Orionides

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Orionidans
Temporal range:
Middle Jurassic Present, 175.6–0  Ma
O
S
D
C
P
T
J
K
Pg
N
Possible Early Jurassic record
Orionides.jpg
1st row (carnosaurs/non-coelurosaurian orionidans):
Allosaurus fragilis , Spinosaurus aegyptiacus , Torvosaurus tanneri ;

2nd row (non-maniraptoran coelurosaurians):
Gallimimus bullatus , Tyrannosaurus rex , Compsognathus longipes ;
3rd row (non-eumaniraptoran maniraptorans):
Alvarezsaurus calvoi , Alxasaurus elesitaiensis , Avimimus portentosus ;
4th row (eumaniraptorans):
Velociraptor mongoliensis , Hesperornis regalis , Ardenna carneipes .

Contents

Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Tetanurae
Clade: Orionides
Carrano, Benson & Sampson, 2012
Clades
Synonyms

Orionides is a clade of tetanuran theropod dinosaurs from the Middle Jurassic to the Present. The clade includes most theropod dinosaurs, including birds. [2]

Relationships

Although many phylogenetic analyses found basal tetanurans that were outside both Megalosauroidea and Avetheropoda, the core dichotomy was named only in 2012. Carrano, Benson and Sampson (2012) named that clade Orionides, and defined it as the node comprising Megalosauroidea, Avetheropoda, their most recent common ancestor, and all its descendants. [2] In 2015, Hendrickx, Hartman and Mateus clarified this definition, specifying it as the least inclusive clade including Allosaurus fragilis , Megalosaurus bucklandii , and Passer domesticus . [3] The cladogram presented below follows a phylogenetic analysis published by Zanno and Makovicky in 2013. [4]

Orionides
Megalosauroidea

Piatnitzkysauridae Piatnitzkysaurus floresi by Paleocolour.jpg

Megalosauria

Spinosauridae Spinosaurus aegyptiacus.png

Megalosauridae Torvosaurus tanner DBi.jpg

Avetheropoda

Coelurosauria Meyers grosses Konversations-Lexikon - ein Nachschlagewerk des allgemeinen Wissens (1908) (Antwerpener Breiftaube).jpg

Allosauroidea

Metriacanthosauridae Yangchuanosaurus NT (flipped).jpg

Allosauria

Allosauridae Allosaurus Revised.jpg

Carcharodontosauria

Neovenatoridae Neovenator.png

Carcharodontosauridae Shaochilong.jpg

In 2019, a basal Allosauroid by the name of Asfaltovenator was discovered that provided more evidence to the idea of a monophyletic Carnosauria that contained both Megalosauroid and Allosauroid groups. While this did cast doubt on the validity of Avetheropoda, Orionides as group was still maintained however this could also imply that the name Orionides is synonymous with Avetheropoda.

Range

Large, predatory spinosaurids and allosaurids flourished during the Late Jurassic and Early Cretaceous, especially in Gondwana, but seem to have died out before the end of the Cretaceous, possibly due to competition from abelisaurid ceratosaurs and tyrannosaurid coelurosaurs. The diverse coelurosaurs persisted until the end of the Mesozoic Era, when all except for crown clade avians died out. Modern birds are the only living representatives of the clade Tetanurae.

Etymology

The clade name "Orionides" was first established by Matthew T. Carrano, Roger B. J. Benson and Scott D. Sampson in 2012. It is derived from Orion, the giant hunter of Greek mythology in references to the large size and carnivorism of basal orionidans. The name also refers to the alternative name for the constellation of Orion, Alektropodion, meaning "rooster foot". [2]

Related Research Articles

<span class="mw-page-title-main">Coelurosauria</span> Clade of dinosaurs

Coelurosauria is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.

<span class="mw-page-title-main">Carnosauria</span> Extinct Infraorder of theropod dinosaurs

Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as Giganotosaurus, Mapusaurus, Carcharodontosaurus and Tyrannotitan which are among the largest known predatory dinosaurs.

