List of myotines

Greater mouse-eared bat (Myotis myotis)

Myotinae is one of the four subfamilies of Vespertilionidae, itself one of twenty families of bats in the mammalian order Chiroptera and part of the microbat suborder. A member of this subfamily is called a myotine, or a vesper bat. They are found in all continents besides Antarctica, primarily in forests and caves, though some species can also be found in shrublands, grasslands, rocky areas, or deserts. They range in size from the Taiwan broad-muzzled bat, at 3 cm (1 in) plus a 3 cm (1 in) tail, to the large myotis, at 10 cm (4 in) plus a 6 cm (2 in) tail. Like all bats, myotines are capable of true and sustained flight, and have wing lengths ranging from 2 cm (1 in) to 7 cm (3 in). They are all insectivorous and eat a variety of insects and spiders, ^[1] but some bats with long toes which trawl for insects on top of the water, such as the long-fingered bat Daubenton's bat, Maluku myotis, Rickett's big-footed bat, and pond bat, may sometimes supplement their diet with small fish from still waters. The fish-eating bat also regularly eats fish and crustaceans as well as insects, and is the only bat species that hunts fish in the ocean. ^[2] Almost no myotines have population estimates, though seven species—the Atacama myotis, eastern small-footed myotis, Findley's myotis, flat-headed myotis, frosted myotis, little brown bat, and peninsular myotis—are categorized as endangered species, and two species—the Nimba myotis and Yanbaru whiskered bat—are categorized as critically endangered.

The 121 extant species of Myotinae are divided between three genera: Eudiscopus and Submyotodon with one species each, and Myotis, or the mouse-eared bats, with the other 119. A few extinct prehistoric myotine species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed. ^[3]

Conventions

IUCN Red List categories

Conservation status
EX	Extinct (0 species)
EW	Extinct in the wild (0 species)
CR	Critically Endangered (2 species)
EN	Endangered (7 species)
VU	Vulnerable (8 species)
NT	Near threatened (11 species)
LC	Least concern (65 species)
Other categories
DD	Data deficient (28 species)
NE	Not evaluated (0 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the myotine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. Population figures are rounded to the nearest hundred.

Classification

Myotinae, one of the four subfamilies of the family Vespertilionidae, contains 121 extant species divided into 3 genera.

Subfamily Myotinae

• Genus Eudiscopus (disk-footed bat): one species
• Genus Myotis (mouse-eared bats): 119 species
• Genus Submyotodon (Taiwan broad-muzzled bat): one species

Myotines

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists. ^[4]

Genus Eudiscopus – Conisbee, 1953 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Disk-footed bat	E. denticulus (Osgood, 1932)	Southeastern Asia	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest [6]	LC  Unknown [6]

