List of cingulates

Last updated

Nine-banded armadillo (Dasypus novemcinctus) Nine-banded Armadillo.jpg
Nine-banded armadillo (Dasypus novemcinctus)

Cingulata is an order of armored placental mammals. Members of this order are called cingulates, or colloquially, armadillos. They are primarily found in South America, though the northern naked-tailed armadillo is found mainly in Central America and the nine-banded armadillo has a range extending into North America. They are generally found in forests, but also savannas, shrublands, and grasslands. They all follow a similar body plan, and range in size from the pink fairy armadillo, at 11 cm (4 in) plus a 2 cm (1 in) tail, to the giant armadillo, at 100 cm (39 in) plus a 50 cm (20 in) tail. No population estimates have been made for any cingulate species, though the giant armadillo and the Brazilian three-banded armadillo are categorized as vulnerable species.

Contents

The twenty-two extant species of Cingulata are divided into two families: Dasypodidae, containing a single genus of nine species in the subfamily Dasypodinae, and Chlamyphoridae, containing thirteen species split between the two genera in the subfamily Chlamyphorinae, three in the subfamily Euphractinae, and three in the subfamily Tolypeutinae. Prior to 2016, all four subfamilies were included in Dasypodidae, with Chlamyphoridae containing only extinct species of glyptodonts. [1] [2] Over one hundred extinct Cingulata species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed. [3]

Conventions

IUCN Red List categories
Conservation status
 EX  Extinct (0 species)
 EW  Extinct in the wild (0 species)
 CR  Critically Endangered (0 species)
 EN  Endangered (0 species)
 VU  Vulnerable (2 species)
 NT  Near threatened (5 species)
 LC  Least concern (8 species)
Other categories
 DD  Data deficient (5 species)
 NE  Not evaluated (2 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the cingulate's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "Dagger-14-plain.png".

Classification

The order Cingulata consists of two families, Dasypodidae and Chlamyphoridae. Dasypodidae contains nine species in a single genus, while Chlamyphoridae contains thirteen species in eight genera, divided into three subfamilies. Many of these species are further subdivided into subspecies. This does not include hybrid species or extinct prehistoric species.

Family Dasypodidae

Family Chlamyphoridae

Cingulata   
Dasypodidae   
Dasypodinae

Dasypus

Chlamyphoridae   
Euphractinae   

Euphractus

Chaetophractus

Zaedyus

Chlamyphorinae   

Calyptophractus

Chlamyphorus

Tolypeutinae   

Priodontes

Cabassous

Tolypeutes

Cingulates

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis. [4]

Dasypodidae

Subfamily Dasypodinae

Genus Dasypus Linnaeus, 1758 – nine species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
East Amazonian long-nosed armadillo D. beniensis
Lönnberg, 1942
North-central South AmericaSize: 51–58 cm (20–23 in) long, plus 33–48 cm (13–19 in) tail [5]

Habitat: Forest [6]

Diet: Insects [5]
 NE 


Unknown Blue question mark (italic).svg [6]

Greater long-nosed armadillo

Kappler Gurteltier (gesamt Ansicht).jpg

D. kappleri
Krauss, 1862
Northeastern South AmericaSize: 51–58 cm (20–23 in) long, plus 33–48 cm (13–19 in) tail [7]

Habitat: Forest [6]

Diet: Insects [7]
 LC 


Unknown Blue question mark (italic).svg [6]

Hairy long-nosed armadillo

Dasypus pilosus - Feijo et al 2018 holotype.tif

D. pilosus
(Fitzinger, 1856)
Western South America
Hairy Long-nosed Armadillo area.png
Size: 32–44 cm (13–17 in) long, plus 23–31 cm (9–12 in) tail [8]

Habitat: Forest [9]

Diet: Insects [8]
 NE 


Unknown Blue question mark (italic).svg [9]

Llanos long-nosed armadillo

Dasypus sabanicola - Feijo et al 2018 holotype.tif

D. sabanicola
Mondolfi, 1968
Northern South America
Llanos Long-nosed Armadillo area.png
Size: 25–31 cm (10–12 in) long, plus 17–21 cm (7–8 in) tail [10]

