List of echimyids

Last updated

From top-left, clockwise: Nutria (Myocastor coypus), Ferreira's spiny tree-rat (Mesomys hispidus), Atlantic spiny rat (Trinomys mirapitanga), and Desmarest's hutia (Capromys pilorides), which is considered for inclusion in the family Echimyidae.jpg
From top-left, clockwise: Nutria (Myocastor coypus), Ferreira's spiny tree-rat (Mesomys hispidus), Atlantic spiny rat (Trinomys mirapitanga), and Desmarest's hutia (Capromys pilorides), which is considered for inclusion in the family

Echimyidae is a family of mammals in the order Rodentia and part of the Caviomorpha parvorder. Members of this family are called echimyids or Neotropical spiny rats, and include spiny rats, tree-rats, and cave rats. They are found in South America and Central America, though the nutria has been introduced to the United States, Europe, and Japan. They live primarily in forests, savannas, and grasslands, though some species can be found in shrublands, wetlands, and rocky areas. They range in size from the Sao Lourenço punaré, at 12 cm (5 in) plus a 13 cm (5 in) tail, to the nutria, at 57 cm (22 in) plus a 40 cm (16 in) tail. Almost no echimyids have population estimates, but six species—the painted tree-rat, giant Atlantic tree-rat, Lund's Atlantic tree-rat, orange-brown Atlantic tree-rat, Moojen's Atlantic spiny rat, and Yonenaga's Atlantic spiny rat—are categorized as an endangered species, and three—the Mantiqueira Atlantic tree-rat, short-furred Atlantic tree-rat, and red-crested tree-rat—are categorized as critically endangered, while six species of Caribbean cave rat, comprising the three genera of the subfamily Heteropsomyinae, were made extinct since 1500 due to the European colonization of the Americas and introduction of non-native rats, with some species surviving until the 1900s. [1]

Contents

The 88 extant species of Echimyidae are divided into three subfamilies: Caterodontinae, containing a single species; Echimyinae, containing 75 species in 18 genera, and Euryzygomatomyinae, containing 12 species in 3 genera. The family Capromyidae, or the hutias, has been proposed to be merged into Echimyidae as the subfamily Capromyinae, but the proposal is not yet universally accepted. [2] A few extinct prehistoric echimyid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed. [3]

Conventions

IUCN Red List categories
Conservation status
 EX  Extinct (6 species)
 EW  Extinct in the wild (0 species)
 CR  Critically endangered (3 species)
 EN  Endangered (6 species)
 VU  Vulnerable (0 species)
 NT  Near threatened (2 species)
 LC  Least concern (48 species)
Other categories
 DD  Data deficient (29 species)
 NE  Not evaluated (0 species)

The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the echimyid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species, subspecies, or genera listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol: "".

Classification

Echimyidae is a family consisting of 88 extant species in 22 genera. These species are divided into three subfamilies: Caterodontinae, containing a single species; Echimyinae, containing 75 species in 18 genera, and Euryzygomatomyinae, containing 12 species in 3 genera. Additionally, the subfamily Heteropsomyinae, containing 6 species in 3 genera, was made extinct since 1500.

Echimyidae [4]

Echimyids

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. [5]

Subfamily Caterodontinae

Genus Carterodon Waterhouse, 1848 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Owl's spiny rat


C. sulcidens
(Lund, 1841)
Eastern BrazilSize: 13–25 cm (5–10 in) long, plus 6–10 cm (2–4 in) tail [6]

Habitat: Forest and savanna [7]

Diet: Vegetation [8]
 DD 


Unknown Blue question mark (italic).svg [7]

Subfamily Echimyinae

Genus Callistomys Emmons & Vucetich, 1998 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Painted tree-rat

Callystomis pictus 1 - CMARF-UESC 1230.png

C. pictus
(Pictet, 1841)
Eastern BrazilSize: 25–29 cm (10–11 in) long, plus 27–32 cm (11–13 in) tail [9]

Habitat: Forest [10]

Diet: Vegetation [11]
 EN 


Unknown Decrease2.svg [10]

Genus Dactylomys Geoffroy, 1838 – three species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Amazon bamboo rat

Dactylomys dactylinus.JPG

D. dactylinus
(Desmarest, 1817)

Three subspecies
  • D. d. canescens
  • D. d. dactylinus
  • D. d. modestus
Northern South America
Dactylomys dactylinus range.svg
Size: About 31 cm (12 in) long, plus about 39 cm (15 in) tail [12]

Habitat: Forest [13]

Diet: Bamboo and other plants [14]
 LC 


Unknown Steady2.svg [13]

Bolivian bamboo rat


D. boliviensis
Anthony, 1920
Western South America
Dactylomys boliviensis range.svg
Size: 27–29 cm (11 in) long, plus about 41 cm (16 in) tail [12]

Habitat: Forest [15]

Diet: Bamboo and other plants [14]
 LC 


Unknown Steady2.svg [15]

Montane bamboo rat


D. peruanus
Allen, 1900
Bolivia and Peru
Dactylomys peruanus range.svg
Size: About 24 cm (9 in) long, plus 32 cm (13 in) tail [12]

Habitat: Forest [16]

Diet: Bamboo and other plants [14]
 DD 


Unknown Blue question mark (italic).svg [16]

Genus Diplomys Thomas, 1916 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Colombian soft-furred spiny rat

LoncheresCanicepsMintern.jpg

D. caniceps
(Günther, 1877)
Northern Colombia Size: 21–39 cm (8–15 in) long, plus 17–27 cm (7–11 in) tail [12]

Habitat: Forest [17]

Diet: Seeds and fruit [11]
 DD 


Unknown Blue question mark (italic).svg [17]

