Cingulata is an order of armored placental mammals. Members of this order are called cingulates, or colloquially, armadillos. They are primarily found in South America, though the northern naked-tailed armadillo is found mainly in Central America and the nine-banded armadillo has a range extending into North America. They are generally found in forests, but also savannas, shrublands, and grasslands. They all follow a similar body plan, and range in size from the pink fairy armadillo, at 11 cm (4 in) plus a 2 cm (1 in) tail, to the giant armadillo, at 100 cm (39 in) plus a 50 cm (20 in) tail. No population estimates have been made for any cingulate species, though the giant armadillo and the Brazilian three-banded armadillo are categorized as vulnerable species.
The twenty-two extant species of Cingulata are divided into two families: Dasypodidae, containing a single genus of nine species in the subfamily Dasypodinae, and Chlamyphoridae, containing thirteen species split between the two genera in the subfamily Chlamyphorinae, three in the subfamily Euphractinae, and three in the subfamily Tolypeutinae. Prior to 2016, all four subfamilies were included in Dasypodidae, with Chlamyphoridae containing only extinct species of glyptodonts. [1] [2] Over one hundred extinct Cingulata species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed. [3]
Conservation status | |
---|---|
EX | Extinct (0 species) |
EW | Extinct in the wild (0 species) |
CR | Critically Endangered (0 species) |
EN | Endangered (0 species) |
VU | Vulnerable (2 species) |
NT | Near threatened (5 species) |
LC | Least concern (8 species) |
Other categories | |
DD | Data deficient (5 species) |
NE | Not evaluated (2 species) |
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the cingulate's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".
The order Cingulata consists of two families, Dasypodidae and Chlamyphoridae. Dasypodidae contains nine species in a single genus, while Chlamyphoridae contains thirteen species in eight genera, divided into three subfamilies. Many of these species are further subdivided into subspecies. This does not include hybrid species or extinct prehistoric species.
Family Dasypodidae
Family Chlamyphoridae
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis. [4]
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
East Amazonian long-nosed armadillo | D. beniensis Lönnberg, 1942 | North-central South America | Size: 51–58 cm (20–23 in) long, plus 33–48 cm (13–19 in) tail [5] Habitat: Forest [6] Diet: Insects [5] | NE
|
Greater long-nosed armadillo | D. kappleri Krauss, 1862 | Northeastern South America | Size: 51–58 cm (20–23 in) long, plus 33–48 cm (13–19 in) tail [7] Habitat: Forest [6] Diet: Insects [7] | LC
|
Hairy long-nosed armadillo | D. pilosus (Fitzinger, 1856) | Western South America | Size: 32–44 cm (13–17 in) long, plus 23–31 cm (9–12 in) tail [8] Habitat: Forest [9] Diet: Insects [8] | NE
|
Llanos long-nosed armadillo | D. sabanicola Mondolfi, 1968 | Northern South America | Size: 25–31 cm (10–12 in) long, plus 17–21 cm (7–8 in) tail [10] Habitat: Forest [11] Diet: Termites, as well as ants, beetles, and worms [10] | NT
|
Nine-banded armadillo | D. novemcinctus Linnaeus, 1758 Six subspecies
| Central and northern South America, and central, southern, and eastern North America | Size: 35–57 cm (14–22 in) long, plus 24–45 cm (9–18 in) tail [12] Habitat: Forest, savanna, shrubland, and grassland [13] Diet: Omnivorous, including invertebrates, birds, fruit, and roots [12] | LC
|
Seven-banded armadillo | D. septemcinctus Linnaeus, 1758 | Eastern South America | Size: 24–31 cm (9–12 in) long, plus 12–17 cm (5–7 in) tail [14] Habitat: Forest, savanna, and grassland [15] Diet: Insects, seeds, and other plant material [14] | LC
|
Southern long-nosed armadillo | D. hybridus Desmarest, 1804 | Southeastern South America | Size: 26–31 cm (10–12 in) long, plus 15–19 cm (6–7 in) tail [16] Habitat: Forest, savanna, shrubland, and grassland [17] Diet: Ants, termites, and beetles, as well as plant material and small vertebrates [16] | NT
|
West Amazonian long-nosed armadillo | D. pastasae (Thomas, 1901) | Northwestern South America | Size: 51–58 cm (20–23 in) long, plus 33–48 cm (13–19 in) tail [18] Habitat: Forest [6] Diet: Insects [18] | DD
|
