List of nesomyids

Last updated

Malagasy giant rat (Hypogeomys antimena) Krecek velky zoo praha 2.jpg
Malagasy giant rat (Hypogeomys antimena)

Nesomyidae is a family of mammals in the order Rodentia and part of the Myomorpha suborder. Members of this family are called nesomyids, and include Malagasy rodents, climbing mice, African rock mice, pouched rats, and tufted-tailed rats. They are found in sub-Saharan Africa, primarily in forests, savannas, shrublands, grasslands, and deserts, though some species can be found in rocky areas. They range in size from Monard's African climbing mouse, at 4 cm (2 in) plus a 5 cm (2 in) tail, to the southern giant pouched rat, at 41 cm (16 in) plus a 45 cm (18 in) tail. Nesomyids are generally omnivores, and eat seeds, fruit, nuts, roots, stems, and insects. Few nesomyids have population estimates, but five species—the white-tipped tufted-tailed rat, greater big-footed mouse, western nesomys, eastern voalavo, and Petter's tufted-tailed rat—are categorized as endangered, while the Mount Kahuzi climbing mouse and Malagasy giant rat are categorized as critically endangered.

Contents

The 67 extant species of Nesomyidae are divided into 21 genera, divided into 6 subfamilies. Cricetomyinae contains 8 species of hamster-rats and pouched rats in 3 genera, Delanymyinae contains a single species, Dendromurinae contains 26 species of climbing mice and fat mice in 6 genera, Mystromyinae contains a single species, Nesomyinae contains 27 species of short-tailed rats, tufted-tailed rats, and big-footed mice in 9 genera, and Petromyscinae contains four species of rock mice in a single genus. Several extinct prehistoric nesomyid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed. [1]

Conventions

IUCN Red List categories
Conservation status
 EX  Extinct (0 species)
 EW  Extinct in the wild (0 species)
 CR  Critically endangered (2 species)
 EN  Endangered (5 species)
 VU  Vulnerable (5 species)
 NT  Near threatened (0 species)
 LC  Least concern (45 species)
Other categories
 DD  Data deficient (8 species)
 NE  Not evaluated (2 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the nesomyid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

Nesomyidae is a family consisting of 67 extant species in 21 genera. These genera are divided into six subfamilies: Cricetomyinae, containing 8 species of in 3 genera; Delanymyinae, containing a single species; Dendromurinae, containing 26 species in 6 genera; Mystromyinae, containing a single species; Nesomyinae, containing 27 species in 9 genera; and Petromyscinae, containing 4 species in a single genus. This does not include hybrid species or extinct prehistoric species.

Family Nesomyidae

Nesomyidae [2]

Nesomyids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. [3]

Subfamily Cricetomyinae

Genus Beamys Thomas, 1909 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Greater hamster-rat


B. major
Dollman, 1914
Southeastern AfricaSize: 13–18 cm (5–7 in) long, plus 12–15 cm (5–6 in) tail [4]

Habitat: Forest [5]

Diet: Seeds and fruit [5]
 NE 


Unknown Blue question mark (italic).svg

Lesser hamster-rat

Beamys hindei 156259379.jpg

B. hindei
Thomas, 1909
Southeastern AfricaSize: 12–16 cm (5–6 in) long, plus 11–13 cm (4–5 in) tail [4]

Habitat: Forest [6]

Diet: Seeds and fruit [5]
 LC 


Unknown Decrease2.svg [6]

Genus Cricetomys Waterhouse, 1840 – four species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Emin's pouched rat

Crycetomis emini.jpg

C. emini
Wroughton, 1910
Central and western AfricaSize: 30–35 cm (12–14 in) long, plus 32–43 cm (13–17 in) tail [4]

Habitat: Forest [7]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails [8]
 LC 


Unknown Steady2.svg [7]

Gambian pouched rat

Cricetomys gambianus as a captive pet.jpg

C. gambianus
Waterhouse, 1840
Central and western AfricaSize: 28–39 cm (11–15 in) long, plus 25–40 cm (10–16 in) tail [4]

Habitat: Forest and savanna [9]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails [8]
 LC 


Unknown Steady2.svg [9]

Kivu giant pouched rat


C. kivuensis
Lönnberg, 1917
Central and western AfricaSize: About 32 cm (13 in) long, plus about 34 cm (13 in) tail [4]

Habitat: Forest [10]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails [8]
 LC 


Unknown Steady2.svg [10]

Southern giant pouched rat

Southern Giant Pouched Rat imported from iNaturalist photo 34781286 on 19 April 2024.jpg

C. ansorgei
Thomas, 1904
Central and southern Africa
Cricetomys ansorgei range.tif
Size: 29–41 cm (11–16 in) long, plus 34–45 cm (13–18 in) tail [4]

Habitat: Forest and savanna [11]

Diet: Omnivorous, including fruit and nuts as well as vegetables, insects, crabs, and snails [8]
 LC 


Unknown Steady2.svg [11]

Genus Saccostomus Peters, 1846 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Mearns's pouched mouse


S. mearnsi
Heller, 1910
Eastern AfricaSize: 10–15 cm (4–6 in) long, plus 3–8 cm (1–3 in) tail [4]

Habitat: Savanna, shrubland, and desert [12]

Diet: Seeds, berries, grain, and nuts, as well as insects [13]
 LC 


Unknown Steady2.svg [12]