Genus Myotis – Kaup, 1829 – 119 species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Alcathoe bat	M. alcathoe von Helversen & Heller, 2001	Europe and Turkey	Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Forest [8]	DD  Unknown [8]
Anjouan myotis	M. anjouanensis Dorst, 1960	Madagascar	Size: About 5 cm (2 in), plus about 5 cm (2 in) tail about 4 cm (2 in) arm/wing length [7] Habitat: Unknown [9]	DD  Unknown [9]
Anna Tess's bat	M. annatessae Kruskop & Borisenko, 2013	Vietnam and Laos	Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest [11]	DD  Unknown [11]
Annamit myotis	M. annamiticus Kruskop & Tsytsulina, 2001	Vietnam	Size: About 4 cm (2 in), plus about 3 cm (1 in) tail about 4 cm (2 in) arm/wing length [10] Habitat: Forest and inland wetlands [12]	DD  Unknown [12]
Arizona myotis	M. occultus Hollister, 1909	Mexico and southwestern United States	Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest, rocky areas, caves, and desert [13]	LC  Unknown [13]
Atacama myotis	M. atacamensis Lataste, 1892	Chile and Peru	Size: 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Shrubland, rocky areas, and desert [15]	EN  Unknown [15]
Australian myotis	M. australis Dobson, 1878	Australia	Size: Unknown [16] Habitat: Unknown [16]	DD  Unknown [16]
Barbados myotis	M. nyctor LaVal & Schwartz, 1974	Lesser Antilles in the Caribbean	Size: 4–5 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest and caves [17]	VU  Unknown [17]
Bechstein's bat	M. bechsteinii (Kuhl, 1817)	Europe and western Asia	Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, shrubland, and caves [19]	NT  Unknown [19]
Beijing mouse-eared bat	M. pequinius Thomas, 1908	Eastern China	Size: About 6 cm (2 in), plus about 4 cm (2 in) tail 4–6 cm (2–2 in) arm/wing length [18] Habitat: Forest and caves [20]	LC  Unknown [20]
Black myotis	M. nigricans Schinz, 1821 Four subspecies M. n. carteri M. n. extremus M. n. nigricans M. n. osculatii	Mexico, Central America, and South America	Size: 3–6 cm (1–2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Forest, savanna, shrubland, and caves [21]	LC  Unknown [21]
Bocharic myotis	M. bucharensis Kuzyakin, 1950	Central Asia	Size: 4–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Caves [22]	DD  Unknown [22]
Bornean whiskered myotis	M. borneoensis Hill & Francis, 1984	Island of Borneo (in green)	Size: Unknown length, plus 4–5 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length [10] Habitat: Forest [23]	DD  Unknown [23]
Brandt's bat	M. brandtii (Eversmann, 1845) Two subspecies M. b. brandtii M. b. gracilis	Europe and western Asia (in red)	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Forest, shrubland, grassland, inland wetlands, and caves [24]	LC  Unknown [24]
Burmese whiskered myotis	M. montivagus (Dobson, 1874)	Southern Asia (in leftmost red)	Size: Unknown length 3–5 cm (1–2 in) arm/wing length [10] Habitat: Forest and caves [25]	DD  Unknown [25]
California myotis	M. californicus (Audubon & Bachman, 1842) Four subspecies M. c. californicus M. c. caurinus M. c. mexicanus M. c. stephensi	Western North America	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest, caves, and desert [26]	LC  Unknown [26]
Cape hairy bat	M. tricolor (Temminck, 1832)	Sub-Saharan Africa	Size: 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail 4–6 cm (2–2 in) arm/wing length [7] Habitat: Forest, savanna, and shrubland [27]	LC  Unknown [27]
Cave myotis	M. velifer (Allen, 1890) Five subspecies M. v. brevis M. v. grandis M. v. incautus M. v. magnamolaris M. v. velifer	United States and Mexico	Size: 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [14] Habitat: Forest and caves [28]	LC  Unknown [28]
Chilean myotis	M. chiloensis (Waterhouse, 1840)	Southern South America	Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [14] Habitat: Rocky areas and forest [29]	LC  Unknown [29]
Chinese water myotis	M. laniger Peters, 1870	Eastern Asia	Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest and caves [30]	LC  Unknown [30]
Cinnamon myotis	M. fortidens Miller & Allen, 1928 Two subspecies M. f. fortidens M. f. sonoriensis	Guatemala and Mexico	Size: 4–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [31]	LC  Unknown [31]
Cryptic myotis	M. crypticus Ruedi, Ibáñez, Salicini, Juste, & Puechmaille, 2019	Southern Europe	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, inland wetlands, and caves [32]	NT  Unknown [32]
Csorba's mouse-eared bat	M. csorbai Topál, 1997	Nepal	Size: Unknown [33] Habitat: Forest [33]	DD  Unknown [33]
Curacao myotis	M. nesopolus Miller, 1900 Two subspecies M. n. larensis M. n. nesopolus	Northern South America	Size: About 3 cm (1 in), plus about 4 cm (2 in) tail 2–4 cm (1–2 in) arm/wing length [14] Habitat: Forest and shrubland [34]	LC  Unknown [34]
Dark-nosed small-footed myotis	M. melanorhinus Merriam, 1890	Western North America	Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Caves and forest [35]	LC  Unknown [35]
Daubenton's bat	M. daubentonii (Kuhl, 1817) Six subspecies M. d. chasanensis M. d. daubentonii M. d. loukashkini M. d. nathalinae M. d. ussuriensis M. d. volgensis	Europe and Asia	Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, shrubland, inland wetlands, rocky areas, caves, and neritic marine [36]	LC  Unknown [36]
David's myotis	M. davidii Peters, 1869	Eastern Asia	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Caves and forest [37]	LC  Unknown [37]
Diminutive bat	M. diminutus Moratelli & Wilson, 2011	Ecuador and Colombia	Size: About 4 cm (2 in), plus about 3 cm (1 in) tail about 3 cm (1 in) arm/wing length [14] Habitat: Forest [38]	DD  Unknown [38]
Dinelli's myotis	M. dinellii Thomas, 1902	Southern South America (in green)	Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest [39]	LC  Unknown [39]
Dominican myotis	M. dominicensis Miller, 1902	Dominica and Guadeloupe in the Caribbean	Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Inland wetlands and caves [40]	VU  Unknown [40]
Eastern long-fingered bat	M. macrodactylus (Temminck, 1840) Three subspecies M. m. continentalis M. m. insularis M. m. macrodactylus	Eastern Asia	Size: 4–8 cm (2–3 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, inland wetlands, and caves [41]	LC  Unknown [41]
Eastern small-footed myotis	M. leibii Audubon & Bachman, 1842	Eastern United States and Canada	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Caves, rocky areas, and forest [42]	EN  Unknown [42]
Eastern water bat	M. petax Hollister, 1912	Eastern Asia	Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [18] Habitat: Caves, inland wetlands, and forest [43]	LC  Unknown [43]
Elegant myotis	M. elegans Hall, 1962	Mexico and Central America	Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest [44]	LC  Unknown [44]
Escalera's bat	M. escalerai A. Cabrera, 1904	Southwestern Europe	Size: 4–5 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, shrubland, and caves [45]	NT  Unknown [45]
Far Eastern myotis	M. bombinus Thomas, 1906 Two subspecies M. b. amurensis M. b. bombinus	Eastern Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest and caves [46]	NT  Unknown [46]
Felten's myotis	M. punicus Felten, 1977	Northern Africa and Mediterranean islands	Size: 6–8 cm (2–3 in), plus tail 5–7 cm (2–3 in) arm/wing length [18] Habitat: Caves, grassland, shrubland, and forest [47]	DD  Unknown [47]
Fish-eating bat	M. vivesi Ménégaux, 1901	Western Mexico	Size: 8–9 cm (3–4 in), plus 5–8 cm (2–3 in) tail 5–7 cm (2–3 in) arm/wing length [14] Habitat: Rocky areas and caves [48]	VU  Unknown [48]
Findley's myotis	M. findleyi Bogan, 1978	Tres Marías Islands west of Mexico	Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 2–4 cm (1–2 in) arm/wing length [5] Habitat: Unknown [49]	EN  Unknown [49]
Flat-headed myotis	M. planiceps Baker, 1955	Central Mexico	Size: 3–6 cm (1–2 in), plus 2–3 cm (1–1 in) tail 2–3 cm (1–1 in) arm/wing length [5] Habitat: Forest and caves [50]	EN  240 [50]
Fraternal myotis	M. frater (Allen, 1923)	Eastern Asia (in red)	Size: 3–5 cm (1–2 in), plus 4–6 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest and caves [51]	LC  Unknown [51]
Fringed long-footed myotis	M. fimbriatus Peters, 1870	Eastern Asia	Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Caves [52]	LC  Unknown [52]