Habitat: Forest [11]

Diet: Termites, as well as ants, beetles, and worms [10]
 NT 


Unknown Decrease2.svg [11]

Nine-banded armadillo

Florida-015.jpg

D. novemcinctus
Linnaeus, 1758

Six subspecies
  • D. n. aequatorialis
  • D. n. fenestratus
  • D. n. hoplites
  • D. n. mexianae
  • D. n. mexicanus
  • D. n. novemcinctus
Central and northern South America, and central, southern, and eastern North America
Common Long-nosed Armadillo area.png
Size: 35–57 cm (14–22 in) long, plus 24–45 cm (9–18 in) tail [12]

Habitat: Forest, savanna, shrubland, and grassland [13]

Diet: Omnivorous, including invertebrates, birds, fruit, and roots [12]
 LC 


Unknown Steady2.svg [13]

Seven-banded armadillo

Dasypus septemcinctus hybridus (also named Dasypus hybridus), at Valizas, Uruguay.jpg

D. septemcinctus
Linnaeus, 1758
Eastern South America
Brazilian Lesser Long-nosed Armadillo area.png
Size: 24–31 cm (9–12 in) long, plus 12–17 cm (5–7 in) tail [14]

Habitat: Forest, savanna, and grassland [15]

Diet: Insects, seeds, and other plant material [14]
 LC 


Unknown Blue question mark (italic).svg [15]

Southern long-nosed armadillo

Dasypus hybridus - Museo Civico di Storia Naturale Giacomo Doria - Genoa, Italy - DSC03055.JPG

D. hybridus
Desmarest, 1804
Southeastern South America
Southern Long-nosed Armadillo area.png
Size: 26–31 cm (10–12 in) long, plus 15–19 cm (6–7 in) tail [16]

Habitat: Forest, savanna, shrubland, and grassland [17]

Diet: Ants, termites, and beetles, as well as plant material and small vertebrates [16]
 NT 


Unknown Decrease2.svg [17]

West Amazonian long-nosed armadillo D. pastasae
(Thomas, 1901)
Northwestern South AmericaSize: 51–58 cm (20–23 in) long, plus 33–48 cm (13–19 in) tail [18]

Habitat: Forest [6]

Diet: Insects [18]
 DD 


Unknown Blue question mark (italic).svg [6]

Yepes's mulita

Dasypus mazzai - Feijo et al 2018 holotype.tif

D. mazzai
Yepes, 1933
South-central South America
Yepes's Mulita area.png
Size: About 31 cm (12 in) long, plus 18–23 cm (7–9 in) tail [19]

Habitat: Forest [20]

Diet: Believed to be omnivorous with a preference for insects [19]
 DD 


Unknown Blue question mark (italic).svg [20]

Chlamyphoridae

Subfamily Chlamyphorinae

Genus Calyptophractus Fitzinger, 1871 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Greater fairy armadillo

Burmeister-Gurtelmull2.png

C. retusus
(Burmeister, 1863)
Central South America
Greater Fairy Armadillo area.png
Size: 14–18 cm (6–7 in) long, plus 4 cm (2 in) tail [21]

Habitat: Savanna, shrubland, and desert [22]

Diet: Insects, worms, snails, roots, and small seeds [21]
 DD 


Unknown Blue question mark (italic).svg [22]

Genus Chlamyphorus Harlan, 1825 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Pink fairy armadillo

Pink Fairy Armadillo (Chlamyphorus truncatus).jpg

C. truncatus
Harlan, 1825
Southern South America
Lesser Fairy Armadillo area.png
Size: 11–15 cm (4–6 in) long, plus 2–3 cm (1–1 in)tail [23] [24]

Habitat: Savanna, shrubland, grassland, and desert [25]

Diet: insects, worms and snails, as well as possibly plants [23]
 DD 


Unknown Blue question mark (italic).svg [25]

Subfamily Euphractinae

Genus Chaetophractus Fitzinger, 1871 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Big hairy armadillo