South African pouched mouse

Saccostomus campestris.jpg

S. campestris
Peters, 1846
Southern AfricaSize: 9–18 cm (4–7 in) long, plus 2–6 cm (1–2 in) tail [4]

Habitat: Savanna, shrubland, grassland, and desert [14]

Diet: Seeds, berries, grain, and nuts, as well as insects [13]
 LC 


Unknown Steady2.svg [14]

Subfamily Delanymyinae

Genus Delanymys Hayman, 1962 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Delany's mouse


D. brooksi
Hayman, 1962
Central Africa
Delanymys brooksi distribution.svg
Size: 5–7 cm (2–3 in) long, plus 8–12 cm (3–5 in) tail [4]

Habitat: Shrubland and inland wetlands [15]

Diet: Seeds [5]
 VU 


Unknown Decrease2.svg [15]

Subfamily Dendromurinae

Genus Dendromus Smith, 1829 – fourteen species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Banana climbing mouse


D. messorius
(Thomas, 1903)
Central and western AfricaSize: 5–7 cm (2–3 in) long, plus 7–10 cm (3–4 in) tail [16]

Habitat: Grassland [17]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Steady2.svg [17]

Brants's climbing mouse

Dendromus mesomelas.jpg

D. mesomelas
(Brants, 1827)
Southern AfricaSize: 6–9 cm (2–4 in) long, plus 7–13 cm (3–5 in) tail [16]

Habitat: Forest, savanna, shrubland, and grassland [19]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Steady2.svg [19]

Cameroon climbing mouse


D. oreas
Osgood, 1936
Cameroon Size: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail [16]

Habitat: Grassland [20]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Blue question mark (italic).svg [20]

Chestnut climbing mouse

Klatre08.jpg

D. mystacalis
(Heuglin, 1863)
Eastern and southern AfricaSize: 4–8 cm (2–3 in) long, plus 6–11 cm (2–4 in) tail [16]

Habitat: Savanna and grassland [21]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Steady2.svg [21]

Gray climbing mouse

Grey Climbing Mouse.jpg

D. melanotis
(Smith, 1834)
Sub-Saharan Africa Size: 5–8 cm (2–3 in) long, plus 5–10 cm (2–4 in) tail [16]

Habitat: Savanna, shrubland, grassland, and desert [22]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Steady2.svg [22]

Kivu climbing mouse


D. nyasae
Thomas, 1916
Southern AfricaSize: 6–8 cm (2–3 in) long, plus 8–11 cm (3–4 in) tail [16]

Habitat: Forest, savanna, shrubland, and grassland [23]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Steady2.svg [23]

Lachaise's climbing mouse


D. lachaisei
Denys & Aniskine, 2012
Western Africa
Distribution of Dendromus lachaisei.png
Size: 5–8 cm (2–3 in) long, plus 7–10 cm (3–4 in) tail [16]

Habitat: Forest and savanna [24]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 VU 


Unknown Blue question mark (italic).svg [24]

Lovat's climbing mouse

DendromysLovatiSmit white background.jpg

D. lovati
(de Winton, 1900)
Ethiopia Size: 5–10 cm (2–4 in) long, plus 5–9 cm (2–4 in) tail [16]

Habitat: Grassland [25]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Decrease2.svg [25]

Monard's African climbing mouse


D. leucostomus
Monard, 1933
Angola Size: 4–7 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail [16]

Habitat: Savanna and grassland [18]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 NE 


Unknown Blue question mark (italic).svg

Montane African climbing mouse


D. insignis
(Thomas, 1903)
East-central AfricaSize: 7–10 cm (3–4 in) long, plus 8–12 cm (3–5 in) tail [16]

Habitat: Forest, shrubland, grassland, and inland wetlands [26]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Steady2.svg [26]

Mount Kahuzi climbing mouse


D. kahuziensis
(Dieterlen, 1969)
Democratic Republic of the Congo Size: 7–9 cm (3–4 in) long, plus 12–14 cm (5–6 in) tail [16]

Habitat: Forest [27]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 CR 


Unknown Decrease2.svg [27]

Nyika climbing mouse


D. nyikae
Wroughton, 1909
Southern AfricaSize: 6–8 cm (2–3 in) long, plus 8–10 cm (3–4 in) tail [16]

Habitat: Forest, savanna, shrubland, and grassland [28]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 LC 


Unknown Blue question mark (italic).svg [28]

Rupp's African climbing mouse


D. ruppi
Dieterlen, 2009
South Sudan Size: 6–9 cm (2–4 in) long, plus 9–12 cm (4–5 in) tail [16]

Habitat: Inland wetlands [29]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 DD 


Unknown Blue question mark (italic).svg [29]

Vernay's climbing mouse


D. vernayi
Hill & Carter, 1937
AngolaSize: 6–8 cm (2–3 in) long, plus 8–9 cm (3–4 in) tail [16]

Habitat: Savanna [30]

Diet: Seeds, berries, insects, eggs, small lizards, and small birds [18]
 DD 


Unknown Blue question mark (italic).svg [30]

Genus Dendroprionomys – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Velvet climbing mouse


D. rousseloti
Petter, 1966
Republic of the Congo Size: About 8 cm (3 in) long, plus about 11 cm (4 in) tail [16]

Habitat: Unknown [31]

Diet: Insects [32]
 DD 


Unknown Blue question mark (italic).svg [31]

Genus Malacothrix Candolle, 1838 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Gerbil mouse