Fringed myotis	M. thysanodes Miller, 1897 Four subspecies M. t. aztecus M. t. pahasapensis M. t. thysanodes M. t. vespertinus	Western North America	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length [5] Habitat: Forest, rocky areas, caves, and desert [53]	LC  Unknown [53]
Frosted myotis	M. pruinosus Yoshiyuki, 1971	Japan	Size: 3–5 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest and caves [54]	EN  Unknown [54]
Geoffroy's bat	M. emarginatus (Geoffroy, 1806) Three subspecies M. e. desertorum M. e. emarginatus M. e. turcomanicus	Europe, northern Africa, and western Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [7] Habitat: Caves, grassland, and shrubland [55]	LC  Unknown [55]
Gomantong myotis	M. gomantongensis Francis & Hill, 1998	Island of Borneo in Malaysia	Size: Unknown length, plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length [10] Habitat: Caves [56]	LC  Unknown [56]
Gray bat	M. grisescens Howell, 1909	Eastern United States	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length [14] Habitat: Forest and caves [57]	VU  Unknown [57]
Greater mouse-eared bat	M. myotis (Borkhausen, 1797) Two subspecies M. m. macrocephalicus M. m. myotis	Europe and western Asia	Size: 6–9 cm (2–4 in), plus 4–6 cm (2–2 in) tail 5–7 cm (2–3 in) arm/wing length [18] Habitat: Forest, shrubland, and caves [58]	LC  Unknown [58]
Guatemalan myotis	M. cobanensis Goodwin, 1955	Guatemala	Size: About 4 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length [14] Habitat: Unknown [59]	DD  Unknown [59]
Hairy-faced bat	M. annectans Dobson, 1871	Southern and southeastern Asia	Size: 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length [10] Habitat: Forest [60]	LC  Unknown [60]
Hairy-legged myotis	M. keaysi Allen, 1914 Two subspecies M. k. keaysi M. k. pilosotibialis	Mexico, Central America, and northern and western South America	Size: 5–7 cm (2–3 in), plus 2–5 cm (1–2 in) tail Unknown arm/wing length [5] Habitat: Forest [61]	LC  Unknown [61]
Herman's myotis	M. hermani Thomas, 1923	Indonesia	Size: 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 4–6 cm (2–2 in) arm/wing length [7] Habitat: Forest [62]	DD  Unknown [62]
Himalayan whiskered bat	M. siligorensis Horsfield, 1855 Four subspecies M. s. alticraniatus M. s. siligorensis M. s. sowerbyi M. s. thaianus	Central, eastern, and southeastern Asia	Size: Unknown length about 3 cm (1 in) arm/wing length [10] Habitat: Forest and caves [63]	LC  Unknown [63]
Hodgson's bat	M. formosus (Hodgson, 1835)	Central and eastern, and southeastern Asia	Size: 5–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 4–6 cm (2–2 in) arm/wing length [7] Habitat: Forest, shrubland, grassland, and caves [64]	NT  Unknown [64]
Horsfield's bat	M. horsfieldii Temminck, 1840 Five subspecies M. h. deignani M. h. dryas M. h. horsfieldii M. h. jeannei M. h. peshwa	Southern and southeastern Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [10] Habitat: Forest and caves [65]	LC  Unknown [65]
Ikonnikov's bat	M. ikonnikovi Ogniov, 1912	Eastern Asia	Size: 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Forest, inland wetlands, and caves [66]	LC  Unknown [66]
Indiana bat	M. sodalis Miller, 1898	Eastern United States	Size: 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [67]	NT  Unknown [67]
Indochinese mouse-eared bat	M. indochinensis Son, Motokawa, Estók, Thong, Dang, Oshida, Csorba, Francis, Görföl, & Endō, 2013	Vietnam and China	Size: Unknown length 4–5 cm (2–2 in) arm/wing length [10] Habitat: Forest [68]	DD  Unknown [68]
Insular myotis	M. insularum Dobson, 1878	American Samoa	Size: About 4 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length [10] Habitat: Unknown [69]	DD  Unknown [69]
Izecksohn's myotis	M. izecksohni Moratelli, Peracchi, Dias, & de Oliveira, 2011	Brazil and Argentina	Size: 4–7 cm (2–3 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest [70]	DD  Unknown [70]
Kashmir cave bat	M. longipes Dobson, 1873	Southern Asia	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest and caves [71]	DD  Unknown [71]
Keen's myotis	M. keenii (Merriam, 1895)	Western Canada and United States	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest and rocky areas [72]	LC  Unknown [72]
Kei myotis	M. stalkeri Thomas, 1910	Indonesia	Size: 4–8 cm (2–3 in), plus 5–6 cm (2–2 in) tail 5–6 cm (2–2 in) arm/wing length [10] Habitat: Forest, inland wetlands, and caves [73]	LC  10,000–11,000 [73]
Kock's mouse-eared bat	M. dieteri Happold, 2005	Republic of the Congo	Size: About 5 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length [7] Habitat: Forest and caves [74]	DD  Unknown [74]
Large myotis	M. chinensis Tomes, 1857	Eastern and southeastern Asia	Size: 9–10 cm (4–4 in), plus 5–6 cm (2–2 in) tail 6–7 cm (2–3 in) arm/wing length [18] Habitat: Forest, inland wetlands, and caves [75]	LC  Unknown [75]
Large-footed bat	M. adversus Horsfield, 1824 Six subspecies M. a. adversus M. a. carimatae M. a. orientis M. a. taiwanensis M. a. tanimbarensis M. a. wetarensis	Southeastern Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [10] Habitat: Forest, inland wetlands, and caves [76]	LC  Unknown [76]
Large-footed myotis	M. macropus (Gould, 1854)	Eastern Australia	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Inland wetlands and caves [77]	LC  Unknown [77]
LaVal's myotis	M. lavali Moratelli, Peracchi, Dias, & de Oliveira, 2011	Central and eastern South America	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest, savanna, and grassland [78]	LC  Unknown [78]
Lesser large-footed bat	M. hasseltii Temminck, 1840 Four subspecies M. h. abboti M. h. continentis M. h. hasseltii M. h. macellus	Southern and southeastern Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest and caves [79]	LC  Unknown [79]
Lesser mouse-eared bat	M. blythii Tomes, 1857 Four subspecies M. b. ancilla M. b. blythii M. b. lesviacus M. b. omari	Europe and Asia	Size: 5–8 cm (2–3 in), plus about 6 cm (2 in) tail 5–7 cm (2–3 in) arm/wing length [18] Habitat: Shrubland, grassland, and caves [80]	LC  Unknown [80]
Little brown bat	M. lucifugus (Conte, 1831) Five subspecies M. l. alascensis M. l. carissima M. l. lucifugus M. l. pernox M. l. relictus	United States and Canada	Size: 3–6 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [81]	EN  Unknown [81]
Long-eared myotis	M. evotis (H. Allen, 1864) Six subspecies M. e. chrysonotus M. e. evotis M. e. jonesorum M. e. micronyx M. e. milleri M. e. pacificus	Western North America	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Caves, rocky areas, and forest [82]	LC  Unknown [82]
Long-fingered bat	M. capaccinii Bonaparte, 1837	Southern Europe, northern Africa, and western Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [10] Habitat: Shrubland, inland wetlands, and caves [83]	VU  Unknown [83]
Long-legged myotis	M. volans H. Allen, 1866 Four subspecies M. v. amotus M. v. interior M. v. longicrus M. v. volans	Western North America	Size: 4–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest, rocky areas, caves, and desert [84]	LC  Unknown [84]
Long-tailed myotis	M. longicaudatus Ogniov, 1927 Four subspecies M. l. eniseensis M. l. kaguyae M. l. longicaudatus	Eastern Asia (excluding red)	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, inland wetlands, and caves [85]	LC  Unknown [85]
Long-toed myotis	M. secundus Ruedi, Csorba, Lin, & Chou, 2015	Taiwan	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest [86]	LC  Unknown [86]
Malagasy mouse-eared bat	M. goudoti Smith, 1834	Madagascar	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [7] Habitat: Forest, rocky areas, and caves [87]	LC  Unknown [87]
Malaysian whiskered myotis	M. federatus Thomas, 1916	Malaysia (in blue)	Size: About 5 cm (2 in), plus about 4 cm (2 in) tail 3–5 cm (1–2 in) arm/wing length [10] Habitat: Forest [88]	DD  Unknown [88]
Maluku myotis	M. moluccarum Thomas, 1915 Three subspecies M. m. moluccarum M. m. richardsi M. m. solomonis	Indonesia	Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [10] Habitat: Forest and inland wetlands [89]	LC  Unknown [89]
Mandelli's mouse-eared bat	M. sicarius Thomas, 1915	India and Nepal	Size: 5–6 cm (2–2 in), plus 4–6 cm (2–2 in) tail 4–6 cm (2–2 in) arm/wing length [18] Habitat: Forest and caves [90]	VU  Unknown [90]