Yepes's mulita | D. mazzai Yepes, 1933 | South-central South America | Size: About 31 cm (12 in) long, plus 18–23 cm (7–9 in) tail [19] Habitat: Forest [20] Diet: Believed to be omnivorous with a preference for insects [19] | DD
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Greater fairy armadillo | C. retusus (Burmeister, 1863) | Central South America | Size: 14–18 cm (6–7 in) long, plus 4 cm (2 in) tail [21] Habitat: Savanna, shrubland, and desert [22] Diet: Insects, worms, snails, roots, and small seeds [21] | DD
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Pink fairy armadillo | C. truncatus Harlan, 1825 | Southern South America | Size: 11–15 cm (4–6 in) long, plus 2–3 cm (1–1 in)tail [23] [24] Habitat: Savanna, shrubland, grassland, and desert [25] Diet: insects, worms and snails, as well as possibly plants [23] | DD
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Big hairy armadillo | C. villosus (Desmarest, 1804) | Southern South America | Size: 22–40 cm (9–16 in) long, plus 9–17 cm (4–7 in) tail [12] Habitat: Forest, savanna, shrubland, grassland, and desert [26] Diet: Omnivorous, including insects, invertebrates, small vertebrates, plants, and carrion [27] | LC
|
Screaming hairy armadillo | C. vellerosus (Gray, 1865) Two subspecies
| Southern South America | Size: 20–30 cm (8–12 in) long, plus tail [28] Habitat: Savanna, shrubland, grassland, and desert [29] Diet: Beetles, butterfly larvae, plants, and small vertebrates [29] | LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Six-banded armadillo | E. sexcinctus (Linnaeus, 1758) Five subspecies
| Central and eastern South America | Size: 40–50 cm (16–20 in) long, plus 20–25 cm (8–10 in) tail [30] Habitat: Forest, savanna, shrubland, and grassland [31] Diet: Carrion, small vertebrates, insects, spiders, bird eggs, and plants [30] | LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Pichi | Z. pichiy (Desmarest, 1804) Two subspecies
| Southern South America | Size: 26–34 cm (10–13 in) long, plus 10–12 cm (4–5 in) tail [12] Habitat: Shrubland, grassland, and desert [32] Diet: Insects, worms, and other invertebrates, as well as carrion [12] | NT
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Chacoan naked-tailed armadillo | C. chacoensis Wetzel, 1980 | South-central South America | Size: 30–35 cm (12–14 in) long, plus 9–10 cm (4–4 in) tail [33] Habitat: Forest, savanna, and shrubland [34] Diet: Ants and termites, as well as seeds and fruit [33] | NT
|
Greater naked-tailed armadillo | C. tatouay (Desmarest, 1804) | Eastern South America | Size: 36–49 cm (14–19 in) long, plus 15–20 cm (6–8 in) tail [35] Habitat: Forest and grassland [36] Diet: Ants and termites [37] | LC
|
Northern naked-tailed armadillo | C. centralis (Miller, 1899) | Northern South America and Central America | Size: 30–40 cm (12–16 in) long, plus 5–7 cm (2–3 in) tail [12] Habitat: Forest, savanna, and shrubland [38] Diet: Termites and ants [12] | DD
|
Southern naked-tailed armadillo | C. unicinctus (Linnaeus, 1758) Two subspecies
| Northern and central South America | Size: 35–44 cm (14–17 in) long, plus 16–20 cm (6–8 in) tail [39] Habitat: Forest, shrubland, and grassland [40] Diet: Ants and termites [39] | LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Giant armadillo | P. maximus (Kerr, 1792) | Northern and central South America | Size: 75–100 cm (30–39 in) long, plus 50 cm (20 in) tail [41] Habitat: Forest, savanna, and grassland [42] Diet: Termites and certain ant species [43] | VU
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
---|---|---|---|---|
Brazilian three-banded armadillo | T. tricinctus (Linnaeus, 1758) | Eastern South America | Size: 23–25 cm (9–10 in) long, plus tail [44] Habitat: Savanna and shrubland [45] Diet: Ants, termites, beetles, and other insects, as well as plants [44] | VU
|
Southern three-banded armadillo | T. matacus (Desmarest, 1804) | South-central South America | Size: 20–25 cm (8–10 in) long, plus tail [46] Habitat: Savanna and shrubland [47] Diet: Insects, as well as fruit and seeds [46] | NT
|
Armadillos are New World placental mammals in the order Cingulata. They form part of the superorder Xenarthra, along with the anteaters and sloths. 21 extant species of armadillo have been described, some of which are distinguished by the number of bands on their armor. All species are native to the Americas, where they inhabit a variety of different environments.