M. typica
(Smith, 1834)
Southern AfricaSize: 6–9 cm (2–4 in) long, plus 2–5 cm (1–2 in) tail [16]

Habitat: Desert, shrubland, and savanna [33]

Diet: Green vegetation [32]
 LC 


Unknown Steady2.svg [33]

Genus Megadendromus Dieterlen & Rupp, 1978 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Nikolaus's mouse


M. nikolausi
Dieterlen & Rupp, 1978
EthiopiaSize: 11–13 cm (4–5 in) long, plus 9–11 cm (4 in) tail [16]

Habitat: Shrubland [34]

Diet: Likely vegetation and insects [35]
 VU 


Unknown Blue question mark (italic).svg [34]

Genus Prionomys Dollman, 1910 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Dollman's tree mouse

Dollman-s-Tree-Mouse-Prionomys-batesi.jpg

P. batesi
Dollman, 1910
West-central AfricaSize: 7–9 cm (3–4 in) long, plus 10–12 cm (4–5 in) tail [16]

Habitat: Forest [36]

Diet: Insects [32]
 DD 


Unknown Blue question mark (italic).svg [36]

Genus Steatomys Peters, 1846 – eight species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Bocage's African fat mouse


S. bocagei
(Thomas, 1892)
Angola and Democratic Republic of the CongoSize: 9–13 cm (4–5 in) long, plus 5–8 cm (2–3 in) tail [16]

Habitat: Forest, savanna, and shrubland [37]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Blue question mark (italic).svg [37]

Dainty fat mouse


S. cuppedius
Thomas & Hinton, 1920
Western AfricaSize: 7–10 cm (3–4 in) long, plus 4–5 cm (2 in) tail [16]

Habitat: Shrubland and grassland [39]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Steady2.svg [39]

Fat mouse

Steatomys pratensis 178572260.jpg

S. pratensis
Peters, 1846
Central and southern AfricaSize: 8–11 cm (3–4 in) long, plus 4–6 cm (2 in) tail [16]

Habitat: Savanna and grassland [40]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Steady2.svg [40]

Jackson's fat mouse


S. jacksoni
Hayman, 1936
Ghana and Nigeria Size: About 12 cm (5 in) long, plus about 5 cm (2 in) tail [16]

Habitat: Unknown [41]

Diet: Seeds, grass bulbs, and insects [38]
 DD 


Unknown Blue question mark (italic).svg [41]

Krebs's fat mouse

Steatomys krebsii krebsii 15204345.jpg

S. krebsii
Peters, 1852
Southern AfricaSize: 7–10 cm (3–4 in) long, plus 3–6 cm (1–2 in) tail [16]

Habitat: Shrubland and grassland [42]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Steady2.svg [42]

Northwestern fat mouse


S. caurinus
Thomas, 1912
Western AfricaSize: 9–13 cm (4–5 in) long, plus 3–5 cm (1–2 in) tail [16]

Habitat: Shrubland [43]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Steady2.svg [43]

Pousargues African fat mouse


S. opimus
(Pousargues, 1894)
Central AfricaSize: 11–13 cm (4–5 in) long, plus 5–7 cm (2–3 in) tail [16]

Habitat: Forest and savanna [44]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Blue question mark (italic).svg [44]

Tiny fat mouse


S. parvus
Rhoads, 1896
Eastern and southern AfricaSize: 5–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail [16]

Habitat: Savanna, shrubland, and grassland [45]

Diet: Seeds, grass bulbs, and insects [38]
 LC 


Unknown Steady2.svg [45]

Subfamily Mystromyinae

Genus Mystromys Wagner, 1841 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
White-tailed rat


M. albicaudatus
(Smith, 1834)
Southern AfricaSize: 11–20 cm (4–8 in) long, plus 4–9 cm (2–4 in) tail [4]

Habitat: Shrubland and grassland [46]

Diet: Seeds, vegetable matter, and insects [47]
 VU 


7,000–14,000 Decrease2.svg [46]

Subfamily Nesomyinae

Genus Brachytarsomys Günther, 1875 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Hairy-tailed antsangy


B. villosa
Petter, 1962
Northern Madagascar
Brachytarsomys villosa range map.svg
Size: 23–24 cm (9 in) long, plus 26–27 cm (10–11 in) tail [48]

Habitat: Forest [49]

Diet: Fruit [50]
 VU 


Unknown Blue question mark (italic).svg [49]

White-tailed antsangy

BrachytarsomysAlbicaudataWolfSmit.jpg

B. albicauda
Günther, 1875
Eastern Madagascar
Brachytarsomys albicauda range map.svg
Size: 22–23 cm (9 in) long, plus 22–24 cm (9 in) tail [48]

Habitat: Forest [51]

Diet: Fruit [50]
 LC 


Unknown Blue question mark (italic).svg [51]

Genus Brachyuromys Forsyth Major, 1896 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Betsileo short-tailed rat


B. betsileoensis
(Bartlett, 1880)
Eastern Madagascar
Brachyuromys betsileoensis range map.svg
Size: 14–19 cm (6–7 in) long, plus 7–10 cm (3–4 in) tail [4]

Habitat: Forest, shrubland, grassland, and inland wetlands [52]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [52]

Gregarious short-tailed rat


B. ramirohitra
Forsyth Major, 1896
Eastern Madagascar
Brachyuromys ramirohitra range map.svg
Size: 14–17 cm (6–7 in) long, plus 8–11 cm (3–4 in) tail [4]

Habitat: Forest [53]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [53]

