List of ctenomyids

Last updated

Silky tuco-tuco (Ctenomys sericeus) Ctenomys sericeus 216930243.jpg
Silky tuco-tuco (Ctenomys sericeus)

Ctenomyidae is a family of fossorial mammals in the order Rodentia and part of the Caviomorpha parvorder. Members of this family are called ctenomyids or tuco-tucos. They are found in southern South America, primarily in shrublands and grasslands, though some species can be found in forests, savannas, deserts, wetlands, and coastal areas. They range in size from the white-toothed tuco-tuco, at 11 cm (4 in) plus a 7 cm (3 in) tail, to Conover's tuco-tuco, at 33 cm (13 in) plus a 11 cm (4 in) tail. Ctenomyids are herbivores and eat a wide variety of vegetation. [1] No ctenomyids have population estimates, but ten species are categorized as an endangered species, and three—Reig's tuco-tuco, Roig's tuco-tuco, and the social tuco-tuco—are categorized as critically endangered.

Contents

The 59 extant species of Ctenomyidae are all contained in a single genus, Ctenomys. A few extinct prehistoric ctenomyid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed. [2]

Conventions

IUCN Red List categories
Conservation status
 EX  Extinct (0 species)
 EW  Extinct in the wild (0 species)
 CR  Critically endangered (3 species)
 EN  Endangered (10 species)
 VU  Vulnerable (1 species)
 NT  Near threatened (5 species)
 LC  Least concern (17 species)
Other categories
 DD  Data deficient (23 species)
 NE  Not evaluated (0 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the ctenomyid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

Ctenomyidae is a family consisting of 59 species in a single genus, Ctenomys.

Ctenomyids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. [3]

Genus Ctenomys Blainville, 1826 – 59 species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Argentine tuco-tuco


C. argentinus
Berry & Contreras, 1982
Northern Argentina Size: 17–18 cm (7 in) long, plus 7–9 cm (3–4 in) tail [4]

Habitat: Shrubland and grassland [5]
 NT 


Unknown Decrease2.svg [5]

Azara's tuco-tuco

Ctenomys azarae 118549264.jpg

C. azarae
Thomas, 1903
Central ArgentinaSize: 16–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail [6]

Habitat: Shrubland [7]
 EN 


Unknown Decrease2.svg [7]

Berg's tuco-tuco


C. bergi
Thomas, 1902
Central ArgentinaSize: 13–17 cm (5–7 in) long, plus about 7 cm (3 in) tail [8]

Habitat: Savanna and grassland [9]
 EN 


Unknown Decrease2.svg [9]

Bolivian tuco-tuco


C. boliviensis
Waterhouse, 1848

Two subspecies
  • C. b. boliviensis
  • C. b. nattereri
Bolivia and western BrazilSize: 18–28 cm (7–11 in) long, plus 7–10 cm (3–4 in) tail [10]

Habitat: Savanna [11]
 LC 


Unknown Steady2.svg [11]

Bonetto's tuco-tuco


C. bonettoi
Berry & Contreras, 1982
Northern ArgentinaSize: 17–19 cm (7 in) long, plus 6–8 cm (2–3 in) tail [8]

Habitat: Shrubland [12]
 EN 


Unknown Decrease2.svg [12]

Brazilian tuco-tuco

Ctenomys brasiliensis 1847 - cropping.jpg

C. brasiliensis
Blainville, 1826
Uruguay Size: Unknown [6]

Habitat: Unknown [13]
 DD 


Unknown Blue question mark (italic).svg [13]

Catamarca tuco-tuco


C. knighti
Thomas, 1919
Northern ArgentinaSize: About 20 cm (8 in) long, plus about 8 cm (3 in) tail [8]

Habitat: Grassland [14]
 DD 


Unknown Blue question mark (italic).svg [14]

Chacoan tuco-tuco


C. dorsalis
Thomas, 1900
Paraguay and BoliviaSize: About 16 cm (6 in) long, plus about 5 cm (2 in) tail [15]

Habitat: Savanna [16]
 DD 


Unknown Blue question mark (italic).svg [16]

Colburn's tuco-tuco

Ctenomys colburni 204747802.jpg

C. colburni
Allen, 1903
Southern ArgentinaSize: Unknown [4]

Habitat: Grassland [17]
 DD 


Unknown Blue question mark (italic).svg [17]