Rufous soft-furred spiny rat

Diplomys labilis.JPG

D. labilis
(Bangs, 1901)
Northwestern South America and Panama Size: About 34 cm (13 in) long, plus 20 cm (8 in) tail [12]

Habitat: Forest [18]

Diet: Seeds and fruit [11]
 LC 


Unknown Steady2.svg [18]

Genus Echimys F. Cuvier, 1809 – three species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Dark spiny tree-rat


E. saturnus
Thomas, 1928
Ecuador and PeruSize: 28–33 cm (11–13 in) long, plus 29–38 cm (11–15 in) tail [12]

Habitat: Forest [19]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [19]

Vieira's tree-rat


E. vieirai
Iack-Ximenes, Vivo, & Percequillo, 2005
Northern BrazilSize: About 24 cm (9 in) long, plus about 34 cm (13 in) tail [12]

Habitat: Forest [20]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [20]

White-faced spiny tree-rat

White-faced tree rat standing on log - DPLA - 5632760bf1692380e93a645b3e162dd4.jpg

E. chrysurus
(Zimmermann, 1780)
Northern South AmericaSize: 24–33 cm (9–13 in) long, plus 25–39 cm (10–15 in) tail [12]

Habitat: Forest [21]

Diet: Vegetation [11]
 LC 


Unknown Steady2.svg [21]

Genus Hoplomys J. A. Allen, 1908 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Armored rat

Hoplomys gymnurus1.jpg

H. gymnurus
(Thomas, 1897)
Central America and northwestern South America
Distribution of Hoplomys gymnurus.png
Size: 21–32 cm (8–13 in) long, plus 11–24 cm (4–9 in) tail [9]

Habitat: Forest [22]

Diet: Wide variety of vegetation, as well as insects [23]
 LC 


Unknown Steady2.svg [22]

Genus Isothrix Wagner, 1845 – six species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Barbara Brown's brush-tailed rat


I. barbarabrownae
Patterson & Velazco, 2006
Southern PeruSize: Unknown [24]

Habitat: Forest [25]

Diet: Vegetation [26]
 DD 


Unknown Blue question mark (italic).svg [25]

Orinoco brush-tailed rat


I. orinoci
Thomas, 1899
Northern South AmericaSize: 21–26 cm (8–10 in) long, plus 16–28 cm (6–11 in) tail [24]

Habitat: Forest [27]

Diet: Vegetation [26]
 DD 


Unknown Blue question mark (italic).svg [27]

Plain brush-tailed rat


I. pagurus
Wagner, 1845
Northern BrazilSize: 17–23 cm (7–9 in) long, plus 17–23 cm (7–9 in) tail [24]

Habitat: Forest [28]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [28]

Rio Negro brush-tailed rat


I. negrensis
Thomas, 1920
Northwestern BrazilSize: 20–29 cm (8–11 in) long, plus 18–27 cm (7–11 in) tail [24]

Habitat: Forest [29]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [29]

Sinnamary brush-tailed rat


I. sinnamariensis
Vié, Volobouev, Patton, & Granjon, 1996
Northern South AmericaSize: About 21 cm (8 in) long, plus about 26 cm (10 in) tail [24]

Habitat: Forest [30]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [30]

Yellow-crowned brush-tailed rat

Isothrix bistriata 74499273 (cropped).jpg

I. bistriata
Wagner, 1845
Western South AmericaSize: 21–27 cm (8–11 in) long, plus 21–27 cm (8–11 in) tail [24]

Habitat: Forest and grassland [31]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [31]

Genus Kannabateomys Jentink, 1891 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Atlantic bamboo rat

Atlantic Bamboo Rat, Porto Belo, SC, 88210-000, Brasil imported from iNaturalist photo 453361837.jpg

K. amblyonyx
Wagner, 1845

Two subspecies
  • K. a. amblyonyx
  • K. a. pallidior
Eastern South AmericaSize: 23–35 cm (9–14 in) long, plus 30–42 cm (12–17 in) tail [24]

Habitat: Forest [32]

Diet: Bamboo [33]
 LC 


Unknown Blue question mark (italic).svg [32]

Genus Lonchothrix Thomas, 1920 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Tuft-tailed spiny tree-rat


L. emiliae
Thomas, 1920
Northern BrazilSize: 15–22 cm (6–9 in) long, plus 15–23 cm (6–9 in) tail [24]

Habitat: Forest and grassland [34]

Diet: Vegetation [26]
 LC 


Unknown Steady2.svg [34]

Genus Makalata Husson, 1978 – three species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Brazilian spiny tree-rat

Makalata didelphoides.jpg

M. didelphoides
(Desmarest, 1817)
Northern South AmericaSize: 15–25 cm (6–10 in) long, plus 15–23 cm (6–9 in) tail [12]

Habitat: Forest [35]

Diet: Fruit and seeds [14]
 LC 


Unknown Blue question mark (italic).svg [35]

Dusky spiny tree-rat


M. obscura
(Wagner, 1840)
BrazilSize: Unknown [12]

Habitat: Unknown [36]

Diet: Fruit and seeds [14]
 DD 


Unknown Blue question mark (italic).svg [36]

Long-tailed armored tree-rat


M. macrura
(Wagner, 1842)
Northwestern South AmericaSize: 21–28 cm (8–11 in) long, plus 17–24 cm (7–9 in) tail [12]

Habitat: Forest [37]

Diet: Fruit and seeds [14]
 LC 


Unknown Blue question mark (italic).svg [37]

Genus Mesomys Wagner, 1845 – four species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Ferreira's spiny tree-rat