Genus Eliurus A. Milne-Edwards, 1885 – twelve species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Ankarana Special Reserve tufted-tailed rat


E. carletoni
Goodman, Raheriarisena, & Jansa, 2009
Northern MadagascarSize: 14–15 cm (6 in) long, plus 16–19 cm (6–7 in) tail [48]

Habitat: Forest [54]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [54]

Daniel's tufted-tailed rat


E. danieli
Goodman & Carleton, 2007
Southern Madagascar
Eliurus danieli range map.svg
Size: 15–16 cm (6 in) long, plus 17–20 cm (7–8 in) tail [48]

Habitat: Shrubland [55]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [55]

Dormouse tufted-tailed rat

Dormouse Tufted-tailed Rat (Eliurus myoxinus) (9579527484).jpg

E. myoxinus
A. Milne-Edwards, 1855
Northern, western, and southern Madagascar
Eliurus myoxinus range map.svg
Size: 11–14 cm (4–6 in) long, plus 12–17 cm (5–7 in) tail [48]

Habitat: Forest [56]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [56]

Ellerman's tufted-tailed rat


E. ellermani
Carleton, 1994
Eastern Madagascar
Eliurus ellermani range map.svg
Size: About 15 cm (6 in) long, plus about 18 cm (7 in) tail [48]

Habitat: Forest [57]

Diet: Likely vegetation and insects [35]
 DD 


Unknown Blue question mark (italic).svg [57]

Grandidier's tufted-tailed rat


E. grandidieri
Goodman & Carleton, 1998
Northeastern Madagascar
Eliurus grandidieri range map.svg
Size: 11–17 cm (4–7 in) long, plus 14–18 cm (6–7 in) tail [48]

Habitat: Forest [58]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [58]

Lesser tufted-tailed rat


E. minor
Forsyth Major, 1896
Northern and eastern Madagascar
Eliurus minor range map.svg
Size: 10–13 cm (4–5 in) long, plus 11–14 cm (4–6 in) tail [48]

Habitat: Forest [59]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [59]

Major's tufted-tailed rat


E. majori
Thomas, 1895
Eastern Madagascar
Eliurus majori range map.svg
Size: 13–17 cm (5–7 in) long, plus 15–20 cm (6–8 in) tail [48]

Habitat: Forest [60]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [60]

Petter's tufted-tailed rat


E. petteri
Carleton, 1994
Eastern MadagascarSize: About 13 cm (5 in) long, plus about 19 cm (7 in) tail [48]

Habitat: Forest [61]

Diet: Likely vegetation and insects [35]
 EN 


Unknown Blue question mark (italic).svg [61]

Tanala tufted-tailed rat


E. tanala
Forsyth Major, 1896
Eastern Madagascar
Eliurus tanala range map.svg
Size: 14–16 cm (6 in) long, plus 15–20 cm (6–8 in) tail [48]

Habitat: Forest [62]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [62]

Tsingy tufted-tailed rat


E. antsingy
Goodman, Rakotondravony, & Carleton, 2001
Northern Madagascar
Eliurus antsingy range map.svg
Size: 14–16 cm (6 in) long, plus 15–20 cm (6–8 in) tail [48]

Habitat: Forest [63]

Diet: Likely vegetation and insects [35]
 DD 


Unknown Blue question mark (italic).svg [63]

Webb's tufted-tailed rat


E. webbi
Ellerman, 1949
Eastern Madagascar
Eliurus webbi range map.svg
Size: 14–16 cm (6 in) long, plus 16–19 cm (6–7 in) tail [48]

Habitat: Forest [64]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [64]

White-tipped tufted-tailed rat


E. penicillatus
Thomas, 1908
East-central Madagascar
Eliurus penicillatus range map.svg
Size: About 15 cm (6 in) long, plus about 17 cm (7 in) tail [48]

Habitat: Forest [65]

Diet: Likely vegetation and insects [35]
 EN 


Unknown Blue question mark (italic).svg [65]

Genus Gymnuromys Forsyth Major, 1896 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Voalavoanala


G. roberti
Forsyth Major, 1896
Eastern Madagascar
Gymnuromys roberti range map.svg
Size: 15–18 cm (6–7 in) long, plus 14–20 cm (6–8 in) tail [48]

Habitat: Forest [66]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [66]

Genus Hypogeomys Grandidier, 1869 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Malagasy giant rat

Malagasy giant rat.jpg

H. antimena
Grandidier, 1869
Western Madagascar
Hypogeomys antimena range map.svg
Size: 30–34 cm (12–13 in) long, plus 21–24 cm (8–9 in) tail [48]

Habitat: Forest [67]

Diet: Fruit [68]
 CR 


5,000 Decrease2.svg [67]

Genus Macrotarsomys A. Milne-Edwards & Grandidier, 1898 – three species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Bastard big-footed mouse

Bastard big-footed mouse, Reserve forestiere de Kirindy, Kirindy Village, Madagascar imported from iNaturalist photo 455479106.jpg

M. bastardi
A. Milne-Edwards & Grandidier, 1898
Western Madagascar
Macrotarsomys bastardi range map.svg
Size: 8–11 cm (3–4 in) long, plus 12–15 cm (5–6 in) tail [48]

Habitat: Forest and shrubland [69]

Diet: Berries, fruit, seeds, roots, and stems [50]
 LC 


Unknown Steady2.svg [69]