Montane myotis	M. oxyotus Peters, 1867 Two subspecies M. o. gardneri M. o. oxyotus	Northern and western South America	Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [14] Habitat: Forest [91]	LC  Unknown [91]
Morris's bat	M. morrisi Hill, 1971	Ethiopia and Nigeria	Size: About 5 cm (2 in), plus about 5 cm (2 in) tail 4–5 cm (2–2 in) arm/wing length [7] Habitat: Savanna and caves [92]	DD  Unknown [92]
Natterer's bat	M. nattereri (Kuhl, 1817) Two subspecies M. n. nattereri M. n. tschuliensis	Europe and western Asia	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, shrubland, grassland, inland wetlands, and caves [93]	LC  Unknown [93]
Nepal myotis	M. nipalensis Dobson, 1871 Three subspecies M. n. nipalensis M. n. przewalskii M. n. transcaspicus	Central and western Asia	Size: About 4 cm (2 in), plus about 33–5 cm (1–2 in) tail about 3 cm (1 in) arm/wing length [7] Habitat: Forest, shrubland, grassland, caves, and desert [94]	LC  Unknown [94]
Nimba myotis	M. nimbaensis Simmons, Flanders, Fils, Parker, Suter, Bamba, Douno, Keita, Morales, & Frick, 2021	Guinea	Size: 6–8 cm (2–3 in), plus 4–5 cm (2–2 in) tail 5–6 cm (2–2 in) arm/wing length [7] Habitat: Forest, grassland, inland wetlands, caves, and shrubland [95]	CR  Unknown [95]
Northern long-eared bat	M. septentrionalis Trouessart, 1897	Canada and eastern United States	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest and caves [96]	NT  Unknown [96]
Orange-fingered myotis	M. rufopictus (Waterhouse, 1845)	Indonesia	Size: About 6 cm (2 in), plus 4–6 cm (2–2 in) tail 4–6 cm (2–2 in) arm/wing length [7] Habitat: Forest [97]	DD  Unknown [97]
Pallid large-footed myotis	M. macrotarsus Waterhouse, 1845 Two subspecies M. m. macrotarsus M. m. saba	Indonesia	Size: About 6 cm (2 in), plus 4–6 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length [10] Habitat: Caves [98]	LC  Unknown [98]
Peninsular myotis	M. peninsularis Miller, 1898	Southern Baja California peninsula	Size: 4–6 cm (2–2 in), plus 3–6 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [14] Habitat: Forest and caves [99]	EN  Unknown [99]
Peters's myotis	M. ater (Peters, 1866) Two subspecies M. a. ater M. a. nugax	Southeastern Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [10] Habitat: Forest [100]	LC  Unknown [100]
Peyton's myotis	M. peytoni Wroughton & Ryley, 1913	Southern India (in yellow)	Size: 5–7 cm (2–3 in), plus 4–5 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length [10] Habitat: Forest [101]	DD  Unknown [101]
Pond bat	M. dasycneme (Boie, 1825)	Europe and northern Asia	Size: 5–7 cm (2–3 in), plus 4–6 cm (2–2 in) tail 4–5 cm (2–2 in) arm/wing length [7] Habitat: Forest, inland wetlands, and caves [102]	NT  Unknown [102]
Red myotis	M. ruber Geoffroy, 1806	Southeastern South America	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [5] Habitat: Forest [103]	NT  Unknown [103]
Reddish myotis	M. soror Ruedi, Csorba, Lin, & Chou, 2015	Taiwan	Size: About 5 cm (2 in), plus about 4 cm (2 in) tail about 4 cm (2 in) arm/wing length [18] Habitat: Forest [104]	DD  Unknown [104]
Reddish-black myotis	M. rufoniger (Tomes, 1858)	Eastern Asia	Size: 5–8 cm (2–3 in), plus 4–6 cm (2–2 in) tail 4–6 cm (2–2 in) arm/wing length [7] Habitat: Forest and caves [105]	LC  Unknown [105]
Rickett's big-footed bat	M. pilosus Peters, 1869	Eastern Asia	Size: 5–7 cm (2–3 in), plus 3–6 cm (1–2 in) tail 5–7 cm (2–3 in) arm/wing length [18] Habitat: Forest and inland wetlands [106]	VU  Unknown [106]
Ridley's bat	M. ridleyi Thomas, 1898	Southeastern Asia	Size: 3–5 cm (1–2 in), plus 2–4 cm (1–2 in) tail 2–4 cm (1–2 in) arm/wing length [10] Habitat: Caves, inland wetlands, and forest [107]	NT  Unknown [107]
Riparian myotis	M. riparius Handley, 1960	Central America and South America	Size: 4–5 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest [108]	LC  Unknown [108]
Rufous mouse-eared bat	M. bocagii Peters, 1870 Three subspecies M. b. bocagii M. b. cupreolus M. b. dogalensis	Scattered sub-Saharan Africa and Yemen	Size: 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [7] Habitat: Forest and savanna [109]	LC  Unknown [109]
Schaub's myotis	M. schaubi Kormos, 1934 Two subspecies M. s. araxenus M. s. schaubi	Armenia and Iran	Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [18] Habitat: Forest, shrubland, and caves [110]	DD  Unknown [110]
Schwartz's myotis	M. martiniquensis LaVal, 1973	Barbados and Martinique in the Caribbean	Size: Unknown length 3–4 cm (1–2 in) arm/wing length [14] Habitat: Caves [111]	NT  Unknown [111]
Scott's mouse-eared bat	M. scotti Thomas, 1927	Ethiopia	Size: 4–6 cm (2–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [7] Habitat: Forest and shrubland [112]	VU  Unknown [112]
Siberian bat	M. sibiricus (Kastschenko, 1905)	Northeastern Asia (in green)	Size: 3–6 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Forest, savanna, rocky areas, and caves [113]	LC  Unknown [113]
Silver-tipped myotis	M. albescens Geoffroy, 1806	Central America and South America	Size: 4–5 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest [114]	LC  Unknown [114]
Singapore whiskered bat	M. oreias (Temminck, 1840)	Singapore	Size: Unknown [115] Habitat: Unknown [115]	DD  Unknown [115]
Southeastern myotis	M. austroriparius (Rhoads, 1897)	Southeastern United States	Size: 4–6 cm (2–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest and caves [116]	LC  Unknown [116]
Southern myotis	M. aelleni Baud, 1979	Argentina	Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–5 cm (1–2 in) arm/wing length [14] Habitat: Unknown [117]	DD  Unknown [117]
Southwestern myotis	M. auriculus Baker & Stains, 1955 Two subspecies M. a. apache M. a. auriculus	Southern United States and Mexico	Size: 5–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest, caves, and desert [118]	LC  Unknown [118]
Szechwan myotis	M. altarium Thomas, 1911	China and Thailand	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 4–5 cm (2–2 in) arm/wing length [7] Habitat: Caves [119]	LC  Unknown [119]
Thick-thumbed myotis	M. rosseti Oei, 1951	Southeastern Asia	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 2–4 cm (1–2 in) arm/wing length [10] Habitat: Forest [120]	LC  Unknown [120]
Velvety myotis	M. simus Thomas, 1901	Western, central, and eastern South America	Size: 5–6 cm (2–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest [121]	DD  Unknown [121]
Wall-roosting mouse-eared bat	M. muricola (Gray, 1846) Seven subspecies M. m. browni M. m. caliginosus M. m. herrei M. m. moupinensis M. m. muricola M. m. niasensis M. m. patriciae	Southeastern Asia	Size: 4–6 cm (2–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest and caves [122]	LC  Unknown [122]
Weber's myotis	M. weberi (Jentink, 1890)	Sulawesi island in Indonesia	Size: About 6 cm (2 in), plus about 4 cm (2 in) tail 4–6 cm (2–2 in) arm/wing length [7] Habitat: Forest [123]	DD  Unknown [123]
Welwitsch's bat	M. welwitschii (Gray, 1866)	Western, eastern, and southern Africa	Size: 5–7 cm (2–3 in), plus 5–7 cm (2–3 in) tail 5–6 cm (2–2 in) arm/wing length [7] Habitat: Forest, savanna, shrubland, and grassland [124]	LC  Unknown [124]
Western small-footed bat	M. ciliolabrum (Merriam, 1842)	Western North America	Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest, rocky areas, caves, and desert [125]	LC  Unknown [125]
Whiskered bat	M. mystacinus (Kuhl, 1817) Three subspecies M. m. caucasicus M. m. mystacinus M. m. occidentalis	Europe, northern Africa, and western Asia	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [7] Habitat: Desert, caves, grassland, shrubland, and forest [126]	LC  Unknown [126]
Yanbaru whiskered bat	M. yanbarensis Maeda & Matsumura, 1998	Ryukyu Islands in Japan	Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [10] Habitat: Forest [127]	CR  Unknown [127]
Yellowish myotis	M. levis Geoffroy, 1806	Southeastern South America (in red)	Size: 3–6 cm (1–2 in), plus 4–5 cm (2–2 in) tail 3–5 cm (1–2 in) arm/wing length [14] Habitat: Forest [128]	LC  Unknown [128]
Yuma myotis	M. yumanensis H. Allen, 1864 Six subspecies M. y. lambi M. y. lutosus M. y. oxalis M. y. saturatus M. y. sociabilis M. y. yumanensis	Western North America	Size: 4–5 cm (2–2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [14] Habitat: Forest and caves [129]	LC  Unknown [129]