Collared tuco-tuco

Ctenomys torquatus.png

C. torquatus
Lichtenstein, 1830
Uruguay and southern BrazilSize: 15–23 cm (6–9 in) long, plus 5–9 cm (2–4 in) tail [15]

Habitat: Forest and shrubland [18]
 LC 


Unknown Blue question mark (italic).svg [18]

Conover's tuco-tuco

Ctenomys conoveri 31638880.jpg

C. conoveri
Osgood, 1946
Paraguay and BoliviaSize: 24–33 cm (9–13 in) long, plus 9–14 cm (4–6 in) tail [19]

Habitat: Shrubland and grassland [20]
 LC 


Unknown Steady2.svg [20]

Coyhaique tuco-tuco


C. coyhaiquensis
Kelt & Gallardo, 1994
Southern Argentina and southern Chile Size: Unknown length, plus 5–9 cm (2–4 in) tail [4]

Habitat: Shrubland [21]
 DD 


Unknown Blue question mark (italic).svg [21]

D'Orbigny's tuco-tuco


C. dorbignyi
Contreras & Contreras, 1984
Northern Argentina
Ctenomys dorbignyi distribution.svg
Size: 19–22 cm (7–9 in) long, plus 8–10 cm (3–4 in) tail [6]

Habitat: Shrubland and grassland [22]
 NT 


Unknown Decrease2.svg [22]

Emily's tuco-tuco

Fam.Ctenomyidae, Ctenomys emilianus -Tuco-tuco neuquino, Salto El Agrio, 2015 03 10 IMG 8452.jpg

C. emilianus
St. Leger, 1926
Western ArgentinaSize: About 21 cm (8 in) long, plus about 9 cm (4 in) tail [10]

Habitat: Desert [23]
 LC 


Unknown Decrease2.svg [23]

Famatina tuco-tuco


C. famosus
Thomas, 1920
Northwestern ArgentinaSize: About 16 cm (6 in) long, plus about 7 cm (3 in) tail [15]

Habitat: Unknown [24]
 DD 


Unknown Blue question mark (italic).svg [24]

Flamarion's tuco-tuco

Ctenomys flamarioni.jpg

C. flamarioni
Travi, 1981
Uruguay and southern Brazil
Ctenomys flamarioni distribution.png
Size: About 25 cm (10 in) long, plus about 7 cm (3 in) tail [6]

Habitat: Coastal marine [25]
 EN 


Unknown Decrease2.svg [25]

Foch's tuco-tuco


C. fochi
Thomas, 1919
Northern ArgentinaSize: About 16 cm (6 in) long, plus about 8 cm (3 in) tail [8]

Habitat: Unknown [26]
 DD 


Unknown Blue question mark (italic).svg [26]

Furtive tuco-tuco


C. occultus
Thomas, 1920
Northern ArgentinaSize: 13–16 cm (5–6 in) long, plus tail [4]

Habitat: Unknown [27]
 EN 


Unknown Decrease2.svg [27]

Goodfellow's tuco-tuco


C. goodfellowi
Thomas, 1921
BoliviaSize: 18–28 cm (7–11 in) long, plus 7–10 cm (3–4 in) tail [10]

Habitat: Forest [28]
 LC 


Unknown Blue question mark (italic).svg [28]

Goya tuco-tuco


C. perrensi
Thomas, 1898
Northeastern ArgentinaSize: About 20 cm (8 in) long, plus about 7 cm (3 in) tail [6]

Habitat: Unknown [29]
 LC 


Unknown Steady2.svg [29]

Haig's tuco-tuco

Haig's Tuco-tuco imported from iNaturalist photo 276685030 on 24 December 2024 (cropped).jpg

C. haigi
Thomas, 1917

Two subspecies
  • C. h. haigi
  • C. h. lentulus
Western ArgentinaSize: 17–20 cm (7–8 in) long, plus 8–9 cm (3–4 in) tail [4]

Habitat: Forest, grassland, and inland wetlands [30]
 LC 


Unknown Blue question mark (italic).svg [30]

Highland tuco-tuco

Highland Tuco-tuco, Abra Pampa, Jujuy, AR imported from iNaturalist photo 257010861.jpg