Greater big-footed mouse


M. ingens
(Petter, 1959)
Northern Madagascar
Macrotarsomys ingens range map.svg
Size: 11–15 cm (4–6 in) long, plus 18–24 cm (7–9 in) tail [48]

Habitat: Forest [70]

Diet: Berries, fruit, seeds, roots, and stems [50]
 EN 


Unknown Decrease2.svg [70]

Petter's big-footed mouse


M. petteri
Goodman & Soarimalala, 2005
Southern Madagascar
Macrotarsomys petteri range.svg
Size: About 16 cm (6 in) long, plus about 24 cm (9 in) tail [48]

Habitat: Forest [71]

Diet: Berries, fruit, seeds, roots, and stems [50]
 DD 


Unknown Blue question mark (italic).svg [71]

Genus Monticolomys Goodman & Carleton, 1996 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Malagasy mountain mouse


M. koopmani
Goodman & Carleton, 1996
Northern, central, and southern Madagascar
Monticolomys range.svg
Size: 9–11 cm (4 in) long, plus 13–15 cm (5–6 in) tail [48]

Habitat: Forest [72]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Blue question mark (italic).svg [72]

Genus Nesomys Peters, 1870 – three species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Island mouse

Nesomys rufus - Red forest rat (15721901477).jpg

N. rufus
Peters, 1870
Eastern Madagascar
Nesomys rufus range map.svg
Size: 17–20 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail [4]

Habitat: Forest [73]

Diet: Fruit and nuts [50]
 LC 


Unknown Blue question mark (italic).svg [73]

Western nesomys

Nesomys lambertoni 243136443.jpg

N. lambertoni
Grandidier, 1928
Western MadagascarSize: 18–20 cm (7–8 in) long, plus 18–20 cm (7–8 in) tail [4]

Habitat: Forest [74]

Diet: Fruit and nuts [50]
 EN 


Unknown Blue question mark (italic).svg [74]

White-bellied nesomys

Nesomys audeberti 260723875.jpg

N. audeberti
Jentink, 1879
Eastern MadagascarSize: 19–20 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail [4]

Habitat: Forest [75]

Diet: Fruit and nuts [50]
 LC 


Unknown Decrease2.svg [75]

Genus Voalavo Goodman & Carleton, 1998 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Eastern voalavo


V. antsahabensis
Goodman, Rakotondravony, Randriamanantsoa, & Rakotomalala-Razanahoera, 2005
Central Madagascar (in green)
Voalavo range.svg
Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail [48]

Habitat: Forest [76]

Diet: Likely vegetation and insects [35]
 EN 


Unknown Blue question mark (italic).svg [76]

Northern voalavo


V. gymnocaudus
Goodman & Carleton, 1998
Northern Madagascar (in red)
Voalavo range.svg
Size: 8–9 cm (3–4 in) long, plus 12–13 cm (5 in) tail [48]

Habitat: Forest [77]

Diet: Likely vegetation and insects [35]
 LC 


Unknown Steady2.svg [77]

Subfamily Petromyscinae

Genus Petromyscus Thomas, 1926 – four species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Barbour's rock mouse


P. barbouri
Shortridge, 1938
South Africa Size: 7–8 cm (3 in) long, plus 7–9 cm (3–4 in) tail [4]

Habitat: Shrubland and rocky areas [78]

Diet: Omnivorous [5]
 LC 


Unknown Steady2.svg [78]

Brukkaros pygmy rock mouse


P. monticularis
(Thomas & Hinton, 1925)
Namibia and South AfricaSize: 6–10 cm (2–4 in) long, plus 6–9 cm (2–4 in) tail [4]

Habitat: Shrubland and rocky areas [79]

Diet: Omnivorous [5]
 LC 


Unknown Steady2.svg [79]

Pygmy rock mouse


P. collinus
(Hinton, 1925)
Southwestern AfricaSize: 7–10 cm (3–4 in) long, plus 7–13 cm (3–5 in) tail [4]

Habitat: Shrubland and rocky areas [80]

Diet: Omnivorous [5]
 LC 


Unknown Steady2.svg [80]

Shortridge's rock mouse


P. shortridgei
Thomas, 1926
Angola and NamibiaSize: 6–10 cm (2–4 in) long, plus 7–11 cm (3–4 in) tail [4]

Habitat: Shrubland and rocky areas [81]

Diet: Omnivorous [5]
 LC 


Unknown Steady2.svg [81]

