Heteromyidae is a family of small mammals in the order Rodentia and part of the Castorimorpha suborder. Members of this family are called heteromyids and include kangaroo rats, kangaroo mice, pocket mice, and spiny pocket mice. They are found in North America, Central America, and the northwest tip of South America, primarily in forests, shrublands, and deserts, though some species can be found in grasslands or rocky areas. They range in size from Merriam's pocket mouse, at 5 cm (2 in) plus a 4 cm (2 in) tail, to Nelson's spiny pocket mouse, at 17 cm (7 in) plus a 20 cm (8 in) tail. Heteromyids are omnivores and primarily eat seeds, leaves, and other vegetation, as well as grain and insects. Almost no heteromyids have population estimates, though four species—the giant kangaroo rat, Jaliscan spiny pocket mouse, Nelson's spiny pocket mouse, and Paraguaná spiny pocket mouse—are categorized as endangered species, while the San Quintin kangaroo rat is categorized as critically endangered with a population of fewer than fifty mature adults.
The sixty-three extant species of Heteromyidae are divided into three subfamilies: Dipodomyinae, containing twenty–two species of kangaroo rats and kangaroo mice in two genera; Heteromyinae, containing a single genus of fourteen spiny pocket mouse species; and Perognathinae, containing twenty-seven pocket mouse species in two genera. A few extinct prehistoric heteromyid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed. [1]
Conservation status | |
---|---|
EX | Extinct (0 species) |
EW | Extinct in the wild (0 species) |
CR | Critically endangered (1 species) |
EN | Endangered (4 species) |
VU | Vulnerable (5 species) |
NT | Near threatened (3 species) |
LC | Least concern (49 species) |
Other categories | |
DD | Data deficient (1 species) |
NE | Not evaluated (0 species) |
The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the heteromyid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Heteromyidae is a family consisting of sixty-three species in five genera. These genera are divided between three subfamilies: Dipodomyinae, containing the kangaroo rats and kangaroo mice; Heteromyinae, or the spiny pocket mice; and Perognathinae, or the pocket mice.
Family Heteromyidae
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. [3]
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Agile kangaroo rat | D. agilis Gambel, 1848 Two subspecies
| Western United States | Size: 10–13 cm (4–5 in) long, plus 17–20 cm (7–8 in) tail [4] Habitat: Forest and shrubland [5] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Banner-tailed kangaroo rat | D. spectabilis Merriam, 1890 Six subspecies
| Southern United States and Mexico | Size: 13–15 cm (5–6 in) long, plus 18–21 cm (7–8 in) tail [4] Habitat: Desert [7] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | NT
|
California kangaroo rat | D. californicus Merriam, 1890 Three subspecies
| Western United States | Size: 10–13 cm (4–5 in) long, plus 15–22 cm (6–9 in) tail [4] Habitat: Forest, shrubland, and grassland [8] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Chisel-toothed kangaroo rat | D. microps (Merriam, 1904) Thirteen subspecies
| Western United States | Size: 11–12 cm (4–5 in) long, plus 13–18 cm (5–7 in) tail [9] Habitat: Shrubland, rocky areas, and desert [10] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Desert kangaroo rat | D. deserti Stephens, 1887 Four subspecies
| Western United States and western Mexico | Size: 13–16 cm (5–6 in) long, plus 19–21 cm (7–8 in) tail [4] Habitat: Desert [11] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Dulzura kangaroo rat | D. simulans (Merriam, 1904) Two subspecies
| Western United States and western Mexico | Size: 11–13 cm (4–5 in) long, plus 16–19 cm (6–7 in) tail [4] Habitat: Forest, shrubland, grassland, and desert [12] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Fresno kangaroo rat | D. nitratoides Merriam, 1894 Three subspecies
| Western United States | Size: 7–12 cm (3–5 in) long, plus 13–15 cm (5–6 in) tail [4] Habitat: Shrubland, grassland, and desert [13] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | VU
|
Giant kangaroo rat | D. ingens (Merriam, 1904) | Western United States | Size: 13–15 cm (5–6 in) long, plus 17–20 cm (7–8 in) tail [4] Habitat: Savanna and grassland [14] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | EN
|
Gulf Coast kangaroo rat | D. compactus True, 1889 Two subspecies
