–2 in) long, plus 1–3 cm (0.4–1.2 in) tail 1–5 cm (0.4–2.0 in) arm/wing length [19] Habitat: Forest, savanna, shrubland, and caves [36]	LC  Unknown [36]
Ethiopian large-eared roundleaf bat	H. megalotis (Heuglin, 1862)	Eastern Africa	Size: About 4 cm (2 in) long, plus 2–3 cm (1 in) tail 3–4 cm (1–2 in) arm/wing length [19] Habitat: Savanna and shrubland [37]	LC  Unknown [37]
Fawn leaf-nosed bat	H. cervinus (Gould, 1854) Four subspecies H. c. batchianus H. c. cervinus H. c. labuanensis H. c. misoriensis	Southeastern Asia	Size: 5–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Forest and caves [38]	LC  Unknown [38]
Fierce roundleaf bat	H. dinops K. Andersen, 1905	Northeastern Oceania	Size: 8–11 cm (3–4 in) long, plus 5–7 cm (2–3 in) tail 8–10 cm (3–4 in) arm/wing length [19] Habitat: Forest and caves [39]	VU  6,000 [39]
Fly River roundleaf bat	H. muscinus (Thomas & Doria, 1886)	New Guinea	Size: 4–6 cm (2 in) long, plus about 2 cm (1 in) tail 4–5 cm (2 in) arm/wing length [5] Habitat: Forest [40]	LC  Unknown [40]
Fulvus roundleaf bat	H. fulvus Gray, 1838	Southern Asia	Size: 4–5 cm (2 in) long, plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [41]	LC  Unknown [41]
Grand roundleaf bat	H. grandis Allen, 1936	Southeastern Asia	Size: 6–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail 5–7 cm (2–3 in) arm/wing length [19] Habitat: Caves and unknown [42]	LC  Unknown [42]
Great roundleaf bat	H. armiger (Hodgson, 1835) Four subspecies H. a. armiger H. a. fujianensis H. a. terasensis H. a. tranninhensis	Eastern and southeastern Asia	Size: 8–11 cm (3–4 in) long, plus 4–7 cm (2–3 in) tail 8–11 cm (3–4 in) arm/wing length [19] Habitat: Forest and caves [43]	LC  Unknown [43]
Griffin's leaf-nosed bat	H. griffini Thong et al., 2012	Vietnam	Size: Unknown length 8–9 cm (3–4 in) arm/wing length [19] Habitat: Forest and caves [44]	NT  Unknown [44]
Ha Long leaf-nosed bat	H. alongensis Bourret, 1942	Vietnam	Size: Unknown length 6–8 cm (2–3 in) arm/wing length [19] Habitat: Forest and caves [45]	VU  9,000 [45]
Hill's roundleaf bat	H. edwardshilli Flannery & Colgan, 1993	Northern Papua New Guinea	Size: About 5 cm (2 in) long, plus 1–2 cm (0.4–0.8 in) tail 4–6 cm (2 in) arm/wing length [5] Habitat: Forest and caves [46]	VU  Unknown [46]
House-dwelling leaf-nosed bat	H. einnaythu Douangboubpha et al., 2011	Myanmar	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [19] Habitat: Unknown [47]	DD  Unknown [47]
Indian roundleaf bat	H. lankadiva Kelaart, 1850	Southern Asia	Size: Unknown length, plus 3–6 cm (1–2 in) tail 7–10 cm (3–4 in) arm/wing length [19] Habitat: Forest, shrubland, and caves [48]	LC  Unknown [48]
Intermediate roundleaf bat	H. larvatus (Horsfield, 1823)	Southeastern Asia	Size: 5–8 cm (2–3 in) long, plus 3–4 cm (1–2 in) tail 5–7 cm (2–3 in) arm/wing length [19] Habitat: Forest and caves [49]	LC  Unknown [49]
Jones's roundleaf bat	H. jonesi Hayman, 1947	Western Africa	Size: 5–6 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 4–5 cm (2 in) arm/wing length [19] Habitat: Forest, savanna, grassland, rocky areas, and caves [50]	NT  Unknown [50]