Genus Submyotodon – Ziegler, 2003 – one species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Taiwan broad-muzzled bat	S. latirostris (Kishida, 1932)	Taiwan	Size: 3–4 cm (1–2 in), plus 3–4 cm (1–2 in) tail 3–4 cm (1–2 in) arm/wing length [5] Habitat: Forest [130]	LC  Unknown [130]

References

1. Nowak, pp. 184–186
2. Levin, E.; Barnea, A.; Yovel, Y.; Yom-Tov, Y. (2006). "Have introduced fish initiated piscivory among the long-fingered bat?". Mammalian Biology . 71 (3): 139–143. Bibcode:2006MamBi..71..139L. doi:10.1016/j.mambio.2006.01.002.
3. "Fossilworks: Myotis". Paleobiology Database . University of Wisconsin–Madison. Archived from the original on September 22, 2022. Retrieved November 2, 2024.
4. Wilson, Reeder, pp. 499–518
5. Chernasky; Motis; Burgin, pp. 556–557
6. Soisook, P.; Csorba, G.; Bumrungsri, S.; Francis, C. M.; Bates, P.; Kingston, T. (2016). "Eudiscopus denticulus". IUCN Red List of Threatened Species . 2016: e.T8168A22028419. doi: 10.2305/IUCN.UK.2016-2.RLTS.T8168A22028419.en .
7. Chernasky; Motis; Burgin, pp. 560–561
8. Hutson, A. M.; Paunović, M. (2016). "Myotis alcathoe". IUCN Red List of Threatened Species . 2016: e.T136680A518740. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136680A518740.en .
9. Jacobs, D. (2019). "Myotis anjouanensis". IUCN Red List of Threatened Species . 2019: e.T44863A22073545. doi: 10.2305/IUCN.UK.2019-3.RLTS.T44863A22073545.en .
10. Chernasky; Motis; Burgin, pp. 562–563
11. Srinivasulu, C.; Srinivasulu, B. (2019). "Myotis annatessae". IUCN Red List of Threatened Species . 2019: e.T85342605A85342608. doi: 10.2305/IUCN.UK.2019-3.RLTS.T85342605A85342608.en .
12. Kruskop, S. V. (2016). "Myotis annamiticus". IUCN Red List of Threatened Species . 2016: e.T136279A22006224. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136279A22006224.en .
13. Solari, S. (2018). "Myotis occultus". IUCN Red List of Threatened Species . 2018: e.T136650A21990499. doi: 10.2305/IUCN.UK.2018-2.RLTS.T136650A21990499.en .
14. Chernasky; Motis; Burgin, pp. 558–559
15. Vargas-Rodríguez, R.; Peñaranda, D.; Ugarte Nuñez, J.; Rodríguez-San Pedro, A.; Ossa Gomez, G.; Gatica Castro, A. (2016). "Myotis atacamensis". IUCN Red List of Threatened Species . 2016: e.T14143A22050638. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14143A22050638.en .
16. Reardon, T. B.; Lumsden, L. F. (2020). "Myotis australis". IUCN Red List of Threatened Species . 2020: e.T14146A22060248. doi: 10.2305/IUCN.UK.2020-3.RLTS.T14146A22060248.en .
17. Larsen, R. (2016). "Myotis nyctor". IUCN Red List of Threatened Species . 2016: e.T76435059A76435083. doi: 10.2305/IUCN.UK.2016-1.RLTS.T76435059A76435083.en .
18. Chernasky; Motis; Burgin, pp. 564–565
19. Paunović, M. (2016). "Myotis bechsteinii". IUCN Red List of Threatened Species . 2016: e.T14123A22053752. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14123A22053752.en .
20. Feng, J.; Jiang, T. L. (2019). "Myotis pequinius". IUCN Red List of Threatened Species . 2019: e.T14190A22066613. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14190A22066613.en .
21. Solari, S. (2019). "Myotis nigricans". IUCN Red List of Threatened Species . 2019: e.T14185A22066939. doi: 10.2305/IUCN.UK.2019-2.RLTS.T14185A22066939.en .
22. Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis bucharensis". IUCN Red List of Threatened Species . 2019: e.T136219A22011494. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136219A22011494.en .
23. Görföl, T.; Csorba, G. (2017). "Myotis borneoensis". IUCN Red List of Threatened Species . 2017: e.T85568289A85568292. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85568289A85568292.en .
24. Gazaryan, S.; Kruskop, S. V.; Godlevska, L. (2021) [errata version of 2020 assessment]. "Myotis brandtii". IUCN Red List of Threatened Species . 2020: e.T85566997A195857637. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85566997A195857637.en .
25. Görföl, T.; Tu, V.; Csorba, G.; Francis, C. M.; Hutson, A. M.; Bates, P. J. J.; Bumrungsri, S.; Molur, S.; Srinivasulu, C. (2020). "Myotis montivagus". IUCN Red List of Threatened Species . 2020: e.T85567622A22065126. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85567622A22065126.en .
26. Arroyo-Cabrales, J.; Perez, S. (2017). "Myotis californicus". IUCN Red List of Threatened Species . 2017: e.T14150A22061366. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14150A22061366.en .
27. Monadjem, A.; Jacobs, D. (2017). "Myotis tricolor". IUCN Red List of Threatened Species . 2017: e.T14207A22063832. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14207A22063832.en .
28. Solari, S. (2019). "Myotis velifer". IUCN Red List of Threatened Species . 2019: e.T14208A22063586. doi: 10.2305/IUCN.UK.2019-1.RLTS.T14208A22063586.en .
29. Barquez, R.; Diaz, M. (2016). "Myotis chiloensis". IUCN Red List of Threatened Species . 2016: e.T14151A22061103. doi: 10.2305/IUCN.UK.2016-3.RLTS.T14151A22061103.en .
30. Feng, J.; Jiang, T. L. (2019). "Myotis laniger". IUCN Red List of Threatened Species . 2019: e.T136429A21984685. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136429A21984685.en .
31. Perez, S.; de Grammont, P. C.; Cuarón, A. D. (2017). "Myotis fortidens". IUCN Red List of Threatened Species . 2017: e.T14161A22056846. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14161A22056846.en .
32. Russo, D.; Cistrone, L. (2024) [errata version of 2023 assessment]. "Myotis crypticus". IUCN Red List of Threatened Species . 2023: e.T215154989A254355251.
33. Csorba, G.; Thapa, S. (2016). "Myotis csorbai". IUCN Red List of Threatened Species . 2016: e.T29420A22070788. doi: 10.2305/IUCN.UK.2016-2.RLTS.T29420A22070788.en .
34. Solari, S. (2016). "Myotis nesopolus". IUCN Red List of Threatened Species . 2016: e.T14184A22065759. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14184A22065759.en .
35. Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis melanorhinus". IUCN Red List of Threatened Species . 2017: e.T136784A22033542. doi: 10.2305/IUCN.UK.2017-2.RLTS.T136784A22033542.en .
36. Kruskop, S. V.; Godlevska, L.; Bücs, S.; Çoraman, E.; Gazaryan, S. (2021) [errata version of 2020 assessment]. "Myotis daubentonii". IUCN Red List of Threatened Species . 2020: e.T85342710A195858793. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85342710A195858793.en .