Mesomys hispidus female.jpg

M. hispidus
(Desmarest, 1817)
Northern South AmericaSize: 17–19 cm (7 in) long, plus 16–20 cm (6–8 in) tail [24]

Habitat: Forest [38]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [38]

Pará spiny tree-rat


M. stimulax
Thomas, 1911
Northern BrazilSize: 15–20 cm (6–8 in) long, plus 12–20 cm (5–8 in) tail [24]

Habitat: Forest [39]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [39]

Tufted-tailed spiny tree-rat


M. occultus
Patton, Silva, & Malcolm, 2000
Northwestern BrazilSize: About 17 cm (7 in) long, plus about 18 cm (7 in) tail [24]

Habitat: Forest [40]

Diet: Vegetation [26]
 LC 


Unknown Blue question mark (italic).svg [40]

Woolly-headed spiny tree-rat


M. leniceps
Thomas & St. Leger, 1926
Western PeruSize: About 18 cm (7 in) long, plus about 21 cm (8 in) tail [24]

Habitat: Forest [41]

Diet: Vegetation [26]
 DD 


Unknown Blue question mark (italic).svg [41]

Genus Myocastor Kerr, 1792 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Nutria

Myocastor coypus 02.jpg

M. coypus
(Molina, 1782)
Southern South America, and introduced in United States, Europe, and Japan
Nutria.svg
Size: 47–57 cm (19–22 in) long, plus 34–40 cm (13–16 in) tail [9]

Habitat: Inland wetlands [42]

Diet: Roots and aquatic plants [43]
 LC 


Unknown Decrease2.svg [42]

Genus Olallamys Emmons, 1988 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Greedy olalla rat


O. edax
(Thomas, 1916)
Western Venezuela Size: About 22 cm (9 in) long, plus about 34 cm (13 in) tail [12]

Habitat: Forest and shrubland [44]

Diet: Vegetation [33]
 DD 


Unknown Blue question mark (italic).svg [44]

White-tailed olalla rat

ThrinacodusAlbicaudaSmit.jpg

O. albicauda
(Günther, 1879)
ColombiaSize: 15–18 cm (6–7 in) long, plus 25–26 cm (10 in) tail [12]

Habitat: Forest [45]

Diet: Vegetation [33]
 DD 


Unknown Blue question mark (italic).svg [45]

Genus Pattonomys Emmons, 2005 – five species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Bare-tailed armored tree-rat


P. occasius
(Thomas, 1921)
EcuadorSize: About 2 cm (1 in) long, plus about 22 cm (9 in) tail [12]

Habitat: Forest [46]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [46]

Carriker's speckled tree-rat


P. carrikeri
(J. A. Allen, 1911)
Northern VenezuelaSize: About 22 cm (9 in) long, plus about 22 cm (9 in) tail [12]

Habitat: Forest and savanna [47]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [47]

Orinocoan speckled tree-rat


P. punctatus
Thomas, 1899
VenezuelaSize: About 27 cm (11 in) long, plus about 23 cm (9 in) tail [12]

Habitat: Forest [48]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [48]

Speckled spiny tree-rat


P. semivillosus
(Geoffroy, 1838)
ColombiaSize: 20–27 cm (8–11 in) long, plus 21–26 cm (8–10 in) tail [12]

Habitat: Forest and savanna [49]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [49]

Yellow speckled tree-rat


P. flavidus
Hollister, 1914
Northern VenezuelaSize: About 25 cm (10 in) long, plus about 25 cm (10 in) tail [12]

Habitat: Unknown [50]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [50]

Genus Phyllomys Lund, 1839 – thirteen species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Black-spined Atlantic tree-rat


P. nigrispinus
(Wagner, 1842)
Southern BrazilSize: 19–26 cm (7–10 in) long, plus 19–26 cm (7–10 in) tail [51]

Habitat: Forest [52]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [52]

Drab Atlantic tree-rat


P. dasythrix
Hensel, 1872
Southern BrazilSize: 18–20 cm (7–8 in) long, plus 20–22 cm (8–9 in) tail [51]

Habitat: Forest [53]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [53]

Giant Atlantic tree-rat

Curarao.jpeg

P. thomasi
(H. von Ihering, 1897)
Southern BrazilSize: 27–30 cm (11–12 in) long, plus 27–34 cm (11–13 in) tail [51]

Habitat: Forest [54]

Diet: Vegetation [11]
 EN 


Unknown Decrease2.svg [54]

Golden Atlantic tree-rat


P. blainvillii
(Jourdan, 1837)
Eastern BrazilSize: 17–22 cm (7–9 in) long, plus 19–27 cm (7–11 in) tail [12]

Habitat: Forest [55]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [55]

Kerr's Atlantic tree-rat


P. kerri
(Oliveira, 1950)
Southern BrazilSize: 19–23 cm (7–9 in) long, plus 20–22 cm (8–9 in) tail [51]

Habitat: Forest [56]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [56]

Long-furred Atlantic tree-rat


P. medius
(Thomas, 1909)
Southern BrazilSize: 22–23 cm (9 in) long, plus 22–26 cm (9–10 in) tail [51]

Habitat: Forest [57]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [57]

Lund's Atlantic tree-rat


P. lundi
Leite, 2003
Southern BrazilSize: 18–21 cm (7–8 in) long, plus about 20 cm (8 in) tail [51]

Habitat: Forest [58]

Diet: Vegetation [11]
 EN 


Unknown Decrease2.svg [58]

Mantiqueira Atlantic tree-rat


P. mantiqueirensis
Leite, 2003
Southern BrazilSize: About 22 cm (9 in) long, plus about 22 cm (9 in) tail [51]

Habitat: Forest [59]