Khajuria's leaf-nosed bat	H. durgadasi Khajuria, 1970	Central India	Size: 3–5 cm (1–2 in) long, plus 2–3 cm (1 in) tail 3–4 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [51]	VU  Unknown [51]
Kolar leaf-nosed bat	H. hypophyllus Kock & Bhat, 1994	Southern India	Size: 4–5 cm (2 in) long, plus 2–3 cm (1 in) tail 3–5 cm (1–2 in) arm/wing length [19] Habitat: Shrubland and caves [52]	CR  150–200 [52]
Lamotte's roundleaf bat	H. lamottei (Brosset, 1984)	Western Africa	Size: 9–11 cm (4 in) long, plus 3–5 cm (1–2 in) tail 5–6 cm (2 in) arm/wing length [19] Habitat: Forest, grassland, and caves [53]	CR  Unknown [53]
Laotian leaf-nosed bat	H. rotalis Francis, Kock, & Habersetzer, 1999	Laos	Size: Unknown length, plus 3–4 cm (1–2 in) tail 4–5 cm (2 in) arm/wing length [19] Habitat: Forest [54]	LC  Unknown [54]
Large Asian roundleaf bat	H. lekaguli Thonglongya & Hill, 1974	Southeastern Asia	Size: Unknown length 6–8 cm (2–3 in) arm/wing length [19] Habitat: Forest and caves [55]	NT  Unknown [55]
Large Mindanao roundleaf bat	H. coronatus Peters, 1871	Philippines	Size: 8–9 cm (3–4 in) long, plus 2–3 cm (1 in) tail About 5 cm (2 in) arm/wing length [19] Habitat: Forest and caves [56]	DD  Unknown [56]
Lesser great leaf-nosed bat	H. turpis Bangs, 1901	Japan	Size: 6–9 cm (2–4 in) long, plus 4–6 cm (2 in) tail 6–8 cm (2–3 in) arm/wing length [19] Habitat: Forest, inland wetlands, and caves [57]	EN  Unknown [57]
Maduran leaf-nosed bat	H. madurae Kitchener & Maryanto, 1993	Indonesia	Size: Unknown length, plus 2–4 cm (1–2 in) tail 5–6 cm (2 in) arm/wing length [19] Habitat: Forest and caves [58]	NT  Unknown [58]
Maggie Taylor's roundleaf bat	H. maggietaylorae Smith & Hill, 1981 Two subspecies H. m. erroris H. m. maggietaylorae	New Guinea and nearby islands	Size: 5–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail 5–7 cm (2–3 in) arm/wing length [19] Habitat: Caves, shrubland, and forest [59]	LC  Unknown [59]
Maghreb Leaf-nosed Bat	H. tephrus Cabrera, 1906	Morocco, Yemen, and Senegal	Size: 4–5 cm (2 in) long, plus 2–4 cm (1–2 in) tail Unknown arm/wing length [19] Habitat: Forest and savanna [60]	LC  Unknown [60]
Makira roundleaf bat	H. demissus K. Andersen, 1909	Solomon Islands	Size: 6–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail About 7 cm (3 in) arm/wing length [19] Habitat: Forest and caves [61]	EN  Unknown [61]
Malayan roundleaf bat	H. nequam K. Andersen, 1918	Malaysia	Size: Unknown length About 5 cm (2 in) arm/wing length [19] Habitat: Unknown [62]	DD  Unknown [62]
Nicobar leaf-nosed bat	H. nicobarulae Miller, 1902	Nicobar Islands	Size: Unknown length 3–5 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [63]	EN  Unknown [63]
Noack's roundleaf bat	H. ruber Noack, 1893 Two subspecies H. r. guineensis H. r. ruber	Sub-Saharan Africa	Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Forest, savanna, and caves [64]	LC  Unknown [64]
Northern leaf-nosed bat	H. stenotis Thomas, 1913	Northern Australia	Size: 4–5 cm (2 in) long, plus 2–3 cm (1 in) tail 4–5 cm (2 in) arm/wing length [5] Habitat: Savanna, rocky areas, and caves [65]	VU  5,000 [65]