37. Jiang, T. L.; Feng, J. (2019). "Myotis davidii". IUCN Red List of Threatened Species . 2019: e.T136250A22003049. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136250A22003049.en .
38. Solari, S. (2017). "Myotis diminutus". IUCN Red List of Threatened Species . 2017: e.T88151417A88151431. doi: 10.2305/IUCN.UK.2017-2.RLTS.T88151417A88151431.en .
39. Barquez, R.; Diaz, M. (2016). "Myotis dinellii". IUCN Red List of Threatened Species . 2016: e.T136204A22009702. doi: 10.2305/IUCN.UK.2016-3.RLTS.T136204A22009702.en .
40. Larsen, R. (2016). "Myotis dominicensis". IUCN Red List of Threatened Species . 2016: e.T14155A22057933. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14155A22057933.en .
41. Fukui, D.; Sun, K.; Kruskop, S. V. (2019). "Myotis macrodactylus". IUCN Red List of Threatened Species . 2019: e.T14177A22065868. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14177A22065868.en .
42. Solari, S. (2018). "Myotis leibii". IUCN Red List of Threatened Species . 2018: e.T14172A22055716. doi: 10.2305/IUCN.UK.2018-2.RLTS.T14172A22055716.en .
43. Fukui, D.; Sano, A. (2020). "Myotis petax". IUCN Red List of Threatened Species . 2020: e.T85342726A85342734. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85342726A85342734.en .
44. Miller, B.; Rodriguez, B. (2017) [errata version of 2016 assessment]. "Myotis elegans". IUCN Red List of Threatened Species . 2016: e.T14156A115121563. doi: 10.2305/IUCN.UK.2016-3.RLTS.T14156A22057814.en .
45. Russo, D.; Cistrone, L. (2023). "Myotis escalerai". IUCN Red List of Threatened Species . 2023: e.T85733126A211003991. doi: 10.2305/IUCN.UK.2023-1.RLTS.T85733126A211003991.en .
46. Fukui, D.; Sano, A.; Kruskop, S. V. (2019). "Myotis bombinus". IUCN Red List of Threatened Species . 2019: e.T14149A22061650. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14149A22061650.en .
47. Juste, J.; Paunović, M. (2016). "Myotis punicus". IUCN Red List of Threatened Species . 2016: e.T44864A22073410. doi: 10.2305/IUCN.UK.2016-2.RLTS.T44864A22073410.en .
48. Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis vivesi". IUCN Red List of Threatened Species . 2016: e.T14209A22069146. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14209A22069146.en .
49. Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis findleyi". IUCN Red List of Threatened Species . 2016: e.T14159A22058800. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14159A22058800.en .
50. Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis planiceps". IUCN Red List of Threatened Species . 2016: e.T14191A22066742. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14191A22066742.en .
51. Vincenot, C. E.; Preble, J. H.; Huang, J. C. -C.; Collazo, A. M.; Kamal, A. (2021). "Myotis frater". IUCN Red List of Threatened Species . 2021: e.T85566806A22056940. doi: 10.2305/IUCN.UK.2021-2.RLTS.T85566806A22056940.en .
52. Jiang, T. L.; Feng, J. (2019). "Myotis fimbriatus". IUCN Red List of Threatened Species . 2019: e.T85735587A22058886. doi: 10.2305/IUCN.UK.2019-3.RLTS.T85735587A22058886.en .
53. Arroyo-Cabrales, J.; de Grammont, P. C. (2017). "Myotis thysanodes". IUCN Red List of Threatened Species . 2017: e.T14206A22063246. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14206A22063246.en .
54. Fukui, D.; Sano, A. (2021) [errata version of 2019 assessment]. "Myotis pruinosus". IUCN Red List of Threatened Species . 2019: e.T14192A209551299. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14192A209551299.en .
55. Piraccini, R. (2016). "Myotis emarginatus". IUCN Red List of Threatened Species . 2016: e.T14129A22051191. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14129A22051191.en .
56. Waldien, D. L.; Santiago, K.; Wortham, G.; Stronsick, S. (2021). "Myotis gomantongensis". IUCN Red List of Threatened Species . 2021: e.T40035A22060096. doi: 10.2305/IUCN.UK.2021-3.RLTS.T40035A22060096.en .
57. Solari, S. (2018). "Myotis grisescens". IUCN Red List of Threatened Species . 2018: e.T14132A22051652. doi: 10.2305/IUCN.UK.2018-2.RLTS.T14132A22051652.en .
58. Coroiu, I.; Juste, J.; Paunović, M. (2016). "Myotis myotis". IUCN Red List of Threatened Species . 2016: e.T14133A22051759. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14133A22051759.en .
59. Cajas C., J.; Miller, B. (2016). "Myotis cobanensis". IUCN Red List of Threatened Species . 2016: e.T14154A22058031. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14154A22058031.en .
60. Görföl, T.; Csorba, G.; Bates, P. J. J.; Francis, C. M.; Molur, S.; Srinivasulu, C. (2020). "Myotis annectans". IUCN Red List of Threatened Species . 2020: e.T14142A22050272. doi: 10.2305/IUCN.UK.2020-2.RLTS.T14142A22050272.en .
61. Barquez, R.; Diaz, M. (2016). "Myotis keaysi". IUCN Red List of Threatened Species . 2016: e.T14170A22056048. doi: 10.2305/IUCN.UK.2016-3.RLTS.T14170A22056048.en .
62. Csorba, G.; Hutson, A. M.; Kingston, T.; Bumrungsri, S.; Francis, C. M. (2016). "Myotis hermani". IUCN Red List of Threatened Species . 2016: e.T14165A22057251. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14165A22057251.en .
63. Santiago, K.; Wortham, G.; Waldien, D. L. (2021). "Myotis siligorensis". IUCN Red List of Threatened Species . 2021: e.T14203A22064839. doi: 10.2305/IUCN.UK.2021-3.RLTS.T14203A22064839.en .
64. Huang, J. C. -C.; Csorba, G.; Chang, H.-C.; Ho, Y.-Y. (2020). "Myotis formosus". IUCN Red List of Threatened Species . 2020: e.T85736120A95642290. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85736120A95642290.en .
65. Phelps, K.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Heaney, L.; Gonzalez, J. C.; Molur, S.; Srinivasulu, C. (2019). "Myotis horsfieldii". IUCN Red List of Threatened Species . 2019: e.T14166A22057415. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14166A22057415.en .
66. Zhigalin, A.; Stubbe, M.; Ariunbold, J.; Buuveibaatar, V.; Dorjderem, S.; Monkhzul, T.; Otgonbaatar, M.; Tsogbadrakh, M. (2020). "Myotis ikonnikovi". IUCN Red List of Threatened Species . 2020: e.T14168A22057122. doi: 10.2305/IUCN.UK.2020-3.RLTS.T14168A22057122.en .
67. Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis sodalis". IUCN Red List of Threatened Species . 2016: e.T14136A22053184. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14136A22053184.en .
68. Son, N.; Görföl, T.; Csorba, G. (2019). "Myotis indochinensis". IUCN Red List of Threatened Species . 2019: e.T85342688A85342691. doi: 10.2305/IUCN.UK.2019-3.RLTS.T85342688A85342691.en .