C. opimus
Wagner, 1848

Three subspecies
  • C. o. luteolus
  • C. o. nigriceps
  • C. o. opimus
Bolivia and northwestern ArgentinaSize: 19–25 cm (7–10 in) long, plus 7–10 cm (3–4 in) tail [15]

Habitat: Grassland [31]
 LC 


Unknown Steady2.svg [31]

Ibicui tuco-tuco


C. ibicui
de Freitas, Fernandes, Fornel, Roratto, 2012
Southern BrazilSize: About 16 cm (6 in) long, plus about 8 cm (3 in) tail [15]

Habitat: Grassland [32]
 NT 


Unknown Blue question mark (italic).svg [32]

Jujuy tuco-tuco


C. juris
Thomas, 1920
Northern ArgentinaSize: About 18 cm (7 in) long, plus about 7 cm (3 in) tail [8]

Habitat: Unknown [33]
 DD 


Unknown Blue question mark (italic).svg [33]

Lago Blanco tuco-tuco


C. fodax
Thomas, 1910
Southern Argentina and southern ChileSize: About 26 cm (10 in) long, plus about 10 cm (4 in) tail [4]

Habitat: Grassland [34]
 DD 


Unknown Blue question mark (italic).svg [34]

Lami tuco-tuco

Ctenomys lami 242270267.jpg

C. lami
Freitas, 2001
Southern BrazilSize: 16–22 cm (6–9 in) long, plus 6–10 cm (2–4 in) tail [15]

Habitat: Coastal marine [35]
 VU 


Unknown Decrease2.svg [35]

Lewis's tuco-tuco

Ctenomys amguya tutu.jpg

C. lewisi
Thomas, 1926
Southern BoliviaSize: 20–22 cm (8–9 in) long, plus 6–8 cm (2–3 in) tail [4]

Habitat: Grassland [36]
 LC 


Unknown Steady2.svg [36]

Magellanic tuco-tuco

Ctenomys magellanicus fueginus, Tierra del Fuego, CL-MA, CL imported from iNaturalist photo 266393735.jpg

C. magellanicus
Bennett, 1836

Four subspecies
  • C. m. dicki
  • C. m. fueginus
  • C. m. magellanicus
  • C. m. osgoodi
Southern Argentina and southern ChileSize: Unknown [4]

Habitat: Grassland [37]
 LC 


Unknown Decrease2.svg [37]

Maule tuco-tuco

Maule Tuco-tuco imported from iNaturalist photo 363214670 on 8 June 2025.jpg

C. maulinus
Philippi, 1872

Two subspecies
  • C. m. brunneus
  • C. m. maulinus
Central ChileSize: Unknown [6]

Habitat: Grassland [38]
 LC 


Unknown Blue question mark (italic).svg [38]

Mendoza tuco-tuco


C. mendocinus
Philippi, 1869
Central ArgentinaSize: 16–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail [6]

Habitat: Shrubland, grassland, and inland wetlands [39]
 LC 


Unknown Blue question mark (italic).svg [39]

Mottled tuco-tuco


C. latro
Thomas, 1918
Northern ArgentinaSize: 16–18 cm (6–7 in) long, plus tail [4]

Habitat: Shrubland and grassland [40]
 EN 


Unknown Decrease2.svg [40]

Pearson's tuco-tuco

Ctenomys pearsoni imported from iNaturalist photo 63212110 on 23 June 2020.jpg

C. pearsoni
Lessa & Langguth, 1983
Southern UruguaySize: 17–20 cm (7–8 in) long, plus 7–9 cm (3–4 in) tail [6]

Habitat: Shrubland, grassland, and coastal marine [41]
 NT 


Unknown Decrease2.svg [41]

Peruvian tuco-tuco


C. peruanus
Sanborn & Pearson, 1947
Southern Peru Size: About 22 cm (9 in) long, plus about 9 cm (4 in) tail [8]

Habitat: Grassland [42]
 LC 


Unknown Steady2.svg [42]

Pilar tuco-tuco


C. pilarensis
Contreras, 1993
Northern Argentina and ParaguaySize: 16–21 cm (6–8 in) long, plus tail [4]

Habitat: Grassland [43]
 EN 


Unknown Decrease2.svg [43]