Diet: Vegetation [11]
 CR 


Unknown Decrease2.svg [59]

Orange-brown Atlantic tree-rat


P. brasiliensis
Lund, 1840
Southern BrazilSize: About 21 cm (8 in) long, plus about 17 cm (7 in) tail [12]

Habitat: Unknown [60]

Diet: Vegetation [11]
 EN 


Unknown Decrease2.svg [60]

Pallid Atlantic tree-rat


P. lamarum
(Thomas, 1916)
Eastern BrazilSize: 18–23 cm (7–9 in) long, plus 18–23 cm (7–9 in) tail [51]

Habitat: Forest [61]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [61]

Rusty-sided Atlantic tree-rat


P. pattoni
Emmons, Leite, Kock, Costa, 2002
Eastern BrazilSize: 21–24 cm (8–9 in) long, plus 19–22 cm (7–9 in) tail [51]

Habitat: Forest [62]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [62]

Short-furred Atlantic tree-rat


P. unicolor
(Wagner, 1842)
Eastern BrazilSize: About 28 cm (11 in) long, plus about 20 cm (8 in) tail [51]

Habitat: Forest [63]

Diet: Vegetation [11]
 CR 


Unknown Decrease2.svg [63]

Southern Atlantic tree-rat


P. sulinus
Leite, Christoff, & Fagundes, 2008
Southern BrazilSize: 20–21 cm (8 in) long, plus 16–25 cm (6–10 in) tail [51]

Habitat: Forest [64]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [64]

Genus Proechimys Allen, 1899 – 22 species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Boyacá spiny rat


P. chrysaeolus
(Thomas, 1898)
ColombiaSize: 21–22 cm (8–9 in) long, plus 14–16 cm (6 in) tail [24]

Habitat: Forest [65]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [65]

Colombian spiny rat


P. canicollis
(Allen, 1899)
Colombia and VenezuelaSize: About 22 cm (9 in) long, plus about 17 cm (7 in) tail [9]

Habitat: Forest [67]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Steady2.svg [67]

Cuvier's spiny rat


P. cuvieri
Petter, 1978
Northern South AmericaSize: 21–23 cm (8–9 in) long, plus 14–17 cm (6–7 in) tail [24]

Habitat: Forest [68]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [68]

Gardner's spiny rat


P. gardneri
Silva, 1998
Bolivia and western BrazilSize: 15–21 cm (6–8 in) long, plus 8–16 cm (3–6 in) tail [24]

Habitat: Forest [69]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [69]

Goeldi's spiny rat


P. goeldii
Thomas, 1905
Central BrazilSize: 18–27 cm (7–11 in) long, plus 10–18 cm (4–7 in) tail [24]

Habitat: Forest [70]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Steady2.svg [70]

Guaira spiny rat


P. guairae
Thomas, 1901
Colombia and VenezuelaSize: 21–24 cm (8–9 in) long, plus 17–20 cm (7–8 in) tail [24]

Habitat: Forest [71]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [71]

Guyanan spiny-rat


P. hoplomyoides
Tate, 1939
Northern South AmericaSize: About 21 cm (8 in) long, plus about 15 cm (6 in) tail [24]

Habitat: Forest [72]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [72]

Guyenne spiny rat

Proechimys guyannensis.jpg

P. guyannensis
(Geoffroy, 1803)

Six subspecies
  • P. g. arabupu
  • P. g. arescens
  • P. g. cherriei
  • P. g. guyannensis
  • P. g. riparum
  • P. g. vacillator
Northern South AmericaSize: 18–23 cm (7–9 in) long, plus 11–19 cm (4–7 in) tail [24]

Habitat: Forest [73]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [73]

Kulina spiny rat


P. kulinae
Silva, 1998
Peru and western BrazilSize: 14–19 cm (6–7 in) long, plus 9–14 cm (4–6 in) tail [24]

Habitat: Forest [74]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [74]

Long-tailed spiny rat

Proechimys longicaudatus.jpg

P. longicaudatus
(Rengger, 1830)
Central South AmericaSize: 22–25 cm (9–10 in) long, plus 13–15 cm (5–6 in) tail [24]

Habitat: Forest [75]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [75]

Minca spiny rat


P. mincae
(Allen, 1899)
Northern ColombiaSize: 22–23 cm (9 in) long, plus 20–21 cm (8 in) tail [24]

Habitat: Forest [76]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [76]

Napo spiny rat


P. quadruplicatus
Hershkovitz, 1948
Northern South America
Proechimys quadruplicatus distribution (colored).png
Size: 21–29 cm (8–11 in) long, plus 12–20 cm (5–8 in) tail [9]

Habitat: Forest [77]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Steady2.svg [77]

O'Connell's spiny rat


P. oconnelli
Allen, 1913
ColombiaSize: About 25 cm (10 in) long, plus 18 cm (7 in) tail [9]

Habitat: Forest [78]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [78]

Pacific spiny rat


P. decumanus
(Thomas, 1899)
Ecuador and PeruSize: 26–30 cm (10–12 in) long, plus about 20 cm (8 in) tail [9]

Habitat: Forest [79]

Diet: Wide variety of vegetation [66]
 NT 


Unknown Decrease2.svg [79]

Patton's spiny rat


P. pattoni
Silva, 1998
Peru and western BrazilSize: 14–20 cm (6–8 in) long, plus 10–15 cm (4–6 in) tail [24]

Habitat: Forest [80]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [80]

Robert's spiny rat


P. roberti
Thomas, 1901
Central Brazil
Proechimys roberti distribution (colored).png
Size: 20–23 cm (8–9 in) long, plus about 16 cm (6 in) tail [24]

Habitat: Forest and savanna [81]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Steady2.svg [81]