Orbiculus leaf-nosed bat	H. orbiculus Francis, Kock, & Habersetzer, 1999	Sumatra island in Indonesia and Malaysia	Size: Unknown length, plus 2–4 cm (1–2 in) tail 4–5 cm (2 in) arm/wing length [19] Habitat: Forest [66]	VU  Unknown [66]
Peleng leaf-nosed bat	H. pelingensis Shamel, 1940	Sulawesi island in Indonesia	Size: Unknown length 9–10 cm (4 in) arm/wing length [19] Habitat: Forest and caves [67]	NT  Unknown [67]
Pendlebury's roundleaf bat	H. pendleburyi Chasen, 1936	Thailand	Size: Unknown length, plus 4–7 cm (2–3 in) tail 7–9 cm (3–4 in) arm/wing length [19] Habitat: Forest and caves [68]	VU  4,700 [68]
Philippine forest roundleaf bat	H. obscurus (Peters, 1861)	Philippines	Size: 5–6 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 4–5 cm (2 in) arm/wing length [19] Habitat: Forest and caves [69]	LC  Unknown [69]
Philippine pygmy roundleaf bat	H. pygmaeus (Waterhouse, 1843)	Philippines	Size: 5–8 cm (2–3 in) long, plus 1–3 cm (0.4–1.2 in) tail 3–5 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [70]	LC  Unknown [70]
Phou Khao Khouay leaf-nosed bat	H. khaokhouayensis Guillén-Servent & Francis, 2009	Laos and Vietnam	Size: Unknown length 4–5 cm (2 in) arm/wing length [19] Habitat: Forest [71]	VU  8,000–10,000 [71]
Pomona roundleaf bat	H. pomona K. Andersen, 1918	India	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [72]	EN  Unknown [72]
Pratt's roundleaf bat	H. pratti Thomas, 1891	Eastern Asia	Size: 9–11 cm (4 in) long, plus 5–7 cm (2–3 in) tail 7–9 cm (3–4 in) arm/wing length [19] Habitat: Caves [73]	LC  Unknown [73]
Ridley's leaf-nosed bat	H. ridleyi Robinson & Kloss, 1911	Southeastern Asia	Size: Unknown length, plus 2–3 cm (1 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Forest [74]	VU  Unknown [74]
Schneider's leaf-nosed bat	H. speoris (Schneider, 1800)	India	Size: 4–7 cm (2–3 in) long 4–6 cm (2 in) arm/wing length [19] Habitat: Forest, shrubland, rocky areas, and caves [75]	LC  Unknown [75]
Semon's leaf-nosed bat	H. semoni Matschie, 1903	Northeastern Australia	Size: 4–5 cm (2 in) long, plus 2–3 cm (1 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest, savanna, and caves [76]	LC  Unknown [76]
Shield-faced roundleaf bat	H. lylei Thomas, 1913	Southeastern Asia	Size: 7–10 cm (3–4 in) long, plus 4–6 cm (2 in) tail 7–9 cm (3–4 in) arm/wing length [19] Habitat: Forest and caves [77]	LC  Unknown [77]
Shield-nosed leaf-nosed bat	H. scutinares Robinson, Jenkins, Francis, & Fulford, 2003	Laos and Vietnam	Size: Unknown length, plus 5–6 cm (2 in) tail 7–9 cm (3–4 in) arm/wing length [19] Habitat: Forest and caves [78]	VU  8,000–10,000 [78]
Short-headed roundleaf bat	H. breviceps Tate, 1941	Indonesia	Size: About 4 cm (2 in) long, plus about 2 cm (1 in) tail About 4 cm (2 in) arm/wing length [19] Habitat: Forest and caves [79]	DD  Unknown [79]
Short-tailed roundleaf bat	H. curtus Allen, 1921	Cameroon and Equatorial Guinea	Size: 5–6 cm (2 in) long, plus 1–3 cm (0.4–1.2 in) tail 4–5 cm (2 in) arm/wing length [19] Habitat: Forest and caves [80]	EN  Unknown [80]