Porteous's tuco-tuco


C. porteousi
Thomas, 1916
Central ArgentinaSize: 16–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail [6]

Habitat: Grassland [44]
 NT 


Unknown Blue question mark (italic).svg [44]

Pundt's tuco-tuco


C. pundti
Nehring, 1900
Central ArgentinaSize: About 17 cm (7 in) long, plus about 4 cm (2 in) tail [6]

Habitat: Grassland [45]
 EN 


Unknown Decrease2.svg [45]

Puntilla tuco-tuco


C. coludo
Thomas, 1920
Northwestern ArgentinaSize: About 20 cm (8 in) long, plus about 10 cm (4 in) tail [15]

Habitat: Unknown [46]
 DD 


Unknown Blue question mark (italic).svg [46]

Reddish tuco-tuco


C. frater
Thomas, 1902

Two subspecies
  • C. f. frater
  • C. f. mordosus
Bolivia and northern ArgentinaSize: 17–20 cm (7–8 in) long, plus 5–9 cm (2–4 in) tail [4]

Habitat: Grassland [47]
 LC 


Unknown Decrease2.svg [47]

Reig's tuco-tuco

Ctenomys osvaldoreigi!.jpg

C. osvaldoreigi
Contreras, 1995
Central ArgentinaSize: 23–25 cm (9–10 in) long, plus tail [8]

Habitat: Grassland [48]
 CR 


Unknown Decrease2.svg [48]

Rio Negro tuco-tuco

Rio Negro Tuco-tuco imported from iNaturalist photo 267673422 on 8 June 2025.jpg

C. rionegrensis
Langguth & Abella, 1970
Eastern Argentina and UruguaySize: 16–19 cm (6–7 in) long, plus 6–9 cm (2–4 in) tail [6]

Habitat: Desert and coastal marine [49]
 EN 


Unknown Decrease2.svg [49]

Robust tuco-tuco

Ctenomys tuconax 38864099.jpg

C. tuconax
Thomas, 1925
Northern ArgentinaSize: Unknown [10]

Habitat: Savanna, shrubland, and grassland [50]
 DD 


Unknown Blue question mark (italic).svg [50]

Roig's tuco-tuco


C. roigi
Contreras, 1988
Northern ArgentinaSize: 17–20 cm (7–8 in) long, plus 7–10 cm (3–4 in) tail [6]

Habitat: Grassland, inland wetlands, and other [51]
 CR 


Unknown Decrease2.svg [51]

Salta tuco-tuco


C. saltarius
Thomas, 1912
Northern ArgentinaSize: About 20 cm (8 in) long, plus about 9 cm (4 in) tail [15]

Habitat: Desert [52]
 DD 


Unknown Blue question mark (italic).svg [52]

San Juan tuco-tuco


C. johannis
Thomas, 1921
Western ArgentinaSize: About 20 cm (8 in) long, plus about 10 cm (4 in) tail [15]

Habitat: Unknown [53]
 DD 


Unknown Blue question mark (italic).svg [53]

San Luis tuco-tuco


C. pontifex
Thomas, 1918
Central Argentina and central ChileSize: About 18 cm (7 in) long, plus about 8 cm (3 in) tail [8]

Habitat: Unknown [54]
 DD 


Unknown Blue question mark (italic).svg [54]

Scaglia's tuco-tuco


C. scagliai
Contreras, 1999
Northern ArgentinaSize: About 20 cm (8 in) long, plus about 9 cm (4 in) tail [15]

Habitat: Unknown [55]
 DD 


Unknown Blue question mark (italic).svg [55]

Sierra Tontal tuco-tuco


C. tulduco
Thomas, 1921
Western ArgentinaSize: About 19 cm (7 in) long, plus about 7 cm (3 in) tail [15]

Habitat: Unknown [56]
 DD 


Unknown Blue question mark (italic).svg [56]

Silky tuco-tuco

Ctenomys sericeus 216930243.jpg

C. sericeus
Allen, 1903
Southern Argentina and southern ChileSize: Unknown length, plus 5–9 cm (2–4 in) tail [4]

Habitat: Forest [57]
 DD 


Unknown Blue question mark (italic).svg [57]

Social tuco-tuco


C. sociabilis
Pearson & Christie, 1985
Central Argentina and central ChileSize: 16–25 cm (6–10 in) long, plus 6–8 cm (2–3 in) tail [10]