| Southern United States | Size: 9–12 cm (4–5 in) long, plus 10–14 cm (4–6 in) tail [4] Habitat: Savanna, shrubland, grassland, and coastal marine [15] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Heermann's kangaroo rat | D. heermanni Conte, 1853 Nine subspecies
| Western United States | Size: 9–12 cm (4–5 in) long, plus 16–20 cm (6–8 in) tail [4] Habitat: Forest, savanna, shrubland, grassland, and desert [16] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Merriam's kangaroo rat | D. merriami Mearns, 1890 Nineteen subspecies
| Southwestern United States and Mexico![]() | Size: 7–10 cm (3–4 in) long, plus 12–19 cm (5–7 in) tail [4] Habitat: Shrubland and desert [17] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Narrow-faced kangaroo rat | D. venustus (Merriam, 1904) Three subspecies
| Western United States | Size: 11–13 cm (4–5 in) long, plus 17–20 cm (7–8 in) tail [4] Habitat: Forest and shrubland [18] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Nelson's kangaroo rat | D. nelsoni Merriam, 1907 | Northern Mexico | Size: 12–14 cm (5–6 in) long, plus 12–20 cm (5–8 in) tail [4] Habitat: Shrubland and desert [19] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Ord's kangaroo rat | D. ordii Woodhouse, 1853 32 subspecies
| Western North America![]() | Size: 7–16 cm (3–6 in) long, plus about 13 cm (5 in) tail [4] Habitat: Shrubland and grassland [20] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Ornate kangaroo rat | D. ornatus Merriam, 1894 | Central Mexico![]() | Size: 10–11 cm (4 in) long, plus 15–20 cm (6–8 in) tail [4] Habitat: Desert [21] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Panamint kangaroo rat | D. panamintinus (Merriam, 1894) Five subspecies
| Western United States | Size: About 12 cm (5 in) long, plus about 17 cm (7 in) tail [9] Habitat: Savanna, shrubland, and desert [22] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
Phillips's kangaroo rat | D. phillipsii Gray, 1841 Four subspecies
| Central Mexico | Size: 8–12 cm (3–5 in) long, plus 14–19 cm (6–7 in) tail [4] Habitat: Shrubland and desert [23] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | LC
|
San Quintin kangaroo rat | D. gravipes Huey, 1925 | Western Mexico | Size: 12–14 cm (5–6 in) long, plus 16–18 cm (6–7 in) tail [4] Habitat: Desert [24] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | CR
|
Stephens's kangaroo rat | D. stephensi (Merriam, 1907) | Western United States | Size: 11–12 cm (4–5 in) long, plus 16–18 cm (6–7 in) tail [9] Habitat: Grassland and shrubland [25] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | VU
|
Texas kangaroo rat | D. elator Merriam, 1894 | Southern United States | Size: 9–14 cm (4–6 in) long, plus 16–20 cm (6–8 in) tail [4] Habitat: Savanna and desert [26] Diet: Seeds, as well as fruit, leaves, stems, buds, and insects [6] | VU
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Dark kangaroo mouse | M. megacephalus Merriam, 1891 Thirteen subspecies
| Western United States | Size: About 7 cm (3 in) long, plus 6–11 cm (2–4 in) tail [4] Habitat: Shrubland and desert [27] Diet: Grains and insects [6] | LC
|
Pale kangaroo mouse | M. pallidus Merriam, 1901 Five subspecies
| Western United States | Size: About 7 cm (3 in) long, plus 7–10 cm (3–4 in) tail [4] Habitat: Desert [28] Diet: Grains and insects [6] | LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Desmarest's spiny pocket mouse | H. desmarestianus Gray, 1868 Twelve subspecies
| Southern Mexico, Central America, and northwestern South America | Size: 13–14 cm (5–6 in) long, plus 14–15 cm (6 in) tail [29] Habitat: Forest [30] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Ecuadoran spiny pocket mouse | H. teleus Anderson & Jarrín-Valladares, 2002 | Ecuador | Size: About 14 cm (6 in) long, plus about 15 cm (6 in) tail [29] Habitat: Forest [32] Diet: Seeds, leaves, twigs, and succulent plants [31] | VU
|
Gaumer's spiny pocket mouse | H. gaumeri Allen & Chapman, 1897 | Eastern Mexico and northern Central America | Size: 12–13 cm (5 in) long, plus 14–16 cm (6 in) tail [29] Habitat: Forest [33] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Jaliscan spiny pocket mouse | H. spectabilis (Genoways, 1971) | Central Mexico | Size: 10–11 cm (4 in) long, plus 12–15 cm (5–6 in) tail [29] Habitat: Forest [34] Diet: Seeds, leaves, twigs, and succulent plants [31] | EN
|
Mexican spiny pocket mouse | H. irroratus Gray, 1868 Seven subspecies