Sooty roundleaf bat	H. fuliginosus (Temminck, 1853)	Central and western Africa	Size: 8–10 cm (3–4 in) long, plus 2–4 cm (1–2 in) tail 5–6 cm (2 in) arm/wing length [19] Habitat: Forest [81]	LC  Unknown [81]
Sorensen's leaf-nosed bat	H. sorenseni Kitchener, 1993	Indonesia	Size: 5–6 cm (2 in) long, plus 3–4 cm (1–2 in) tail 5–6 cm (2 in) arm/wing length [19] Habitat: Caves [82]	EN  Unknown [82]
Spurred roundleaf bat	H. calcaratus (Dobson, 1877) Two subspecies H. c. calcaratus H. c. cupidus	Papua New Guinea and the Solomon Islands	Size: 4–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Forest and caves [83]	LC  Unknown [83]
Sumba roundleaf bat	H. sumbae (Oei, 1960)	Indonesia and East Timor	Size: Unknown length, plus 2–4 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Caves [84]	LC  Unknown [84]
Sundevall's roundleaf bat	H. caffer (Sundevall, 1846) Three subspecies H. c. angolensis H. c. caffer H. c. nanus	Africa and southern Arabian Peninsula	Size: 4–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail 4–6 cm (2 in) arm/wing length [19] Habitat: Forest, savanna, shrubland, inland wetlands, and caves [85]	LC  Unknown [85]
Telefomin roundleaf bat	H. corynophyllus Hill, 1985	New Guinea	Size: 5–7 cm (2–3 in) long, plus 0.5–2 cm (0.2–0.8 in) tail 4–6 cm (2 in) arm/wing length [5] Habitat: Forest and caves [86]	LC  Unknown [86]
Thailand roundleaf bat	H. halophyllus Hill & Yenbutra, 1984	Thailand	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [19] Habitat: Forest and caves [87]	VU  2,500–10,000 [87]
Timor roundleaf bat	H. crumeniferus Lesueur & Petit, 1807	Indonesia	Size: Unknown [19] Habitat: Forest [88]	DD  Unknown [88]
Wollaston's roundleaf bat	H. wollastoni Thomas, 1913 Three subspecies H. w. fasensis H. w. parnabyi H. w. wollastoni	New Guinea	Size: 4–5 cm (2 in) long, plus about 3 cm (1 in) tail About 4 cm (2 in) arm/wing length [5] Habitat: Forest and caves [89]	LC  Unknown [89]

Genus Macronycteris – Gray, 1866 – four species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Commerson's roundleaf bat	M. commersonii Geoffroy, 1813	Madagascar	Size: 10–11 cm (4 in) long, plus 3–5 cm (1–2 in) tail 8–11 cm (3–4 in) arm/wing length [5] Habitat: Forest and caves [90]	NT  Unknown [90]
Giant roundleaf bat	M. gigas Wagner, 1845	Central and western Africa	Size: 9–12 cm (4–5 in) long, plus 2–5 cm (1–2 in) tail 9–13 cm (4–5 in) arm/wing length [5] Habitat: Forest, savanna, rocky areas, and caves [91]	LC  Unknown [91]
São Tomé leaf-nosed bat	M. thomensis Bocage, 1891	São Tomé Island	Size: 10–11 cm (4 in) long, plus 2–3 cm (1 in) tail [5] Habitat: Forest and caves [92]	LC  Unknown [92]
Striped leaf-nosed bat	M. vittata (Peters, 1852)	Sub-Saharan Africa	Size: 9–13 cm (4–5 in) long, plus 2–4 cm (1–2 in) tail 8–11 cm (3–4 in) arm/wing length [5] Habitat: Forest, savanna, and caves [93]	NT  Unknown [93]

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53. Monadjem, A.; Cooper-Bohannon, R. (2020). "Hipposideros lamottei". IUCN Red List of Threatened Species . 2020: e.T10141A22091938. doi: 10.2305/IUCN.UK.2020-2.RLTS.T10141A22091938.en .