Short-tailed spiny rat


P. brevicauda
(Günther, 1877)
Western South AmericaSize: 18–25 cm (7–10 in) long, plus 13–17 cm (5–7 in) tail [24]

Habitat: Forest [82]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [82]

Simons's spiny rat


P. simonsi
Thomas, 1900
Western South America
Proechimys simonsi distribution (colored).png
Size: 16–27 cm (6–11 in) long, plus 11–23 cm (4–9 in) tail [9]

Habitat: Forest [83]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [83]

Steere's spiny rat


P. steerei
Goldman, 1911
Western South America
Proechimys steerei distribution (colored).png
Size: 21–29 cm (8–11 in) long, plus 12–21 cm (5–8 in) tail [24]

Habitat: Forest [84]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [84]

Stiff-spine spiny rat


P. echinothrix
Silva, 1998
Western BrazilSize: 14–24 cm (6–9 in) long, plus 10–21 cm (4–8 in) tail [9]

Habitat: Forest [85]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [85]

Tome's spiny rat

Proechimys semispinosus 254496464.jpg

P. semispinosus
(Tomes, 1860)

Ten subspecies
  • P. s. burrus
  • P. s. calidior
  • P. s. centralis
  • P. s. colombianus
  • P. s. goldmani
  • P. s. ignotus
  • P. s. panamensis
  • P. s. rosa
  • P. s. rubellus
  • P. s. semispinosus
Central America and northwestern South America
Proechimys semispinosus distribution (gray).png
Size: About 29 cm (11 in) long, plus about 24 cm (9 in) tail [9]

Habitat: Forest [86]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Steady2.svg [86]

Trinidad spiny rat


P. trinitatus
(Allen & Chapman, 1893)
Northern Venezuela
Proechimys trinitatus distribution (gray).png
Size: 26–27 cm (10–11 in) long, plus 20–21 cm (8–8 in) tail [24]

Habitat: Forest [87]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [87]

Genus Santamartamys Emmons, 2005 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Red-crested tree-rat

Santamartamys rufodorsalis.jpg

S. rufodorsalis
(Allen, 1899)
Northern Colombia
Santamartamys rufodorsalis distribution.png
Size: About 19 cm (7 in) long, plus about 27 cm (11 in) tail [12]

Habitat: Unknown [88]

Diet: Seeds and fruit [11]
 CR 


1–50 Blue question mark (italic).svg [88]

Genus Thrichomys Trouessart, 1880 – four species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Common punaré

Thrichomys apereoides no Parque Nacional da Serra da Canastra.jpg

T. apereoides
(Lund, 1839)
Eastern Brazil
Thrichomys apereoides distribution (gray).png
Size: 19–21 cm (7–8 in) long, plus 16–18 cm (6–7 in) tail [9]

Habitat: Savanna and shrubland [89]

Diet: Seeds, fruit, cacti, and insects [8]
 LC 


Unknown Blue question mark (italic).svg [89]

Highlands punaré


T. inermis
Pictet, 1841
Eastern Brazil
Thrichomys inermis distribution (colored).png
Size: 18–22 cm (7–9 in) long, plus 11–20 cm (4–8 in) tail [9]

Habitat: Forest, savanna, and rocky areas [90]

Diet: Seeds, fruit, cacti, and insects [8]
 LC 


Unknown Blue question mark (italic).svg [90]

Paraguayan punaré


T. pachyurus
(Wagner, 1845)
Southern Brazil
Thrichomys pachyurus distribution (colored).png
Size: Unknown [9]

Habitat: Forest and savanna [91]

Diet: Seeds, fruit, cacti, and insects [8]
 LC 


Unknown Blue question mark (italic).svg [91]

Sao Lourenço punaré

Rabudo, Sitio Pau Preto, Potengi - CE, Brasil imported from iNaturalist photo 418722425.jpg

T. laurentius
Thomas, 1904
Eastern BrazilSize: 12–24 cm (5–9 in) long, plus 13–21 cm (5–8 in) tail [9]

Habitat: Forest [92]

Diet: Seeds, fruit, cacti, and insects [8]
 DD 


Unknown Blue question mark (italic).svg [92]

Genus Toromys Iack-Ximenes & Vivo & Percequillo, 2005 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Giant tree-rat


T. grandis
(Wagner, 1845)
Northern BrazilSize: About 30 cm (12 in) long, plus about 28 cm (11 in) tail [12]

Habitat: Forest [93]

Diet: Vegetation [11]
 LC 


Unknown Blue question mark (italic).svg [93]

Peruvian toro


T. rhipidura
(Thomas, 1928)
PeruSize: 21–26 cm (8–10 in) long, plus 18–21 cm (7–8 in) tail [12]

Habitat: Forest [94]

Diet: Vegetation [11]
 DD 


Unknown Blue question mark (italic).svg [94]

Subfamily Euryzygomatomyinae

Genus Clyomys Thomas, 1916 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Broad-headed spiny rat


C. laticeps
Thomas, 1909
Paraguay and southern BrazilSize: 15–29 cm (6–11 in) long, plus 4–9 cm (2–4 in) tail [6]

Habitat: Savanna [95]

Diet: Vegetation [8]
 LC 


Unknown Blue question mark (italic).svg [95]

Genus Euryzygomatomys Goeldi, 1901 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Fischer's guiara

Euryzygomatomys spinosus.jpg

E. spinosus
(G. von Waldheim, 1814)
Southern Brazil and northeastern Argentina Size: 16–20 cm (6–8 in) long, plus 6–7 cm (2–3 in) tail [6]

Habitat: Forest and grassland [96]

Diet: Vegetation [23]
 LC 


Unknown Blue question mark (italic).svg [96]