Habitat: Savanna and inland wetlands [58]
 CR 


Unknown Decrease2.svg [58]

Southern tuco-tuco

Dune Tuco-tuco imported from iNaturalist photo 57358841 on 11 October 2021.jpg

C. australis
Rusconi, 1934
Eastern ArgentinaSize: About 21 cm (8 in) long, plus 9–11 cm (4 in) tail [6]

Habitat: Coastal marine [59]
 EN 


Unknown Decrease2.svg [59]

Steinbach's tuco-tuco


C. steinbachi
Thomas, 1907
BoliviaSize: 23–25 cm (9–10 in) long, plus tail [10]

Habitat: Savanna [60]
 LC 


Unknown Steady2.svg [60]

Strong tuco-tuco


C. validus
Contreras, Roig, & Suzarte, 1977
Central ArgentinaSize: Unknown [15]

Habitat: Unknown [61]
 DD 


Unknown Blue question mark (italic).svg [61]

Talas tuco-tuco

Ctenomys talarum.jpg

C. talarum
Thomas, 1898

Four subspecies
  • C. t. antonii
  • C. t. occidentalis
  • C. t. recessus
  • C. t. talarum
Central and eastern ArgentinaSize: 15–18 cm (6–7 in) long, plus 5–8 cm (2–3 in) tail [6]

Habitat: Grassland and coastal marine [62]
 LC 


Unknown Blue question mark (italic).svg [62]

Tawny tuco-tuco


C. fulvus
Philippi, 1860

Two subspecies
  • C. f. fulvus
  • C. f. robustus
Western South AmericaSize: Unknown [15]

Habitat: Forest and desert [63]
 DD 


Unknown Blue question mark (italic).svg [63]

Tiny tuco-tuco

Tuco-tuco, Tiny Tuco Tuco (Ctenomys minutus).jpg

C. minutus
Nehring, 1887

Two subspecies
  • C. m. bicolor
  • C. m. minutus
Southern BrazilSize: About 7 cm (3 in) long, plus tail [15]

Habitat: Shrubland and grassland [64]
 DD 


Unknown Steady2.svg [64]

Tucuman tuco-tuco

Ctenomys tucumanus.jpg

C. tucumanus
Thomas, 1900
Northern ArgentinaSize: About 17 cm (7 in) long, plus about 7 cm (3 in) tail [4]

Habitat: Forest [65]
 DD 


Unknown Blue question mark (italic).svg [65]

Vipos tuco-tuco


C. viperinus
Thomas, 1926
Northern ArgentinaSize: About 21 cm (8 in) long, plus about 8 cm (3 in) tail [8]

Habitat: Unknown [66]
 DD 


Unknown Blue question mark (italic).svg [66]

White-toothed tuco-tuco


C. leucodon
Waterhouse, 1848
Western Bolivia and southern PeruSize: 11–18 cm (4–7 in) long, plus 7–9 cm (3–4 in) tail [4]

Habitat: Shrubland and grassland [67]
 LC 


Unknown Steady2.svg [67]

Yolanda's tuco-tuco

Ctenomys yolandae 237804760.jpg

C. yolandae
Contreras & Berry, 1984
Northern ArgentinaSize: Unknown [8]

Habitat: Unknown [68]
 DD 


Unknown Blue question mark (italic).svg [68]