54. Francis, C. M. (2019). "Hipposideros rotalis". IUCN Red List of Threatened Species . 2019: e.T136477A21985931. doi: 10.2305/IUCN.UK.2019-2.RLTS.T136477A21985931.en .
55. Csorba, G.; Bumrungsri, S.; Francis, C.; Bates, P.; Gumal, M.; Kingston, T.; Soisook, P. (2019). "Hipposideros lekaguli". IUCN Red List of Threatened Species . 2019: e.T10144A22091565. doi: 10.2305/IUCN.UK.2019-3.RLTS.T10144A22091565.en .
56. Phelps, K.; Heaney, L.; Sedlock, J. (2016). "Hipposideros coronatus". IUCN Red List of Threatened Species . 2016: e.T10121A22097259. doi: 10.2305/IUCN.UK.2016-3.RLTS.T10121A22097259.en .
57. Fukui, D.; Sano, A. (2019). "Hipposideros turpis". IUCN Red List of Threatened Species . 2019: e.T80224148A22099660. doi: 10.2305/IUCN.UK.2019-3.RLTS.T80224148A22099660.en .
58. Santiago, K.; Wortham, G.; Waldien, D. L. (2021). "Hipposideros madurae". IUCN Red List of Threatened Species . 2021: e.T10147A22100964. doi: 10.2305/IUCN.UK.2021-2.RLTS.T10147A22100964.en .
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60. Monadjem, A.; Shapiro, J. (2017). "Hipposideros tephrus". IUCN Red List of Threatened Species . 2017: e.T85646524A85646528. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85646524A85646528.en .
61. Pennay, M.; Leary, T. (2020). "Hipposideros demissus". IUCN Red List of Threatened Species . 2020: e.T10127A22095744. doi: 10.2305/IUCN.UK.2020-2.RLTS.T10127A22095744.en .
62. Senawi, J.; Bates, P.; Francis, C. (2016). "Hipposideros nequam". IUCN Red List of Threatened Species . 2016: e.T10152A22101545. doi: 10.2305/IUCN.UK.2016-2.RLTS.T10152A22101545.en .
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66. Francis, C; Bates, P.; Kingston, T.; Senawi, J. (2016). "Hipposideros orbiculus". IUCN Red List of Threatened Species . 2016: e.T136192A22008477. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136192A22008477.en .
67. Wiantoro, S.; Waldien, D. L. (2021). "Hipposideros pelingensis". IUCN Red List of Threatened Species . 2021: e.T136600A21996457. doi: 10.2305/IUCN.UK.2021-2.RLTS.T136600A21996457.en .
68. Soisook, P. (2019). "Hipposideros pendlebury". IUCN Red List of Threatened Species . 2019: e.T80224655A95642195. doi: 10.2305/IUCN.UK.2019-3.RLTS.T80224655A95642195.en .
69. Alviola, P. A.; Sedlock, J.; Alvarez, J.; Fidelino, J.; Pedregosa, M.; Jakosalem, P. G.; Tanalgo, K. (2019). "Hipposideros obscurus". IUCN Red List of Threatened Species . 2019: e.T10153A22101961. doi: 10.2305/IUCN.UK.2019-3.RLTS.T10153A22101961.en .
70. Sedlock, J.; Alviola, P. A.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Pedregosa, M.; Veluz, M. J.; Jakosalem, P. G.; Tanalgo, K. (2019). "Hipposideros pygmaeus". IUCN Red List of Threatened Species . 2019: e.T10156A22102078. doi: 10.2305/IUCN.UK.2019-3.RLTS.T10156A22102078.en .