References

  1. "Fossilworks: Nesomyidae". Paleobiology Database . University of Wisconsin–Madison. Archived from the original on November 26, 2024. Retrieved September 18, 2025.
  2. Steppan, S. J.; Schenk, J. J. (2017). "Muroid rodent phylogenetics: 900-species tree reveals increasing diversification rates". PLOS One . 12 (8) e0183070. doi: 10.1371/journal.pone.0183070 . PMC   5559066 .
  3. Wilson; Reeder, pp. 930–954
  4. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 Chernasky; Motis; Burgin, pp. 216–217
  5. 1 2 3 4 5 6 7 8 Nowak, pp. 1492–1493
  6. 1 2 Cassola, F. (2017) [errata version of 2016 assessment]. "Beamys hindei". IUCN Red List of Threatened Species . 2016 e.T2676A115063057. doi: 10.2305/IUCN.UK.2016-3.RLTS.T2676A22238241.en .
  7. 1 2 Cassola, F. (2017) [errata version of 2016 assessment]. "Cricetomys emini". IUCN Red List of Threatened Species . 2016 e.T5521A115072329. doi: 10.2305/IUCN.UK.2016-3.RLTS.T5521A22236579.en .
  8. 1 2 3 4 Nowak, p. 1496
  9. 1 2 Kennerley, R. (2023) [amended version of 2019 assessment]. "Cricetomys gambianus". IUCN Red List of Threatened Species . 2023 e.T112169507A243173564. doi: 10.2305/IUCN.UK.2023-1.RLTS.T112169507A243173564.en .
  10. 1 2 Cassola, F. (2017) [errata version of 2016 assessment]. "Cricetomys emini". IUCN Red List of Threatened Species . 2016 e.T5521A115072329. doi: 10.2305/IUCN.UK.2016-3.RLTS.T5521A22236579.en .
  11. 1 2 Dando, T.; Kennerley, R. (2023) [amended version of 2019 assessment]. "Cricetomys ansorgei". IUCN Red List of Threatened Species . 2023 e.T112256552A243175763. doi: 10.2305/IUCN.UK.2023-1.RLTS.T112256552A243175763.en .
  12. 1 2 Cassola, F. (2017) [errata version of 2016 assessment]. "Saccostomus mearnsi". IUCN Red List of Threatened Species . 2016 e.T19809A115153561. doi: 10.2305/IUCN.UK.2016-3.RLTS.T19809A22234073.en .
  13. 1 2 Nowak, p. 1495
  14. 1 2 Cassola, F.; Child, M. F. (2017) [errata version of 2016 assessment]. "Saccostomus campestris". IUCN Red List of Threatened Species . 2016 e.T19808A115153363. doi: 10.2305/IUCN.UK.2016-3.RLTS.T19808A22233928.en .
  15. 1 2 Kennerley, R.; Kerbis Peterhans, J. (2020) [errata version of 2016 assessment]. "Delanymys brooksi". IUCN Red List of Threatened Species . 2016 e.T6313A184098557. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6313A184098557.en .
  16. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 Chernasky; Motis; Burgin, pp. 218–219
  17. 1 2 Schlitter, D.; Dieterlen, F. (2017) [errata version of 2016 assessment]. "Dendromus messorius". IUCN Red List of Threatened Species . 2016 e.T6445A115081283. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6445A22235137.en .
  18. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 Nowak, p. 1487
  19. 1 2 Child, M. F.; Monadjem, A. (2016). "Dendromus mesomelas". IUCN Red List of Threatened Species . 2016 e.T6444A22235226. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6444A22235226.en .
  20. 1 2 Kennerley, R. (2017) [errata version of 2016 assessment]. "Dendromus oreas". IUCN Red List of Threatened Species . 2016 e.T6448A115081767. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6448A22234774.en .
  21. 1 2 Child, M. F.; Monadjem, A. (2017) [errata version of 2016 assessment]. "Dendromus mystacalis". IUCN Red List of Threatened Species . 2016 e.T6446A115081411. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6446A22234993.en .
  22. 1 2 Child, M. F.; Monadjem, A. (2016). "Dendromus melanotis". IUCN Red List of Threatened Species . 2016 e.T6443A22235350. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6443A22235350.en .
  23. 1 2 Child, M. F.; Monadjem, A. (2016). "Dendromus mesomelas". IUCN Red List of Threatened Species . 2016 e.T6444A22235226. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6444A22235226.en .
  24. 1 2 Denys, C.; Kennerley, R. (2024). "Dendromus lachaisei". IUCN Red List of Threatened Species . 2024 e.T45958985A258446545. doi: 10.2305/IUCN.UK.2024-2.RLTS.T45958985A258446545.en .
  25. 1 2 Lavrenchenko, L. (2016). "Dendromus lovati". IUCN Red List of Threatened Species . 2016 e.T6442A22235513. doi: 10.2305/IUCN.UK.2016-2.RLTS.T6442A22235513.en .
  26. 1 2 Cassola, F. (2016). "Dendromus insignis". IUCN Red List of Threatened Species . 2016 e.T6440A22234456. doi: 10.2305/IUCN.UK.2016-2.RLTS.T6440A22234456.en .
  27. 1 2 Dando, T.; Demos, T. (2021). "Dendromus kahuziensis". IUCN Red List of Threatened Species . 2021 e.T6439A22234587. doi: 10.2305/IUCN.UK.2021-2.RLTS.T6439A22234587.en .
  28. 1 2 Cassola, F. (2017) [errata version of 2016 assessment]. "Dendromus nyikae". IUCN Red List of Threatened Species . 2016 e.T6447A115081603. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6447A22234880.en .