69. Helgen, K.; Bonaccorso, F. J. (2020). "Myotis insularum". IUCN Red List of Threatened Species . 2020: e.T14169A22055968. doi: 10.2305/IUCN.UK.2020-2.RLTS.T14169A22055968.en .
70. Solari, S. (2017). "Myotis izecksohni". IUCN Red List of Threatened Species . 2017: e.T88151563A88151572. doi: 10.2305/IUCN.UK.2017-2.RLTS.T88151563A88151572.en .
71. Kruskop, S. V. (2016). "Myotis longipes". IUCN Red List of Threatened Species . 2016: e.T14175A22056206. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14175A22056206.en .
72. Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis keenii". IUCN Red List of Threatened Species . 2017: e.T14171A22055579. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14171A22055579.en .
73. Bouillard, N. (2021). "Myotis stalkeri". IUCN Red List of Threatened Species . 2021: e.T14205A22063416. doi: 10.2305/IUCN.UK.2021-3.RLTS.T14205A22063416.en .
74. Happold. M. (2019). "Myotis dieteri". IUCN Red List of Threatened Species . 2019: e.T136678A22038629. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136678A22038629.en .
75. Jiang, T. L.; Feng, J. (2019). "Myotis chinensis". IUCN Red List of Threatened Species . 2019: e.T14152A22060946. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14152A22060946.en .
76. Bouillard, N. (2021). "Myotis adversus". IUCN Red List of Threatened Species . 2021: e.T85735326A22049231. doi: 10.2305/IUCN.UK.2021-2.RLTS.T85735326A22049231.en .
77. Gorecki, V.; Pennay, M. (2021). "Myotis macropus". IUCN Red List of Threatened Species . 2021: e.T136697A22039960. doi: 10.2305/IUCN.UK.2021-1.RLTS.T136697A22039960.en .
78. Solari, S. (2017). "Myotis lavali". IUCN Red List of Threatened Species . 2017: e.T88151601A88151604. doi: 10.2305/IUCN.UK.2017-2.RLTS.T88151601A88151604.en .
79. Bates, P. J. J.; Hutson, A. M.; Carino, A.; Kingston, T.; Lee, B. P. Y-H; Maryanto, I.; Molur, S.; Srinivasulu, C. (2020). "Myotis hasseltii". IUCN Red List of Threatened Species . 2020: e.T14164A22056644. doi: 10.2305/IUCN.UK.2020-2.RLTS.T14164A22056644.en .
80. Juste, J.; Paunović, M. (2016). "Myotis blythii". IUCN Red List of Threatened Species . 2016: e.T14124A22053297. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14124A22053297.en .
81. Solari, S. (2021) [amended version of 2018 assessment]. "Myotis lucifugus". IUCN Red List of Threatened Species . 2021: e.T14176A208031565. doi: 10.2305/IUCN.UK.2021-3.RLTS.T14176A208031565.en .
82. Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis evotis". IUCN Red List of Threatened Species . 2017: e.T14157A22059133. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14157A22059133.en .
83. Paunović, M. (2016). "Myotis capaccinii". IUCN Red List of Threatened Species . 2016: e.T14126A22054131. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14126A22054131.en .
84. Solari, S. (2019). "Myotis volans". IUCN Red List of Threatened Species . 2019: e.T14210A22069325. doi: 10.2305/IUCN.UK.2019-1.RLTS.T14210A22069325.en .
85. Vincenot, C. E.; Preble, J. H.; Collazo, A. M. (2021). "Myotis longicaudatus". IUCN Red List of Threatened Species . 2021: e.T85566977A85566980. doi: 10.2305/IUCN.UK.2021-2.RLTS.T85566977A85566980.en .
86. Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Myotis secundus". IUCN Red List of Threatened Species . 2017: e.T85342651A85342654. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85342651A85342654.en .
87. Monadjem, A.; Razafimanahaka, J.; Ranivo, J.; Kofoky, A.; Hutson, A. M.; Cardiff, S. G.; Andriafidison, D.; Goodman, S.; Jenkins, R. K. B.; Racey, P. A.; Ratrimomanarivo, F. H. (2017). "Myotis goudoti". IUCN Red List of Threatened Species . 2017: e.T14163A22056541. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14163A22056541.en .
88. Görföl, T.; Csorba, G. (2017). "Myotis federatus". IUCN Red List of Threatened Species . 2017: e.T85568302A85568305. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85568302A85568305.en .
89. Armstrong, K. N. (2021). "Myotis moluccarum". IUCN Red List of Threatened Species . 2021: e.T136770A22033795. doi: 10.2305/IUCN.UK.2021-3.RLTS.T136770A22033795.en .
90. Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis sicarius". IUCN Red List of Threatened Species . 2019: e.T14202A22063965. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14202A22063965.en .
91. Solari, S. (2018). "Myotis oxyotus". IUCN Red List of Threatened Species . 2018: e.T14187A22067211. doi: 10.2305/IUCN.UK.2018-2.RLTS.T14187A22067211.en .
92. Jacobs, D.; Cotterill, F. P. D.; Taylor, P. J. (2019). "Myotis morrisi". IUCN Red List of Threatened Species . 2019: e.T14182A22065314. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14182A22065314.en .
93. Russo, D.; Cistrone, L. (2023). "Myotis nattereri". IUCN Red List of Threatened Species . 2023: e.T215492021A211005466. doi: 10.2305/IUCN.UK.2023-1.RLTS.T215492021A211005466.en .
94. Srinivasulu, B.; Srinivasulu, C. (2019). "Myotis nipalensis". IUCN Red List of Threatened Species . 2019: e.T136495A21976309. doi: 10.2305/IUCN.UK.2019-3.RLTS.T136495A21976309.en .
95. Bakwo Fils, E. M.; Flanders, J.; Frick, W. F.; Simmons, N. (2022). "Myotis nimbaensis". IUCN Red List of Threatened Species . 2022: e.T216617275A216617367. doi: 10.2305/IUCN.UK.2022-2.RLTS.T216617275A216617367.en .
96. Solari, S. (2018). "Myotis septentrionalis". IUCN Red List of Threatened Species . 2018: e.T14201A22064312. doi: 10.2305/IUCN.UK.2018-2.RLTS.T14201A22064312.en .
97. Csorba, G.; Rosell-Ambal, R. G. B.; Tabaranza, B.; Sedlock, J.; Ingle, N. R.; Heaney, L.; Balete, D. S.; Ong, P. (2016). "Myotis rufopictus". IUCN Red List of Threatened Species . 2016: e.T136411A22017446. doi: 10.2305/IUCN.UK.2016-2.RLTS.T136411A22017446.en .
98. Duya, M. R.; Alvarez, J.; Fidelino, J.; Gatan-Balbas, M.; Pedregosa, M.; Veluz, M. J.; Jakosalem, P. G.; Tanalgo, K.; Garcia, J. J. L.; Cacho, M. A. (2019). "Myotis macrotarsus". IUCN Red List of Threatened Species . 2019: e.T14178A22065997. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14178A22065997.en .
99. Arroyo-Cabrales, J.; Ospina-Garces, S. (2016). "Myotis peninsularis". IUCN Red List of Threatened Species . 2016: e.T14189A22066405. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14189A22066405.en .