Chaetophractus villosus (Wroclaw zoo).JPG

C. villosus
(Desmarest, 1804)
Southern South America
Large Hairy Armadillo area.png
Size: 22–40 cm (9–16 in) long, plus 9–17 cm (4–7 in) tail [12]

Habitat: Forest, savanna, shrubland, grassland, and desert [26]

Diet: Omnivorous, including insects, invertebrates, small vertebrates, plants, and carrion [27]
 LC 


Unknown Steady2.svg [26]

Screaming hairy armadillo

Chaetophractus vellerosus - Swedish Museum of Natural History - Stockholm, Sweden - DSC00653.JPG

C. vellerosus
(Gray, 1865)

Two subspecies
  • C. v. pannosus
  • C. v. vellerosus
Southern South America
Screaming Hairy Armadillo area.png
Size: 20–30 cm (8–12 in) long, plus tail [28]

Habitat: Savanna, shrubland, grassland, and desert [29]

Diet: Beetles, butterfly larvae, plants, and small vertebrates [29]
 LC 


Unknown Steady2.svg [29]

Genus Euphractus Wagler, 1830 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Six-banded armadillo

Six-banded armadillo (Euphractus sexcinctus).JPG

E. sexcinctus
(Linnaeus, 1758)

Five subspecies
  • E. s. boliviae
  • E. s. flavimanus
  • E. s. setosus
  • E. s. sexcinctus
  • E. s. tucumanus
Central and eastern South America
Six-banded Armadillo area.png
Size: 40–50 cm (16–20 in) long, plus 20–25 cm (8–10 in) tail [30]

Habitat: Forest, savanna, shrubland, and grassland [31]

Diet: Carrion, small vertebrates, insects, spiders, bird eggs, and plants [30]
 LC 


Unknown Steady2.svg [31]

Genus Zaedyus Ameghino, 1889 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Pichi

Zaedyus pichiy - Museo Civico di Storia Naturale Giacomo Doria - Genoa, Italy - DSC03056.JPG

Z. pichiy
(Desmarest, 1804)

Two subspecies
  • Z. p. caurinus
  • Z. p. pichiy
Southern South America
Pichi area.png
Size: 26–34 cm (10–13 in) long, plus 10–12 cm (4–5 in) tail [12]

Habitat: Shrubland, grassland, and desert [32]

Diet: Insects, worms, and other invertebrates, as well as carrion [12]
 NT 


Unknown Decrease2.svg [32]

Subfamily Tolypeutinae

Genus Cabassous McMurtrie, 1831 – four species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Chacoan naked-tailed armadillo

P02-03-12 10.00 1.jpg

C. chacoensis
Wetzel, 1980
South-central South America
Chacoan Naked-tailed Armadillo area.png
Size: 30–35 cm (12–14 in) long, plus 9–10 cm (4–4 in) tail [33]

Habitat: Forest, savanna, and shrubland [34]

Diet: Ants and termites, as well as seeds and fruit [33]
 NT 


Unknown Blue question mark (italic).svg [34]

Greater naked-tailed armadillo

Cabassous tatouay at University of Pisa's Natural History Museum.jpg

C. tatouay
(Desmarest, 1804)
Eastern South America
Greater Naked-tailed Armadillo area.png
Size: 36–49 cm (14–19 in) long, plus 15–20 cm (6–8 in) tail [35]

Habitat: Forest and grassland [36]

Diet: Ants and termites [37]
 LC 


Unknown Blue question mark (italic).svg [36]

Northern naked-tailed armadillo

Cabassous centralis (Miller, 1899) head from side.png

C. centralis
(Miller, 1899)
Northern South America and Central America
Northern Naked-tailed Armadillo area.png
Size: 30–40 cm (12–16 in) long, plus 5–7 cm (2–3 in) tail [12]

Habitat: Forest, savanna, and shrubland [38]

Diet: Termites and ants [12]
 DD 


Unknown Blue question mark (italic).svg [38]