References

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  2. "Fossilworks: Ctenomyidae". Paleobiology Database . University of Wisconsin–Madison. Archived from the original on December 4, 2024. Retrieved June 30, 2025.
  3. Wilson, Reeder, pp. 1560-1569
  4. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Chernasky; Motis; Burgin, p. 363
  5. 1 2 Bidau, C. J. (2019). "Ctenomys argentinus". IUCN Red List of Threatened Species . 2019: e.T5795A22191874. doi: 10.2305/IUCN.UK.2019-3.RLTS.T5795A22191874.en .
  6. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Chernasky; Motis; Burgin, p. 362
  7. 1 2 Bidau, C. J. (2019). "Ctenomys azarae". IUCN Red List of Threatened Species . 2019: e.T5797A22192079. doi: 10.2305/IUCN.UK.2019-3.RLTS.T5797A22192079.en .
  8. 1 2 3 4 5 6 7 8 9 10 Chernasky; Motis; Burgin, p. 364
  9. 1 2 Bidau, C. J. (2019). "Ctenomys bergi". IUCN Red List of Threatened Species . 2019: e.T136687A22196510. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136687A22196510.en .
  10. 1 2 3 4 5 6 Chernasky; Motis; Burgin, p. 360
  11. 1 2 Dunnum, J.; Bernal, N. (2017) [errata version of 2016 assessment]. "Ctenomys boliviensis". IUCN Red List of Threatened Species . 2016: e.T5798A115079917. doi: 10.2305/IUCN.UK.2016-3.RLTS.T5798A22192171.en .
  12. 1 2 Bidau, C. J. (2018). "Ctenomys bonettoi". IUCN Red List of Threatened Species . 2018: e.T5799A22192271. doi: 10.2305/IUCN.UK.2018-1.RLTS.T5799A22192271.en .
  13. 1 2 Ojeda, R. (2019). "Ctenomys brasiliensis". IUCN Red List of Threatened Species . 2019: e.T5800A22192465. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5800A22192465.en .
  14. 1 2 Ojeda, R. (2019). "Ctenomys knighti". IUCN Red List of Threatened Species . 2019: e.T5808A22193333. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5808A22193333.en .
  15. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 Chernasky; Motis; Burgin, p. 361
  16. 1 2 Ojeda, R. (2019). "Ctenomys dorsalis". IUCN Red List of Threatened Species . 2019: e.T5803A22192708. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5803A22192708.en .
  17. 1 2 Bidau, C. J. (2019). "Ctenomys colburni". IUCN Red List of Threatened Species . 2019: e.T5801A22192538. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5801A22192538.en .
  18. 1 2 Bidau, C. (2017) [errata version of 2016 assessment]. "Ctenomys torquatus". IUCN Red List of Threatened Species . 2016: e.T5829A115080157. doi: 10.2305/IUCN.UK.2016-3.RLTS.T5829A78319489.en .
  19. Brashear, Katie (2004). "Ctenomys conoveri". Animal Diversity Web . University of Michigan . Retrieved June 30, 2025.
  20. 1 2 Dunnum, J.; Bernal, N. (2016). "Ctenomys conoveri". IUCN Red List of Threatened Species . 2016: e.T5802A22192609. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5802A22192609.en .
  21. 1 2 Ojeda, R. (2019). "Ctenomys coyhaiquensis". IUCN Red List of Threatened Species . 2019: e.T136295A22196080. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136295A22196080.en .
  22. 1 2 Bidau, C. J. (2019). "Ctenomys dorbignyi". IUCN Red List of Threatened Species . 2019: e.T136234A22192383. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136234A22192383.en .
  23. 1 2 Bidau, C. J.; Ojeda, R. (2019). "Ctenomys emilianus". IUCN Red List of Threatened Species . 2019: e.T5804A22192803. doi: 10.2305/IUCN.UK.2019-3.RLTS.T5804A22192803.en .
  24. 1 2 Bidau, C. J. (2019). "Ctenomys famosus". IUCN Red List of Threatened Species . 2019: e.T136588A22197042. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136588A22197042.en .
  25. 1 2 de Frietas, T.; Weksler, M.; Catzeflis, F.; Percequillo, A. (2019). "Ctenomys flamarioni". IUCN Red List of Threatened Species . 2019: e.T136464A22196926. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136464A22196926.en .
  26. 1 2 Ojeda, R. (2019). "Ctenomys fochi". IUCN Red List of Threatened Species . 2019: e.T136499A22193166. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136499A22193166.en .