Genus Trinomys Thomas, 1921 – ten species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Dark-caped Atlantic spiny rat


T. mirapitanga
Lara, Patton, & Hingst-Zaher, 2002
Eastern Brazil
Trinomys mirapitanga distribution (colored).png
Size: About 22 cm (9 in) long, plus 18–20 cm (7–8 in) tail [9]

Habitat: Forest [97]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [97]

Elias's Atlantic spiny rat


T. eliasi
Pessoa & dos Reis, 1993
Southeastern Brazil
Trinomys eliasi distribution (colored).png
Size: 17–24 cm (7–9 in) long, plus 15–22 cm (6–9 in) tail [9]

Habitat: Forest and shrubland [98]

Diet: Wide variety of vegetation [66]
 NT 


Unknown Decrease2.svg [98]

Gracile Atlantic spiny rat


T. gratiosus
(Oliveira, 1948)

Two subspecies
  • T. g. bonafidei
  • T. g. gratiosus
Southeastern Brazil
Trinomys gratiosus distribution (colored).png
Size: 16–23 cm (6–9 in) long, plus 15–22 cm (6–9 in) tail [9]

Habitat: Forest [99]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [99]

Hairy Atlantic spiny rat


T. setosus
(Desmarest, 1817)

Three subspecies
  • T. s. denigratus
  • T. s. elegans
  • T. s. setosus
Southeastern Brazil
Trinomys setosus distribution (colored).png
Size: 18–23 cm (7–9 in) long, plus 16–23 cm (6–9 in) tail [9]

Habitat: Forest and savanna [100]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [100]

Ihering's Atlantic spiny rat


T. iheringi
(Thomas, 1911)
Eastern BrazilSize: 18–22 cm (7–9 in) long, plus 17–22 cm (7–9 in) tail [9]

Habitat: Forest [101]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Steady2.svg [101]

Moojen's Atlantic spiny rat


T. moojeni
Pessôa, Oliveira, & Reis, 1992
Southeastern Brazil
Trinomys moojeni distribution (colored).png
Size: 15–18 cm (6–7 in) long, plus 14–18 cm (6–7 in) tail [9]

Habitat: Forest and savanna [102]

Diet: Wide variety of vegetation [66]
 EN 


Unknown Decrease2.svg [102]

Soft-spined Atlantic spiny rat

Trinomys dimidiatus consuming jackfruit in Ilha Grande (Brazil).png

T. dimidiatus
(Günther, 1877)
Southeastern Brazil
Trinomys dimidiatus distribution (colored).png
Size: 17–24 cm (7–9 in) long, plus 15–22 cm (6–9 in) tail [9]

Habitat: Shrubland [103]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [103]

Spiked Atlantic spiny rat


T. paratus
Oliveira, 1948
Southeastern Brazil
Trinomys paratus distribution (colored).png
Size: 18–27 cm (7–11 in) long, plus 17–23 cm (7–9 in) tail [9]

Habitat: Forest [104]

Diet: Wide variety of vegetation [66]
 DD 


Unknown Blue question mark (italic).svg [104]

White-spined Atlantic spiny rat


T. albispinus
(Geoffroy, 1838)

Three subspecies
  • T. a. albispinus
  • T. a. minor
  • T. a. serotinus
Eastern Brazil
Trinomys albispinus distribution (gray).png
Size: 15–19 cm (6–7 in) long, plus 14–18 cm (6–7 in) tail [6]

Habitat: Forest, savanna, shrubland, grassland, and rocky areas [105]

Diet: Wide variety of vegetation [66]
 LC 


Unknown Blue question mark (italic).svg [105]

Yonenaga's Atlantic spiny rat


T. yonenagae
(Rocha, 1995)
Eastern Brazil
Trinomys yonenagae distribution (colored).png
Size: 14–20 cm (6–8 in) long, plus 16–22 cm (6–9 in) tail [9]

Habitat: Savanna [106]

Diet: Wide variety of vegetation [66]
 EN 


Unknown Decrease2.svg [106]

Subfamily Heteropsomyinae

Genus Boromys Miller, 1916 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Oriente cave rat

Boromys.offella.bmcz.jpg

B. offella
Miller, 1916
Cuba Size: Unknown [107]

Habitat: Unknown [108]

Diet: Unknown [1]
 EX 


0 Steady2.svg [108]

Torre's cave rat

Boromys.torrei.bmcz.jpg

B. torrei
Allen, 1917
CubaSize: Unknown [107]

Habitat: Unknown [109]

Diet: Unknown [1]
 EX 


0 Steady2.svg [109]

Genus Brotomys Miller, 1916 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Haitian edible rat


B. contractus
Miller, 1916
Haiti Size: Unknown [107]

Habitat: Unknown

Diet: Unknown [1]
 EX 


0 Steady2.svg

Hispaniolan edible rat


B. voratus
Miller, 1916
Dominican Republic Size: Unknown [107]

Habitat: Unknown [110]

Diet: Unknown [1]
 EX 


0 Steady2.svg [110]

Genus Heteropsomys Anthony, 1916 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Antillean cave rat


H. antillensis
Anthony, 1916
Puerto Rico Size: Unknown [107]

Habitat: Unknown

Diet: Unknown [1]
 EX 


0 Steady2.svg

Insular cave rat


H. insulans
Anthony, 1916
Puerto RicoSize: Unknown [107]

Habitat: Unknown [111]