Southern naked-tailed armadillo

Nacktschwanzgurteltier (Cabassous unicinctus).jpg

C. unicinctus
(Linnaeus, 1758)

Two subspecies
  • C. u. squamicaudis
  • C. u. unicinctus
Northern and central South America
Cabasu area.png
Size: 35–44 cm (14–17 in) long, plus 16–20 cm (6–8 in) tail [39]

Habitat: Forest, shrubland, and grassland [40]

Diet: Ants and termites [39]
 LC 


Unknown Blue question mark (italic).svg [40]

Genus Priodontes F. Cuvier, 1825 – one species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Giant armadillo

Giant armadillo.jpg

P. maximus
(Kerr, 1792)
Northern and central South America
Giant Armadillo area.png
Size: 75–100 cm (30–39 in) long, plus 50 cm (20 in) tail [41]

Habitat: Forest, savanna, and grassland [42]

Diet: Termites and certain ant species [43]
 VU 


Unknown Decrease2.svg [42]

Genus Tolypeutes Illiger, 1811 – two species
Common nameScientific name and subspeciesRangeSize and ecologyIUCN status and estimated population
Brazilian three-banded armadillo

EdmontonZooArmadillo.JPG

T. tricinctus
(Linnaeus, 1758)
Eastern South America
Tolypeutes tricinctus distribution.svg
Size: 23–25 cm (9–10 in) long, plus tail [44]

Habitat: Savanna and shrubland [45]

Diet: Ants, termites, beetles, and other insects, as well as plants [44]
 VU 


Unknown Decrease2.svg [45]

Southern three-banded armadillo

SouthernThreeBandedArmadillo065b.jpg

T. matacus
(Desmarest, 1804)
South-central South America
Southern Three-banded Armadillo area.png
Size: 20–25 cm (8–10 in) long, plus tail [46]

Habitat: Savanna and shrubland [47]

Diet: Insects, as well as fruit and seeds [46]
 NT 


Unknown Decrease2.svg [47]

Related Research Articles

<span class="mw-page-title-main">Armadillo</span> New World placental mammals in the order Cingulata

Armadillos are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along with the anteaters and sloths. Nine extinct genera and 21 extant species of armadillo have been described, some of which are distinguished by the number of bands on their armor. All species are native to the Americas, where they inhabit a variety of different environments.

<span class="mw-page-title-main">Xenarthra</span> Superorder of mammals including anteaters, sloths, and armadillos

Xenarthra is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 Mya, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene.

<span class="mw-page-title-main">Pink fairy armadillo</span> Species of mammals belonging to the armadillo order of xenarthrans

The pink fairy armadillo or pichiciego is the smallest species of armadillo, first described by Richard Harlan in 1825. This solitary, desert-adapted animal is endemic to central Argentina and can be found inhabiting sandy plains, dunes, and scrubby grasslands.

<span class="mw-page-title-main">Six-banded armadillo</span> Species of mammals belonging to the armadillo order of xenarthrans

The six-banded armadillo, also known as the yellow armadillo, is an armadillo found in South America. The sole extant member of its genus, it was first described by Swedish zoologist Carl Linnaeus in 1758. The six-banded armadillo is typically between 40 and 50 centimeters in head-and-body length, and weighs 3.2 to 6.5 kilograms. The carapace is pale yellow to reddish brown, marked by scales of equal length, and scantily covered by buff to white bristle-like hairs. The forefeet have five distinct toes, each with moderately developed claws.

<span class="mw-page-title-main">Southern naked-tailed armadillo</span> Species of mammal

The southern naked-tailed armadillo is a species of small armadillo from South America.

<span class="mw-page-title-main">Seven-banded armadillo</span> Species of armadillo

The seven-banded armadillo, also known as the Brazilian lesser long-nosed armadillo, is a species of armadillo from South America found in Paraguay, Argentina, Bolivia and Brazil. It is a solitary nocturnal, terrestrial animal, living mostly in dry habitats, outside of rainforest regions.

<span class="mw-page-title-main">Southern long-nosed armadillo</span> Species of mammal

The southern long-nosed armadillo is a species of armadillo native to South America.