71. Douangboubpha, B. (2020) [amended version of 2019 assessment]. "Hipposideros khaokhouayensis". IUCN Red List of Threatened Species . 2020: e.T136819A166602959. doi: 10.2305/IUCN.UK.2020-1.RLTS.T136819A166602959.en .
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73. Jiang, T. L.; Feng, J. (2019). "Hipposideros pratti". IUCN Red List of Threatened Species . 2019: e.T10155A22102257. doi: 10.2305/IUCN.UK.2019-3.RLTS.T10155A22102257.en .
74. Khan, F. A. A.; Rajasegaran, P.; Rosli, Q. (2020). "Hipposideros ridleyi". IUCN Red List of Threatened Species . 2020: e.T10108A22098446. doi: 10.2305/IUCN.UK.2020-2.RLTS.T10108A22098446.en .
75. Srinivasulu, B.; Srinivasulu, C. (2019). "Hipposideros speoris". IUCN Red List of Threatened Species . 2019: e.T10162A22099260. doi: 10.2305/IUCN.UK.2019-2.RLTS.T10162A22099260.en .
76. Armstrong, K. N.; Aplin, K. (2021) [amended version of 2017 assessment]. "Hipposideros semoni". IUCN Red List of Threatened Species . 2021: e.T10160A209537564. doi: 10.2305/IUCN.UK.2021-3.RLTS.T10160A209537564.en .
77. Senawi, J.; Bates, P.; Bumrungsri, S.; Francis, C. (2019). "Hipposideros lylei". IUCN Red List of Threatened Species . 2019: e.T10145A22100391. doi: 10.2305/IUCN.UK.2019-2.RLTS.T10145A22100391.en .
78. Furey, N.; Görföl, T.; Francis, C.; Bates, P.; Csorba, G.; Tu, V. (2019). "Hipposideros scutinares". IUCN Red List of Threatened Species . 2019: e.T136586A22000133. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136586A22000133.en .
79. Huang, J. C.-C.; Sigit W. (2016). "Hipposideros breviceps". IUCN Red List of Threatened Species . 2016: e.T10114A22094935. doi: 10.2305/IUCN.UK.2016-2.RLTS.T10114A22094935.en .
80. Tanshi, I. (2020). "Hipposideros curtus". IUCN Red List of Threatened Species . 2020: e.T10125A22096364. doi: 10.2305/IUCN.UK.2020-2.RLTS.T10125A22096364.en .
81. Monadjem, A.; Juste, J.; Bergmans, W.; Mickleburgh, S.; Hutson, A. M.; Fahr, J. (2017). "Hipposideros fuliginosus". IUCN Red List of Threatened Species . 2017: e.T10134A22090466. doi: 10.2305/IUCN.UK.2017-2.RLTS.T10134A22090466.en .
82. Waldien, D. L.; Wiantoro, S. (2021). "Hipposideros sorenseni". IUCN Red List of Threatened Species . 2021: e.T10161A22099115. doi: 10.2305/IUCN.UK.2021-2.RLTS.T10161A22099115.en .
83. Armstrong, K. N.; Wiantoro, S.; Lavery, T. H. (2021). "Hipposideros calcaratus". IUCN Red List of Threatened Species . 2021: e.T10116A22094185. doi: 10.2305/IUCN.UK.2021-2.RLTS.T10116A22094185.en .
84. Santiago, K.; Wortham, G.; Gill, J.; Waldien, D. L. (2021). "Hipposideros sumbae". IUCN Red List of Threatened Species . 2021: e.T10164A22099540. doi: 10.2305/IUCN.UK.2021-3.RLTS.T10164A22099540.en .
85. Richards, L. R.; Cooper-Bohannon, R.; Kock, D.; Amr, Z. S. S.; Mickleburgh, S.; Hutson, A. M.; Bergmans, W.; Aulagnier, S. (2020). "Hipposideros caffer". IUCN Red List of Threatened Species . 2020: e.T80459007A22094271. doi: 10.2305/IUCN.UK.2020-2.RLTS.T80459007A22094271.en .
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