Xenarthra is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Extinct xenarthrans include the glyptodonts, pampatheres and ground sloths. Xenarthrans originated in South America during the late Paleocene about 60 million years ago. They evolved and diversified extensively in South America during the continent's long period of isolation in the early to mid Cenozoic Era. They spread to the Antilles by the early Miocene and, starting about 3 million years ago, spread to Central and North America as part of the Great American Interchange. Nearly all of the formerly abundant megafaunal xenarthrans became extinct at the end of the Pleistocene.
The six-banded armadillo, also known as the yellow armadillo, is an armadillo found in South America. The sole extant member of its genus, it was first described by Swedish zoologist Carl Linnaeus in 1758. The six-banded armadillo is typically between 40 and 50 centimeters in head-and-body length, and weighs 3.2 to 6.5 kilograms. The carapace is pale yellow to reddish brown, marked by scales of equal length, and scantily covered by buff to white bristle-like hairs. The forefeet have five distinct toes, each with moderately developed claws.
The seven-banded armadillo, also known as the Brazilian lesser long-nosed armadillo, is a species of armadillo from South America found in Paraguay, Argentina, Bolivia and Brazil. It is a solitary nocturnal, terrestrial animal, living mostly in dry habitats, outside of rainforest regions.
The southern long-nosed armadillo is a species of armadillo native to South America.
The southern three-banded armadillo, also known as La Plata three-banded armadillo or Azara's domed armadillo, is an armadillo species from South America. It is found in parts of northern Argentina, southwestern Brazil, Paraguay and Bolivia, at elevations from sea level to 770 m (2,530 ft).
The Andean hairy armadillo is an armadillo found mainly in Bolivia, in the region of the Puna; the departments of Oruro, La Paz, and Cochabamba. Nowark (1991) describes it as distributed in Bolivia and northern Chile. A recent publication of Pacheco (1995) also locates the species in the Puno Region of Peru. This species is also thought to be present in northern Argentina. However, this location may actually only contain a population of C. vellerosus.
The northern naked-tailed armadillo is a species of armadillo. It is one of only two species of armadillo found outside of South America, the other being the more widely distributed nine-banded armadillo.
The Chacoan naked-tailed armadillo is a species of South American armadillo.
Chaetophractus is a small genus of armadillos in the family Chlamyphoridae. It contains the following three species:
The bighairy armadillo is one of the largest and most numerous armadillos in South America. It lives from sea level to altitudes of up to 1,300 meters across the southern portion of South America, and can be found in grasslands, forests, and savannahs, and has even started claiming agricultural areas as its home. It is an accomplished digger and spends most of its time below ground. It makes both temporary and long-term burrows, depending on its food source. In Spanish it is colloquially known as "peludo".
The hairy long-nosed armadillo or woolly armadillo is a species of armadillo in the family Dasypodidae. It is endemic to Peru. Its natural habitats are subtropical or tropical moist lowland forest and subtropical or tropical moist montane forest. The International Union for Conservation of Nature used to consider it a "vulnerable species" but has changed this assessment to "data deficient" because so little is known about the animal and the threats it faces.
Yepes's mulita or the Yungas lesser long-nosed armadillo is a species of armadillo in the family Dasypodidae. It is endemic to Argentina and Bolivia. Its natural habitat is subtropical dry forests. The species was renamed D. yepesi because the type of D. mazzai was suspected to correspond of other species of Dasypus, which it was later proved wrong, becoming D. yepesi a synonym of D. mazzai.
Tolypeutinae is a subfamily of armadillos in the family Chlamyphoridae, consisting of the giant, three-banded and naked-tailed armadillos.
The Llanos long-nosed armadillo or northern long-nosed armadillo is a species of armadillo in the family Dasypodidae. It is endemic to Colombia and Venezuela, where its habitat is the intermittently flooded grassland of the Llanos. The species is closely related to the nine-banded armadillo and the great long-nosed armadillo. It has very little hair and can weigh up to 22 pounds (9.5 kg), and can grow to about 2.1 feet (60 cm) long. It lives in dense cover near limestone formations. Like most other armadillos, it eats ants.
Chlamyphoridae is a family of cingulate mammals. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, there had been speculation that the extant family Dasypodidae could be paraphyletic based on morphological evidence. In 2016, an analysis of Doedicurus mtDNA found it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae. For this reason, all extant armadillos but Dasypus were relocated to a new family.
Chlamyphorinae is a subfamily of South American armadillos in the family Chlamyphoridae. Members of this subfamily, the fairy armadillos, are largely fossorial and have reduced eyes and robust forearms with large claws for digging.