<span class="mw-page-title-main">Southern three-banded armadillo</span> Type of Armadillo

The southern three-banded armadillo, also known as La Plata three-banded armadillo or Azara's domed armadillo, is an armadillo species from South America. It is found in parts of northern Argentina, southwestern Brazil, Paraguay and Bolivia, at elevations from sea level to 770 m (2,530 ft).

<span class="mw-page-title-main">Andean hairy armadillo</span> Species of mammal

The Andean hairy armadillo is an armadillo found mainly in Bolivia, in the region of the Puna; the departments of Oruro, La Paz, and Cochabamba. Nowark (1991) describes it as distributed in Bolivia and northern Chile. A recent publication of Pacheco (1995) also locates the species in the Puno Region of Peru. This species is also thought to be present in northern Argentina. However, this location may actually only contain a population of C. vellerosus.

<span class="mw-page-title-main">Northern naked-tailed armadillo</span> Species of mammal

The northern naked-tailed armadillo is a species of armadillo. It is one of only two species of armadillo found outside of South America, the other being the more widely distributed nine-banded armadillo.

<span class="mw-page-title-main">Chacoan naked-tailed armadillo</span> Species of mammal

The Chacoan naked-tailed armadillo is a species of South American armadillo.

<span class="mw-page-title-main">Big hairy armadillo</span> Species of mammal

The bighairy armadillo is one of the largest and most numerous armadillos in South America. It lives from sea level to altitudes of up to 1,300 meters across the southern portion of South America, and can be found in grasslands, forests, and savannahs, and has even started claiming agricultural areas as its home. It is an accomplished digger and spends most of its time below ground. It makes both temporary and long-term burrows, depending on its food source.

<span class="mw-page-title-main">Hairy long-nosed armadillo</span> Species of mammal

The hairy long-nosed armadillo or woolly armadillo is a species of armadillo in the family Dasypodidae. It is endemic to Peru. Its natural habitats are subtropical or tropical moist lowland forest and subtropical or tropical moist montane forest. The International Union for Conservation of Nature used to consider it a "vulnerable species" but has changed this assessment to "data deficient" because so little is known about the animal and the threats it faces.

<span class="mw-page-title-main">Yepes's mulita</span> Species of mammal

Yepes's mulita or the Yungas lesser long-nosed armadillo is a species of armadillo in the family Dasypodidae. It is endemic to Argentina and Bolivia. Its natural habitat is subtropical dry forests. The species was renamed D. yepesi because the type of D. mazzai was suspected to correspond of other species of Dasypus, which it was later proved wrong, becoming D. yepesi a synonym of D. mazzai.

<span class="mw-page-title-main">Tolypeutinae</span> Subfamily of mammals

Tolypeutinae is a subfamily of armadillos in the family Chlamyphoridae, consisting of the giant, three-banded and naked-tailed armadillos.

<span class="mw-page-title-main">Llanos long-nosed armadillo</span> Species of mammal

The Llanos long-nosed armadillo or northern long-nosed armadillo is a species of armadillo in the family Dasypodidae. It is endemic to Colombia and Venezuela, where its habitat is the intermittently flooded grassland of the Llanos. The species is closely related to the nine-banded armadillo and the great long-nosed armadillo. It has very little hair and can weigh up to 22 pounds (9.5 kg), and can grow to about 2.1 feet (60 cm) long. It lives in dense cover near limestone formations. Like most other armadillos, it eats ants.

<span class="mw-page-title-main">Chlamyphoridae</span> Family of armadillos

Chlamyphoridae is a family of cingulate mammals. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, there had been speculation that the extant family Dasypodidae could be paraphyletic based on morphological evidence. In 2016, an analysis of Doedicurus mtDNA found it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae. For this reason, all extant armadillos but Dasypus were relocated to a new family.

<span class="mw-page-title-main">Chlamyphorinae</span> Subfamily of mammals

Chlamyphorinae is a subfamily of South American armadillos in the family Chlamyphoridae. Members of this subfamily, the fairy armadillos, are largely fossorial and have reduced eyes and robust forearms with large claws for digging.

References

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Sources