  27. 1 2 Bidau, C. J. (2018). "Ctenomys occultus". IUCN Red List of Threatened Species . 2018: e.T5817A22194328. doi: 10.2305/IUCN.UK.2018-1.RLTS.T5817A22194328.en .
  28. 1 2 Roach, N.; Naylor, L. (2016). "Ctenomys goodfellowi". IUCN Red List of Threatened Species . 2016: e.T136607A22196835. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136607A22196835.en .
  29. 1 2 Bidau, C. (2016). "Ctenomys perrensi". IUCN Red List of Threatened Species . 2016: e.T5820A22194434. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5820A22194434.en .
  30. 1 2 Bidau, C. (2016). "Ctenomys haigi". IUCN Red List of Threatened Species . 2016: e.T5807A22193246. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5807A22193246.en .
  31. 1 2 Dunnum, J.; Vargas, J.; Bernal, N.; Zeballos, N.; Vivar, E. (2016). "Ctenomys opimus". IUCN Red List of Threatened Species . 2016: e.T5818A22194246. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5818A22194246.en .
  32. 1 2 Hurtado Materon, M. A.; de Frietas, T.; Lawing, M.; Gonçalves, G. L.; Fornel, R. (2024). "Ctenomys ibicuiensis". IUCN Red List of Threatened Species . 2024: e.T45958852A259145694. doi: 10.2305/IUCN.UK.2024-2.RLTS.T45958852A259145694.en .
  33. 1 2 Ojeda, R. (2019). "Ctenomys juris". IUCN Red List of Threatened Species . 2019: e.T136556A22196428. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136556A22196428.en .
  34. 1 2 Ojeda, R. (2019). "Ctenomys fodax". IUCN Red List of Threatened Species . 2019: e.T136601A22196762. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136601A22196762.en .
  35. 1 2 Bidau, C. J. (2018). "Ctenomys lami". IUCN Red List of Threatened Species . 2018: e.T136567A22196245. doi: 10.2305/IUCN.UK.2018-1.RLTS.T136567A22196245.en .
  36. 1 2 Roach, N.; Naylor, L. (2016). "Ctenomys lewisi". IUCN Red List of Threatened Species . 2016: e.T5811A22193810. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5811A22193810.en .
  37. 1 2 Bidau, C. J. (2019). "Ctenomys magellanicus". IUCN Red List of Threatened Species . 2019: e.T5812A22193726. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5812A22193726.en .
  38. 1 2 Bidau, C. (2016). "Ctenomys maulinus". IUCN Red List of Threatened Species . 2016: e.T5813A22193961. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5813A22193961.en .
  39. 1 2 Bidau, Ojeda; R. (2017) [errata version of 2016 assessment]. "Ctenomys mendocinus". IUCN Red List of Threatened Species . 2016: e.T5814A115080043. doi: 10.2305/IUCN.UK.2016-3.RLTS.T5814A22193878.en .
  40. 1 2 Bidau, C. J. (2019). "Ctenomys latro". IUCN Red List of Threatened Species . 2019: e.T5809A22193629. doi: 10.2305/IUCN.UK.2019-3.RLTS.T5809A22193629.en .
  41. 1 2 Bidau, C. J. (2019). "Ctenomys pearsoni". IUCN Red List of Threatened Species . 2019: e.T5819A22194513. doi: 10.2305/IUCN.UK.2019-3.RLTS.T5819A22194513.en .
  42. 1 2 Zeballos, H.; Vivar, E. (2016). "Ctenomys peruanus". IUCN Red List of Threatened Species . 2016: e.T5821A22194718. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5821A22194718.en .
  43. 1 2 Bidau, C. J. (2019). "Ctenomys pilarensis". IUCN Red List of Threatened Species . 2019: e.T136289A22195989. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136289A22195989.en .
  44. 1 2 Bidau, C. J. (2020). "Ctenomys porteousi". IUCN Red List of Threatened Species . 2020: e.T5823A22194905. doi: 10.2305/IUCN.UK.2020-3.RLTS.T5823A22194905.en .
  45. 1 2 Bidau, C. J. (2019). "Ctenomys pundti". IUCN Red List of Threatened Species . 2019: e.T136370A22194037. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136370A22194037.en .
  46. 1 2 Bidau, C. J.; Ojeda, R. (2019). "Ctenomys coludo". IUCN Red List of Threatened Species . 2019: e.T136308A22196340. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136308A22196340.en .