100. Wiles, G.; Furey, N. (2021). "Myotis ater". IUCN Red List of Threatened Species . 2021: e.T14144A22050847. doi: 10.2305/IUCN.UK.2021-2.RLTS.T14144A22050847.en .
101. Csorba, G.; Görföl, T. (2017). "Myotis peytoni". IUCN Red List of Threatened Species . 2017: e.T85568321A85568324. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85568321A85568324.en .
102. Piraccini, R. (2016). "Myotis dasycneme". IUCN Red List of Threatened Species . 2016: e.T14127A22055164. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14127A22055164.en .
103. Solari, S. (2019). "Myotis ruber". IUCN Red List of Threatened Species . 2019: e.T14197A22062092. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14197A22062092.en .
104. Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Myotis soror". IUCN Red List of Threatened Species . 2017: e.T85342662A85342666. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85342662A85342666.en .
105. Csorba, G.; Görföl, T. (2020). "Myotis rufoniger". IUCN Red List of Threatened Species . 2020: e.T85735909A85735913. doi: 10.2305/IUCN.UK.2020-3.RLTS.T85735909A85735913.en .
106. Jiang, T. L.; Feng, J.; Csorba, G.; Bates, P. (2019). "Myotis pilosus". IUCN Red List of Threatened Species . 2019: e.T14193A22062554. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14193A22062554.en .
107. Azhar, M. I. (2020). "Myotis ridleyi". IUCN Red List of Threatened Species . 2020: e.T14194A22062376. doi: 10.2305/IUCN.UK.2020-2.RLTS.T14194A22062376.en .
108. Barquez, R.; Perez, S.; Diaz, M. (2016). "Myotis riparius". IUCN Red List of Threatened Species . 2016: e.T14195A22062950. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14195A22062950.en .
109. Monadjem, A.; Jacobs, D. (2017). "Myotis bocagii". IUCN Red List of Threatened Species . 2017: e.T14148A22059585. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14148A22059585.en .
110. Piraccini, R. (2016). "Myotis schaubi". IUCN Red List of Threatened Species . 2016: e.T14198A22061746. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14198A22061746.en .
111. Larsen, R. (2016). "Myotis martiniquensis". IUCN Red List of Threatened Species . 2016: e.T76435251A22066280. doi: 10.2305/IUCN.UK.2016-1.RLTS.T76435251A22066280.en .
112. Benda, P.; Lavrenchenko, L. (2017). "Myotis scotti". IUCN Red List of Threatened Species . 2017: e.T14199A22062198. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14199A22062198.en .
113. Zhigalin, A. (2020). "Myotis sibiricus". IUCN Red List of Threatened Species . 2020: e.T85567062A85567065. doi: 10.2305/IUCN.UK.2020-2.RLTS.T85567062A85567065.en .
114. Barquez, R.; Diaz, M.; Samudio Jr, R.; Arroyo-Cabrales, J. (2016). "Myotis albescens". IUCN Red List of Threatened Species . 2016: e.T14140A22049892. doi: 10.2305/IUCN.UK.2016-1.RLTS.T14140A22049892.en .
115. Csorba, G.; Bates, P.; Lee, B.; Soisook, P. (2016). "Myotis oreias". IUCN Red List of Threatened Species . 2016: e.T14186A22067080. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14186A22067080.en .
116. Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis austroriparius". IUCN Red List of Threatened Species . 2017: e.T14147A22059907. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14147A22059907.en .
117. Barquez, R.; Diaz, M. (2017) [errata version of 2016 assessment]. "Myotis aelleni". IUCN Red List of Threatened Species . 2016: e.T14139A115121458. doi: 10.2305/IUCN.UK.2016-3.RLTS.T14139A22049723.en .
118. Arroyo-Cabrales, J.; Miller, B.; Reid, F.; Cuarón, A. D.; de Grammont, P. C. (2017). "Myotis auriculus". IUCN Red List of Threatened Species . 2017: e.T14145A22060698. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14145A22060698.en .
119. Jiang, T. L.; Feng, J. (2019). "Myotis altarium". IUCN Red List of Threatened Species . 2019: e.T14141A22050057. doi: 10.2305/IUCN.UK.2019-3.RLTS.T14141A22050057.en .
120. Csorba, G.; Bumrungsri, S.; Bates, P. (2020). "Myotis rosseti". IUCN Red List of Threatened Species . 2020: e.T14196A22062800. doi: 10.2305/IUCN.UK.2020-2.RLTS.T14196A22062800.en .
121. Barquez, R.; Diaz, M. (2016). "Myotis simus". IUCN Red List of Threatened Species . 2016: e.T14204A22064642. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14204A22064642.en .
122. Srinivasulu, C.; Srinivasulu, B. (2019). "Myotis muricola". IUCN Red List of Threatened Species . 2019: e.T85537578A22065403. doi: 10.2305/IUCN.UK.2019-3.RLTS.T85537578A22065403.en .
123. Bouillard, N.; Csorba, G.; Görföl, T. (2021). "Myotis weberi". IUCN Red List of Threatened Species . 2021: e.T85736011A85736023. doi: 10.2305/IUCN.UK.2021-2.RLTS.T85736011A85736023.en .
124. Monadjem, A.; Taylor, P. J.; Jacobs, D.; Cotterill, F. (2017). "Myotis welwitschii". IUCN Red List of Threatened Species . 2017: e.T14211A22068792. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14211A22068792.en .
125. Arroyo-Cabrales, J.; Álvarez-Castañeda, S. T. (2017). "Myotis ciliolabrum". IUCN Red List of Threatened Species . 2017: e.T14153A22058110. doi: 10.2305/IUCN.UK.2017-2.RLTS.T14153A22058110.en .
126. Coroiu, I. (2016). "Myotis mystacinus". IUCN Red List of Threatened Species . 2016: e.T14134A22052250. doi: 10.2305/IUCN.UK.2016-2.RLTS.T14134A22052250.en .
127. Fukui, D.; Sano, A. (2021) [errata version of 2019 assessment]. "Myotis yanbarensis". IUCN Red List of Threatened Species . 2019: e.T29484A209551473. doi: 10.2305/IUCN.UK.2019-3.RLTS.T29484A209551473.en .
128. Barquez, R.; Diaz, M. (2017) [errata version of 2016 assessment]. "Myotis levis". IUCN Red List of Threatened Species . 2016: e.T14174A115121699. doi: 10.2305/IUCN.UK.2016-3.RLTS.T14174A22056440.en .
129. Solari, S. (2019). "Myotis yumanensis". IUCN Red List of Threatened Species . 2019: e.T14213A22068335. doi: 10.2305/IUCN.UK.2019-1.RLTS.T14213A22068335.en .
130. Ruedi, M.; Csorba, G.; Liang-Kong, L.; Cheng-Han, C. (2017). "Submyotodon latirostris". IUCN Red List of Threatened Species . 2017: e.T85537971A85537974. doi: 10.2305/IUCN.UK.2017-2.RLTS.T85537971A85537974.en .

Sources

• Chernasky, Amy; Motis, Anna; Burgin, Connor, eds. (2023). All the Mammals of the World . Lynx Nature Books. ISBN   978-84-16728-66-4.
• Simmons, Nancy B. (2005). Wilson, Don E.; Reeder, DeeAnn M. (eds.). Mammal Species of the World . Vol. 1 (3rd ed.). Johns Hopkins University Press. ISBN   978-0-8018-8221-0.
• Nowak, Ronald M. (1994). Walker's Bats of the World . Johns Hopkins University Press. ISBN   978-0-8018-4986-2.