  29. 1 2 Denys, C.; Kennerley, R. (2024). "Dendromus ruppi". IUCN Red List of Threatened Species . 2024 e.T45959000A258446746. doi: 10.2305/IUCN.UK.2024-2.RLTS.T45959000A258446746.en .
  30. 1 2 Kennerley, R. (2017) [errata version of 2016 assessment]. "Dendromus vernayi". IUCN Red List of Threatened Species . 2016 e.T6449A115081904. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6449A22234694.en .
  31. 1 2 Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Dendroprionomys rousseloti". IUCN Red List of Threatened Species . 2016 e.T6483A115082012. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6483A22234171.en .
  32. 1 2 3 Nowak, pp. 1488–1489
  33. 1 2 Cassola, F. (2017) [errata version of 2016 assessment]. "Malacothrix typica". IUCN Red List of Threatened Species . 2016 e.T12712A115105728. doi: 10.2305/IUCN.UK.2016-3.RLTS.T12712A22238026.en .
  34. 1 2 Gerrie, R.; Kennerley, R. (2019). "Megadendromus nikolausi". IUCN Red List of Threatened Species . 2019 e.T12937A22237949. doi: 10.2305/IUCN.UK.2019-1.RLTS.T12937A22237949.en .
  35. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 Nowak, p. 1346
  36. 1 2 Gerrie, R.; Kennerley, R. (2016). "Prionomys batesi". IUCN Red List of Threatened Species . 2016 e.T18164A22236213. doi: 10.2305/IUCN.UK.2016-3.RLTS.T18164A22236213.en .
  37. 1 2 Hoffmann, M.; Cox, N. A. (2017). "Steatomys bocagei". IUCN Red List of Threatened Species . 2017 e.T136625A22233115. doi: 10.2305/IUCN.UK.2017-2.RLTS.T136625A22233115.en .
  38. 1 2 3 4 5 6 7 8 Nowak, p. 1490
  39. 1 2 Schlitter, D.; Coetzee, N. (2017). "Steatomys cuppedius". IUCN Red List of Threatened Species . 2017 e.T20719A22233546. doi: 10.2305/IUCN.UK.2017-2.RLTS.T20719A22233546.en .
  40. 1 2 Child, M. F.; Monadjem, A. (2017) [errata version of 2016 assessment]. "Steatomys pratensis". IUCN Red List of Threatened Species . 2016 e.T20722A115159593. doi: 10.2305/IUCN.UK.2016-3.RLTS.T20722A22233313.en .
  41. 1 2 Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Steatomys jacksoni". IUCN Red List of Threatened Species . 2016 e.T20717A115159338. doi: 10.2305/IUCN.UK.2016-3.RLTS.T20717A22233758.en .
  42. 1 2 Monadjem, A.; Schlitter, D. (2017) [errata version of 2016 assessment]. "Steatomys krebsii". IUCN Red List of Threatened Species . 2016 e.T20720A115159464. doi: 10.2305/IUCN.UK.2016-3.RLTS.T20720A22233457.en .
  43. 1 2 Schlitter, D.; Coetzee, N. (2017). "Steatomys caurinus". IUCN Red List of Threatened Species . 2017 e.T20718A22233646. doi: 10.2305/IUCN.UK.2017-2.RLTS.T20718A22233646.en .
  44. 1 2 Hoffmann, M.; Cox, N. A. (2017). "Steatomys opimus". IUCN Red List of Threatened Species . 2017 e.T136513A22233848. doi: 10.2305/IUCN.UK.2017-2.RLTS.T136513A22233848.en .
  45. 1 2 Schlitter, D.; Monadjem, A. (2017). "Steatomys parvus". IUCN Red List of Threatened Species . 2017 e.T20721A22233188. doi: 10.2305/IUCN.UK.2017-2.RLTS.T20721A22233188.en .
  46. 1 2 Avenant, N. L.; Wilson, B.; Power, J.; Palmer, G.; Child, M. F. (2019). "Mystromys albicaudatus". IUCN Red List of Threatened Species . 2019 e.T14262A22237378. doi: 10.2305/IUCN.UK.2019-1.RLTS.T14262A22237378.en .
  47. Nowak, p. 1425
  48. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 Chernasky; Motis; Burgin, pp. 214–215
  49. 1 2 Kennerley, R.; Goodman, S. (2016). "Brachytarsomys villosa". IUCN Red List of Threatened Species . 2016 e.T136399A22237177. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136399A22237177.en .
  50. 1 2 3 4 5 6 7 8 Nowak, pp. 1434–1435
  51. 1 2 Kennerley, R. (2016). "Brachytarsomys albicauda". IUCN Red List of Threatened Species . 2016 e.T2991A22237052. doi: 10.2305/IUCN.UK.2016-2.RLTS.T2991A22237052.en .
  52. 1 2 Kennerley, R. (2016). "Brachyuromys betsileoensis". IUCN Red List of Threatened Species . 2016 e.T2997A22234256. doi: 10.2305/IUCN.UK.2016-2.RLTS.T2997A22234256.en .
  53. 1 2 Kennerley, R. (2016). "Brachyuromys ramirohitra". IUCN Red List of Threatened Species . 2016 e.T2998A22234363. doi: 10.2305/IUCN.UK.2016-2.RLTS.T2998A22234363.en .
  54. 1 2 Kennerley, R. (2016). "Eliurus carletoni". IUCN Red List of Threatened Species . 2016 e.T48266951A48266956. doi: 10.2305/IUCN.UK.2016-3.RLTS.T48266951A48266956.en .
  55. 1 2 Hoffmann, M.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Eliurus danieli". IUCN Red List of Threatened Species . 2016 e.T136251A115205056. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136251A22238907.en .
  56. 1 2 Goodman, S. (2016). "Eliurus myoxinus". IUCN Red List of Threatened Species . 2016 e.T7623A22238673. doi: 10.2305/IUCN.UK.2016-2.RLTS.T7623A22238673.en .