| Southern United States and Mexico![]() | Size: 11–13 cm (4–5 in) long, plus 9–17 cm (4–7 in) tail [29] Habitat: Shrubland [35] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Mountain spiny pocket mouse | H. oresterus Harris Jr., 1932 | Costa Rica | Size: 14–16 cm (6 in) long, plus 16–18 cm (6–7 in) tail [29] Habitat: Forest [36] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Nelson's spiny pocket mouse | H. nelsoni Merriam, 1902 | Eastern Mexico and Guatemala | Size: 15–17 cm (6–7 in) long, plus 18–20 cm (7–8 in) tail [29] Habitat: Forest [37] Diet: Seeds, leaves, twigs, and succulent plants [31] | EN
|
Overlook spiny pocket mouse | H. catopterius Anderson & Gutiérrez, 2009 | Venezuela | Size: About 13 cm (5 in) long, plus about 17 cm (7 in) tail [29] Habitat: Forest [38] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Painted spiny pocket mouse | H. pictus Thomas, 1893 Four subspecies
| Mexico![]() | Size: About 11 cm (4 in) long, plus 9–17 cm (4–7 in) tail [29] Habitat: Forest and shrubland [39] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Panamanian spiny pocket mouse | H. adspersus Peters, 1874 | Panama ![]() | Size: 12–13 cm (5 in) long, plus 10–15 cm (4–6 in) tail [29] Habitat: Forest and shrubland [40] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Paraguaná spiny pocket mouse | H. oasicus Anderson, 2003 | Northern Venezuela | Size: About 12 cm (5 in) long, plus about 12 cm (5 in) tail [29] Habitat: Forest [41] Diet: Seeds, leaves, twigs, and succulent plants [31] | EN
|
Salvin's spiny pocket mouse | H. salvini Thomas, 1893 Three subspecies
| Eastern Mexico and Central America![]() | Size: 10–12 cm (4–5 in) long, plus 8–16 cm (3–6 in) tail [29] Habitat: Forest [42] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Southern spiny pocket mouse | H. australis Thomas, 1901 Three subspecies
| Panama and northwestern South America | Size: 12–13 cm (5 in) long, plus 13–14 cm (5–6 in) tail [29] Habitat: Forest [43] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Trinidad spiny pocket mouse | H. anomalus (Thompson, 1815) Four subspecies
| Northern South America | Size: 12–14 cm (5–6 in) long, plus 15–16 cm (6 in) tail [29] Habitat: Forest [44] Diet: Seeds, leaves, twigs, and succulent plants [31] | LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Bailey's pocket mouse | C. baileyi (Merriam, 1894) Two subspecies
| Southwestern United States and Mexico | Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail [45] Habitat: Desert [46] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Baja pocket mouse | C. rudinoris (Elliot, 1903) Six subspecies
| Western United States and western Mexico | Size: 8–10 cm (3–4 in) long, plus 10–13 cm (4–5 in) tail [45] Habitat: Rocky areas and desert [48] Diet: Seeds, as well as vegetation and insects [47] | LC
|
California pocket mouse | C. californicus (Merriam, 1889) Eight subspecies
| Western United States and western Mexico | Size: 8–10 cm (3–4 in) long, plus 10–15 cm (4–6 in) tail [45] Habitat: Shrubland, grassland, and desert [49] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Cerralvo Island pocket mouse | C. siccus (Osgood, 1907) | Western Mexico | Size: About 7 cm (3 in) long, plus about 9 cm (4 in) tail [45] Habitat: Shrubland [50] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Chihuahuan pocket mouse | C. eremicus (Mearns, 1898) Two subspecies
| Southern United States and Mexico | Size: 7–9 cm (3–4 in) long, plus 7–10 cm (3–4 in) tail [45] Habitat: Shrubland and desert [51] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Dalquest's pocket mouse | C. ammophilus Osgood, 1907 | Western Mexico | Size: About 7 cm (3 in) long, plus about 9 cm (4 in) tail [45] Habitat: Desert [52] Diet: Seeds, as well as vegetation and insects [47] | NT
|
Desert pocket mouse | C. penicillatus (Woodhouse, 1852) Six subspecies
| Southwestern United States and Mexico | Size: 7–9 cm (3–4 in) long, plus 8–13 cm (3–5 in) tail [45] Habitat: Shrubland and desert [53] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Goldman's pocket mouse | C. goldmani (Osgood, 1900) | Western Mexico | Size: About 8 cm (3 in) long, plus about 11 cm (4 in) tail [4] Habitat: Forest and shrubland [54] Diet: Seeds, as well as vegetation and insects [47] | NT
|
Hispid pocket mouse | C. hispidus (Baird, 1858) Four subspecies
| United States and Mexico | Size: 9–12 cm (4–5 in) long, plus 9–12 cm (4–5 in) tail [45] Habitat: Shrubland and grassland [55] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Lined pocket mouse | C. lineatus (Dalquest, 1951) | Central Mexico | Size: About 7 cm (3 in) long, plus about 9 cm (4 in) tail [4] Habitat: Desert [56] Diet: Seeds, as well as vegetation and insects [47] | DD