  47. 1 2 Vivar, E. (2018) [errata version of 2017 assessment]. "Ctenomys frater". IUCN Red List of Threatened Species . 2017: e.T115553730A123796865. doi: 10.2305/IUCN.UK.2017-3.RLTS.T115553730A115553734.en .
  48. 1 2 Bidau, C. J. (2018). "Ctenomys osvaldoreigi". IUCN Red List of Threatened Species . 2018: e.T136390A22195057. doi: 10.2305/IUCN.UK.2018-1.RLTS.T136390A22195057.en .
  49. 1 2 Bidau, C. J. (2018). "Ctenomys rionegrensis". IUCN Red List of Threatened Species . 2018: e.T136635A22193418. doi: 10.2305/IUCN.UK.2018-1.RLTS.T136635A22193418.en .
  50. 1 2 Ojeda, R. (2019). "Ctenomys tuconax". IUCN Red List of Threatened Species . 2019: e.T5830A22195740. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5830A22195740.en .
  51. 1 2 Bidau, C. J. (2018). "Ctenomys roigi". IUCN Red List of Threatened Species . 2018: e.T136633A22193077. doi: 10.2305/IUCN.UK.2018-1.RLTS.T136633A22193077.en .
  52. 1 2 Ojeda, R. (2019). "Ctenomys saltarius". IUCN Red List of Threatened Species . 2019: e.T5824A22194818. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5824A22194818.en .
  53. 1 2 Bidau, C. J. (2019). "Ctenomys johannis". IUCN Red List of Threatened Species . 2019: e.T136281A22195842. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136281A22195842.en .
  54. 1 2 Ojeda, R. (2019). "Ctenomys pontifex". IUCN Red List of Threatened Species . 2019: e.T5822A22194645. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5822A22194645.en .
  55. 1 2 Ojeda, R. (2019). "Ctenomys scagliai". IUCN Red List of Threatened Species . 2019: e.T136568A22196158. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136568A22196158.en .
  56. 1 2 Bidau, C. J. (2019). "Ctenomys tulduco". IUCN Red List of Threatened Species . 2019: e.T136826A22197188. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136826A22197188.en .
  57. 1 2 Bidau, C. J. (2019). "Ctenomys sericeus". IUCN Red List of Threatened Species . 2019: e.T5825A22195256. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5825A22195256.en .
  58. 1 2 Bidau, C. J. (2018). "Ctenomys sociabilis". IUCN Red List of Threatened Species . 2018: e.T5826A22195323. doi: 10.2305/IUCN.UK.2018-1.RLTS.T5826A22195323.en .
  59. 1 2 Bidau, C. J. (2018). "Ctenomys australis". IUCN Red List of Threatened Species . 2018: e.T5796A78319377. doi: 10.2305/IUCN.UK.2018-1.RLTS.T5796A78319377.en .
  60. 1 2 Roach, N.; Naylor, L. (2016). "Ctenomys steinbachi". IUCN Red List of Threatened Species . 2016: e.T5827A22194994. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5827A22194994.en .
  61. 1 2 Ojeda, R. (2019). "Ctenomys validus". IUCN Red List of Threatened Species . 2019: e.T5832A22195532. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5832A22195532.en .
  62. 1 2 Bidau, C. (2016). "Ctenomys talarum". IUCN Red List of Threatened Species . 2016: e.T5828A22195175. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5828A22195175.en .
  63. 1 2 Bidau, C. J.; Ojeda, R. (2019). "Ctenomys fulvus". IUCN Red List of Threatened Species . 2019: e.T5806A22192983. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5806A22192983.en .
  64. 1 2 Dunnum, J.; Bernal, N. (2019). "Ctenomys minutus". IUCN Red List of Threatened Species . 2019: e.T5815A22194130. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5815A22194130.en .
  65. 1 2 Ojeda, R. (2019). "Ctenomys tucumanus". IUCN Red List of Threatened Species . 2019: e.T5831A22195435. doi: 10.2305/IUCN.UK.2019-1.RLTS.T5831A22195435.en .
  66. 1 2 Bidau, C. J. (2018). "Ctenomys viperinus". IUCN Red List of Threatened Species . 2018: e.T136758A22195904. doi: 10.2305/IUCN.UK.2018-2.RLTS.T136758A22195904.en .
  67. 1 2 Zeballos, H.; Vivar, E. (2016). "Ctenomys leucodon". IUCN Red List of Threatened Species . 2016: e.T5810A22193526. doi: 10.2305/IUCN.UK.2016-2.RLTS.T5810A22193526.en .
  68. 1 2 Bidau, C. J.; Ojeda, R. (2019). "Ctenomys yolandae". IUCN Red List of Threatened Species . 2019: e.T136681A22196679. doi: 10.2305/IUCN.UK.2019-1.RLTS.T136681A22196679.en .

Sources