Diet: Unknown [1]
 EX 


0 Steady2.svg [111]

References

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  54. 1 2 Leite, Y.; Loss, C. (2016). "Phyllomys thomasi". IUCN Red List of Threatened Species . 2016: e.T6989A22210762. doi: 10.2305/IUCN.UK.2016-2.RLTS.T6989A22210762.en .
  55. 1 2 Leite, Y.; Loss, C. (2016). "Phyllomys blainvillii". IUCN Red List of Threatened Species . 2016: e.T6977A22209935. doi: 10.2305/IUCN.UK.2016-2.RLTS.T6977A22209935.en .
  56. 1 2 Leite, Y.; Loss, C. (2016). "Phyllomys kerri". IUCN Red List of Threatened Species . 2016: e.T136682A22210378. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136682A22210378.en .
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  60. 1 2 Loss, C.; Leite, Y. (2016). "Phyllomys brasiliensis". IUCN Red List of Threatened Species . 2016: e.T6978A22209830. doi: 10.2305/IUCN.UK.2016-2.RLTS.T6978A22209830.en .
  61. 1 2 Leite, Y.; Loss, C. (2016). "Phyllomys lamarum". IUCN Red List of Threatened Species . 2016: e.T6982A22210201. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6982A22210201.en .
  62. 1 2 Loss, C.; Leite, Y. (2016). "Phyllomys pattoni". IUCN Red List of Threatened Species . 2016: e.T136801A22210466. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136801A22210466.en .
  63. 1 2 Loss, C.; Leite, Y. (2016). "Phyllomys unicolor". IUCN Red List of Threatened Species . 2016: e.T6990A22210643. doi: 10.2305/IUCN.UK.2016-3.RLTS.T6990A22210643.en .
  64. 1 2 Roach, N.; Naylor, L. (2016). "Phyllomys sulinus". IUCN Red List of Threatened Species . 2016: e.T47792718A47792815. doi: 10.2305/IUCN.UK.2016-2.RLTS.T47792718A47792815.en .
  65. 1 2 Weksler, M. (2016). "Proechimys chrysaeolus". IUCN Red List of Threatened Species . 2016: e.T18278A22209011. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18278A22209011.en .
  66. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 Nowak, p. 1689
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  68. 1 2 Catzeflis, F.; Patton, J. (2016). "Proechimys cuvieri". IUCN Red List of Threatened Species . 2016: e.T18279A22208920. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18279A22208920.en .
  69. 1 2 Patton, J. L.; Percequillo, A. (2016). "Proechimys gardneri". IUCN Red List of Threatened Species . 2016: e.T29466A22208352. doi: 10.2305/IUCN.UK.2016-3.RLTS.T29466A22208352.en .
  70. 1 2 Patton, J. (2016). "Proechimys goeldii". IUCN Red List of Threatened Species . 2016: e.T18282A22209354. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18282A22209354.en .
  71. 1 2 Roach, N.; Naylor, L. (2016). "Proechimys guairae". IUCN Red List of Threatened Species . 2016: e.T18284A22209221. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18284A22209221.en .
  72. 1 2 Patton, J.; Catzeflis, F. (2016). "Proechimys hoplomyoides". IUCN Red List of Threatened Species . 2016: e.T18287A22209472. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18287A22209472.en .
  73. 1 2 Catzeflis, F.; Patton, J. (2016). "Proechimys guyannensis". IUCN Red List of Threatened Species . 2016: e.T18277A22209103. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18277A22209103.en .
  74. 1 2 Patton, J.; Percequillo, A. (2016). "Proechimys kulinae". IUCN Red List of Threatened Species . 2016: e.T29464A22207750. doi: 10.2305/IUCN.UK.2016-2.RLTS.T29464A22207750.en .
  75. 1 2 Patton, J.; Bernal, N.; Patterson, B. (2016). "Proechimys longicaudatus". IUCN Red List of Threatened Species . 2016: e.T18289A22207395. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18289A22207395.en .
  76. 1 2 Weksler, M. (2016). "Proechimys mincae". IUCN Red List of Threatened Species . 2016: e.T18291A22207336. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18291A22207336.en .
  77. 1 2 Roach, N.; Naylor, L. (2016). "Proechimys quadruplicatus". IUCN Red List of Threatened Species . 2016: e.T18296A22207674. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18296A22207674.en .
  78. 1 2 Roach, N.; Naylor, L. (2016). "Proechimys oconnelli". IUCN Red List of Threatened Species . 2016: e.T18293A22207828. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18293A22207828.en .
  79. 1 2 Roach, N.; Naylor, L. (2016). "Proechimys decumanus". IUCN Red List of Threatened Species . 2016: e.T18280A22208822. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18280A22208822.en .
  80. 1 2 Patton, J.; Percequillo, A. (2016). "Proechimys pattoni". IUCN Red List of Threatened Species . 2016: e.T29465A22208196. doi: 10.2305/IUCN.UK.2016-2.RLTS.T29465A22208196.en .
  81. 1 2 Weksler, M. (2016). "Proechimys roberti". IUCN Red List of Threatened Species . 2016: e.T18294A22207886. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18294A22207886.en .
  82. 1 2 Patton, J.; Percequillo, A. (2016). "Proechimys brevicauda". IUCN Red List of Threatened Species . 2016: e.T18275A22208647. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18275A22208647.en .
  83. 1 2 Patton, J.; Percequillo, A. (2016). "Proechimys simonsi". IUCN Red List of Threatened Species . 2016: e.T18299A22208022. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18299A22208022.en .