|
Little desert pocket mouse | C. arenarius (Merriam, 1894) Eleven subspecies
| Western Mexico | Size: 6–7 cm (2–3 in) long, plus 8–9 cm (3–4 in) tail [45] Habitat: Desert [57] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Long-tailed pocket mouse | C. formosus (Merriam, 1889) Seven subspecies
| Western United States and western Mexico | Size: 7–9 cm (3–4 in) long, plus 8–13 cm (3–5 in) tail [45] Habitat: Shrubland, rocky areas, and desert [58] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Narrow-skulled pocket mouse | C. artus (Osgood, 1900) | Western Mexico | Size: 8–10 cm (3–4 in) long, plus 9–10 cm (4 in) tail [4] Habitat: Desert [59] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Nelson's pocket mouse | C. nelsoni (Merriam, 1894) Two subspecies
| Southern United States and Mexico | Size: 7–9 cm (3–4 in) long, plus 8–12 cm (3–5 in) tail [45] Habitat: Shrubland, rocky areas, and desert [60] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Rock pocket mouse | C. intermedius (Merriam, 1889) Eight subspecies
| Southwestern United States and Mexico | Size: 7–8 cm (3 in) long, plus 8–12 cm (3–5 in) tail [45] Habitat: Grassland, rocky areas, and desert [61] Diet: Seeds, as well as vegetation and insects [47] | LC
|
San Diego pocket mouse | C. fallax (Merriam, 1889) Six subspecies
| Western United States and western Mexico | Size: 8–9 cm (3–4 in) long, plus 8–12 cm (3–5 in) tail [45] Habitat: Shrubland and desert [62] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Sinaloan pocket mouse | C. pernix (Allen, 1898) Two subspecies
| Western Mexico | Size: About 8 cm (3 in) long, plus about 8 cm (3 in) tail [45] Habitat: Shrubland [63] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Spiny pocket mouse | C. spinatus (Merriam, 1889) Eighteen subspecies
| Western United States and western Mexico | Size: 7–10 cm (3–4 in) long, plus 8–13 cm (3–5 in) tail [45] Habitat: Rocky areas and desert [64] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Arizona pocket mouse | P. amplus Osgood, 1900 Four subspecies
| Southwestern United States and western Mexico | Size: About 7 cm (3 in) long, plus about 8 cm (3 in) tail [29] Habitat: Forest, grassland, and desert [65] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Great Basin pocket mouse | P. parvus (Peale, 1848) Twelve subspecies
| Western United States and southwestern Canada![]() | Size: Unknown [45] Habitat: Forest, shrubland, grassland, and desert [66] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Little pocket mouse | P. longimembris (Coues, 1875) Sixteen subspecies
| Western United States and western Mexico | Size: About 6 cm (2 in) long, plus about 7 cm (3 in) tail [29] Habitat: Shrubland, grassland, and desert [67] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Merriam's pocket mouse | P. merriami Allen, 1892 Two subspecies
| Southern United States and northern Mexico | Size: 5–6 cm (2 in) long, plus 4–7 cm (2–3 in) tail [29] Habitat: Shrubland, grassland, and desert [68] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Olive-backed pocket mouse | P. fasciatus Wied-Neuwied, 1839 Two subspecies
| Northern-central United States and southern-central Canada![]() | Size: About 7 cm (3 in) long, plus about 7 cm (3 in) tail [29] Habitat: Shrubland and grassland [69] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Plains pocket mouse | P. flavescens Merriam, 1889 Eight subspecies
| Central United States and northern Mexico | Size: About 7 cm (3 in) long, plus about 6 cm (2 in) tail [45] Habitat: Shrubland, grassland, and desert [70] Diet: Seeds, as well as vegetation and insects [47] | LC
|
San Joaquin pocket mouse | P. inornatus Merriam, 1889 Three subspecies
| Western United States | Size: About 7 cm (3 in) long, plus about 8 cm (3 in) tail [29] Habitat: Savanna, grassland, and desert [71] Diet: Seeds, as well as vegetation and insects [47] | LC
|
Silky pocket mouse | P. flavus Baird, 1855 Fourteen subspecies
| Southern-central United States and Mexico![]() | Size: About 6 cm (2 in) long, plus about 5 cm (2 in) tail [29] Habitat: Shrubland, grassland, and desert [72] Diet: Seeds, as well as vegetation and insects [47] | LC
|
White-eared pocket mouse | P. alticolus Rhoads, 1894 Two subspecies
| Western United States | Size: 7–8 cm (3 in) long, plus 7–9 cm (3–4 in) tail [45] Habitat: Forest, shrubland, grassland, and desert [73] Diet: Seeds, as well as vegetation and insects [47] | VU
|