  57. 1 2 Kennerley, R. (2017) [errata version of 2016 assessment]. "Eliurus ellermani". IUCN Red List of Threatened Species . 2016 e.T136325A115205656. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136325A22238824.en .
  58. 1 2 Kennerley, R. (2016). "Eliurus grandidieri". IUCN Red List of Threatened Species . 2016 e.T29460A22238512. doi: 10.2305/IUCN.UK.2016-3.RLTS.T29460A22238512.en .
  59. 1 2 Kennerley, R. (2016). "Eliurus minor". IUCN Red List of Threatened Species . 2016 e.T7621A22238349. doi: 10.2305/IUCN.UK.2016-2.RLTS.T7621A22238349.en .
  60. 1 2 Kennerley, R. (2016). "Eliurus majori". IUCN Red List of Threatened Species . 2016 e.T7622A22238442. doi: 10.2305/IUCN.UK.2016-2.RLTS.T7622A22238442.en .
  61. 1 2 Kennerley, R.; Goodman, S. (2016). "Eliurus petteri". IUCN Red List of Threatened Species . 2016 e.T136316A22238985. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136316A22238985.en .
  62. 1 2 Kennerley, R. (2016). "Eliurus tanala". IUCN Red List of Threatened Species . 2016 e.T7625A22239073. doi: 10.2305/IUCN.UK.2016-2.RLTS.T7625A22239073.en .
  63. 1 2 Kennerley, R. (2016). "Eliurus antsingy". IUCN Red List of Threatened Species . 2016 e.T48267436A22238593. doi: 10.2305/IUCN.UK.2016-3.RLTS.T48267436A22238593.en .
  64. 1 2 Kennerley, R. (2016). "Eliurus webbi". IUCN Red List of Threatened Species . 2016 e.T7626A22239153. doi: 10.2305/IUCN.UK.2016-2.RLTS.T7626A22239153.en .
  65. 1 2 Kennerley, R. (2016). "Eliurus penicillatus". IUCN Red List of Threatened Species . 2016 e.T7624A22238746. doi: 10.2305/IUCN.UK.2016-3.RLTS.T7624A22238746.en .
  66. 1 2 Kennerley, R. (2016). "Gymnuromys roberti". IUCN Red List of Threatened Species . 2016 e.T9581A22237297. doi: 10.2305/IUCN.UK.2016-2.RLTS.T9581A22237297.en .
  67. 1 2 Wilmet, L.; Volahy, A.; Hudson, M. A. (2022). "Hypogeomys antimena". IUCN Red List of Threatened Species . 2022 e.T10714A216087357. doi: 10.2305/IUCN.UK.2022-1.RLTS.T10714A216087357.en .
  68. Nowak, p. 1437
  69. 1 2 Kennerley, R. (2016). "Macrotarsomys bastardi". IUCN Red List of Threatened Species . 2016 e.T12647A22236959. doi: 10.2305/IUCN.UK.2016-2.RLTS.T12647A22236959.en .
  70. 1 2 Dollar, L.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Macrotarsomys ingens". IUCN Red List of Threatened Species . 2016 e.T12646A115105272. doi: 10.2305/IUCN.UK.2016-3.RLTS.T12646A22236876.en .
  71. 1 2 Kennerley, R. (2017) [errata version of 2016 assessment]. "Macrotarsomys petteri". IUCN Red List of Threatened Species . 2016 e.T136510A115209149. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136510A22236785.en .
  72. 1 2 Kennerley, R. (2016). "Monticolomys koopmani". IUCN Red List of Threatened Species . 2016 e.T136539A22233042. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136539A22233042.en .
  73. 1 2 Kennerley, R. (2016). "Nesomys rufus". IUCN Red List of Threatened Species . 2016 e.T14671A22237658. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14671A22237658.en .
  74. 1 2 Kennerley, R. (2017) [errata version of 2016 assessment]. "Nesomys lambertoni". IUCN Red List of Threatened Species . 2016 e.T136330A115206046. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136330A22237750.en .
  75. 1 2 Kennerley, R. (2016). "Nesomys audeberti". IUCN Red List of Threatened Species . 2016 e.T136475A22237564. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136475A22237564.en .
  76. 1 2 Kennerley, R. (2019). "Voalavo antsahabensis". IUCN Red List of Threatened Species . 2019 e.T165925A22236108. doi: 10.2305/IUCN.UK.2019-1.RLTS.T165925A22236108.en .
  77. 1 2 Kennerley, R. (2017) [errata version of 2016 assessment]. "Voalavo gymnocaudus". IUCN Red List of Threatened Species . 2016 e.T29461A115169008. doi: 10.2305/IUCN.UK.2016-3.RLTS.T29461A22236010.en .
  78. 1 2 Schlitter, D. (2016). "Petromyscus barbouri". IUCN Red List of Threatened Species . 2016 e.T16777A22235766. doi: 10.2305/IUCN.UK.2016-2.RLTS.T16777A22235766.en .
  79. 1 2 Child, M. F. (2017) [errata version of 2016 assessment]. "Petromyscus monticularis". IUCN Red List of Threatened Species . 2016 e.T16779A115139351. doi: 10.2305/IUCN.UK.2016-3.RLTS.T16779A22235937.en .
  80. 1 2 Child, M. F. (2017) [errata version of 2016 assessment]. "Petromyscus collinus". IUCN Red List of Threatened Species . 2016 e.T16778A115139225. doi: 10.2305/IUCN.UK.2016-3.RLTS.T16778A22235686.en .
  81. 1 2 Schlitter, D. (2016). "Petromyscus shortridgei". IUCN Red List of Threatened Species . 2016 e.T16780A22235864. doi: 10.2305/IUCN.UK.2016-2.RLTS.T16780A22235864.en .

Sources