  84. 1 2 Patton, J.; Percequillo, A. (2016). "Proechimys steerei". IUCN Red List of Threatened Species . 2016: e.T18300A22208111. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18300A22208111.en .
  85. 1 2 Patton, J.; Percequillo, A. (2016). "Proechimys echinothrix". IUCN Red List of Threatened Species . 2016: e.T29463A22207611. doi: 10.2305/IUCN.UK.2016-2.RLTS.T29463A22207611.en .
  86. 1 2 Roach, N.; Naylor, L. (2016). "Proechimys semispinosus". IUCN Red List of Threatened Species . 2016: e.T18297A22208264. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18297A22208264.en .
  87. 1 2 Emmons, L. (2016). "Proechimys trinitatis". IUCN Red List of Threatened Species . 2016: e.T18301A22208438. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18301A22208438.en .
  88. 1 2 Lacher, T.; Roach, N. (2017). "Santamartamys rufodorsalis". IUCN Red List of Threatened Species . 2017: e.T6664A22210948. doi: 10.2305/IUCN.UK.2017-2.RLTS.T6664A22210948.en .
  89. 1 2 Roach, N.; Naylor, L. (2016). "Thrichomys apereoides". IUCN Red List of Threatened Species . 2016: e.T21839A22206242. doi: 10.2305/IUCN.UK.2016-2.RLTS.T21839A22206242.en .
  90. 1 2 Roach, N.; Naylor, L. (2016). "Thrichomys inermis". IUCN Red List of Threatened Species . 2016: e.T136355A22206143. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136355A22206143.en .
  91. 1 2 Lacher, T. (2016). "Thrichomys pachyurus". IUCN Red List of Threatened Species . 2016: e.T136245A22206322. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136245A22206322.en .
  92. 1 2 Roach, N. (2016). "Thrichomys laurentius". IUCN Red List of Threatened Species . 2016: e.T90386381A90386384. doi: 10.2305/IUCN.UK.2016-2.RLTS.T90386381A90386384.en .
  93. 1 2 Cassola, F. (2016). "Toromys grandis". IUCN Red List of Threatened Species . 2016: e.T6981A22212728. doi: 10.2305/IUCN.UK.2016-2.RLTS.T6981A22212728.en .
  94. 1 2 Vivar, E.; Patterson, B. (2016). "Toromys rhipidurus". IUCN Red List of Threatened Species . 2016: e.T90386329A22206649. doi: 10.2305/IUCN.UK.2016-2.RLTS.T90386329A22206649.en .
  95. 1 2 Marinho-Filho, J. (2016). "Clyomys laticeps". IUCN Red List of Threatened Species . 2016: e.T4989A22205489. doi: 10.2305/IUCN.UK.2016-2.RLTS.T4989A22205489.en .
  96. 1 2 Catzeflis, F.; Patton, J.; Percequillo, A.; Weksler, M. (2016). "Euryzygomatomys spinosus". IUCN Red List of Threatened Species . 2016: e.T8418A22205855. doi: 10.2305/IUCN.UK.2016-2.RLTS.T8418A22205855.en .
  97. 1 2 Moura, R.; Patterson, B. (2016). "Trinomys mirapitanga". IUCN Red List of Threatened Species . 2016: e.T136305A22211934. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136305A22211934.en .
  98. 1 2 Roach, N.; Naylor, L. (2016). "Trinomys eliasi". IUCN Red List of Threatened Species . 2016: e.T136407A22211833. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136407A22211833.en .
  99. 1 2 Patterson, B. (2016). "Trinomys gratiosus". IUCN Red List of Threatened Species . 2016: e.T136221A22212027. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136221A22212027.en .
  100. 1 2 Bonvicino, C. R.; Catzeflis, F. (2016). "Trinomys setosus". IUCN Red List of Threatened Species . 2016: e.T18298A22212292. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18298A22212292.en .
  101. 1 2 Patterson, B. (2016). "Trinomys iheringi". IUCN Red List of Threatened Species . 2016: e.T18288A22212466. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18288A22212466.en .
  102. 1 2 Roach, N.; Naylor, L. (2016). "Trinomys moojeni". IUCN Red List of Threatened Species . 2016: e.T136543A22211597. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136543A22211597.en .
  103. 1 2 Bonvicino, C. R.; Catzeflis, F. (2016). "Trinomys dimidiatus". IUCN Red List of Threatened Species . 2016: e.T18281A22212372. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18281A22212372.en .
  104. 1 2 Patterson, B. (2016). "Trinomys paratus". IUCN Red List of Threatened Species . 2016: e.T136492A22212105. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136492A22212105.en .
  105. 1 2 Bonvicino, C. R.; Catzeflis, F.; Patton, J. L.; Percequillo, A.; Weksler, M. (2016). "Trinomys albispinus". IUCN Red List of Threatened Species . 2016: e.T18272A22211739. doi: 10.2305/IUCN.UK.2016-2.RLTS.T18272A22211739.en .
  106. 1 2 Roach, N.; Naylor, L. (2016). "Trinomys yonenagae". IUCN Red List of Threatened Species . 2016: e.T136414A22212204. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136414A22212204.en .
  107. 1 2 3 4 5 6 Chernasky; Motis; Burgin, p. 706
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  109. 1 2 Turvey, S.; Helgen, K. (2016). "Boromys torrei". IUCN Red List of Threatened Species . 2016: e.T2886A22206099. doi: 10.2305/IUCN.UK.2016-2.RLTS.T2886A22206099.en .
  110. 1 2 Turvey, S.; Helgen, K. (2016). "Brotomys voratus". IUCN Red List of Threatened Species . 2016: e.T3121A22205792. doi: 10.2305/IUCN.UK.2016-2.RLTS.T3121A22205792.en .
  111. 1 2 Turvey, S.; Helgen, K. (2016). "Heteropsomys insulans". IUCN Red List of Threatened Species . 2016: e.T10025A22205961. doi: 10.2305/IUCN.UK.2016-2.RLTS.T10025A22205961.en .

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