<span class="mw-page-title-main">Tetanurae</span> Clade containing most theropod dinosaurs

Tetanurae is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans. Tetanurans are defined as all theropods more closely related to modern birds than to Ceratosaurus and contain the majority of predatory dinosaur diversity. Tetanurae likely diverged from its sister group, Ceratosauria, during the late Triassic. Tetanurae first appeared in the fossil record by the Early Jurassic about 190 mya and by the Middle Jurassic had become globally distributed.

<i>Afrovenator</i> Extinct genus of dinosaurs

Afrovenator is a genus of megalosaurid theropod dinosaur from the Middle or Late Jurassic Period on the Tiourarén Formation and maybe the Irhazer II Formation of the Niger Sahara region in northern Africa. Afrovenator represents the only properly identified Gondwanan megalosaur, with proposed material of the group present in the Late Jurassic on Tacuarembó Formation of Uruguay and the Tendaguru Formation of Tanzania.

<span class="mw-page-title-main">Allosauridae</span> Extinct family of theropod dinosaurs

Allosauridae is a family of medium to large bipedal, carnivorous allosauroid theropod dinosaurs from the Late Jurassic. Allosauridae is a fairly old taxonomic group, having been first named by the American paleontologist Othniel Charles Marsh in 1878. Allosaurids are characterized by an astragalus with a restriction of the ascending process to the lateral part of the bone, a larger medial than lateral condyle, and a horizontal groove across the face of the condyles.

<span class="mw-page-title-main">Megalosauridae</span> Extinct family of dinosaurs

Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively primitive group of basal tetanurans containing two main subfamilies, Megalosaurinae and Afrovenatorinae, along with the basal genus Eustreptospondylus, an unresolved taxon which differs from both subfamilies.

<i>Megaraptor</i> Extinct genus of dinosaurs

Megaraptor is a genus of large theropod dinosaur that lived in the ages of the Late Cretaceous. Its fossils have been discovered in the Patagonian Portezuelo Formation of Argentina, South America. Initially thought to have been a giant dromaeosaur-like coelurosaur, it was classified as a neovenatorid allosauroid in previous phylogenies, but more recent phylogeny and discoveries of related megaraptoran genera has placed it as either a basal tyrannosauroid or a basal coelurosaur with some studies still considering it a neovenatorid.

<i>Monolophosaurus</i> Extinct genus of dinosaurs

Monolophosaurus is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China. It was named for the single crest on top of its skull. Monolophosaurus was a mid-sized theropod at about 5–5.5 metres (16–18 ft) long and weighed 475 kilograms (1,047 lb).

<i>Piatnitzkysaurus</i> Extinct genus of dinosaurs

Piatnitzkysaurus is a genus of megalosauroid theropod dinosaur that lived approximately 179 to 177 million years ago during the lower part of the Jurassic Period in what is now Argentina. Piatnitzkysaurus was a moderately large, lightly built, bipedal, ground-dwelling carnivore that could grow up to 6.6 m (21.7 ft) long.

<i>Siamotyrannus</i> Extinct genus of dinosaurs

Siamotyrannus is a genus of theropod dinosaur from the early Cretaceous of Thailand.

<span class="mw-page-title-main">Megalosauroidea</span> Extinct superfamily of Dinosaurs

Megalosauroidea is a superfamily of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as Megalosaurus bucklandii and all taxa sharing a more recent common ancestor with it than with Allosaurus fragilis or Passer domesticus. Members of the group include Spinosaurus, Megalosaurus, and Torvosaurus. They are possibly paraphyletic in nature with respect to Allosauroidea.

Chuandongocoelurus is a genus of carnivorous tetanuran theropod dinosaur from the Jurassic of China.

<span class="mw-page-title-main">Metriacanthosauridae</span> Extinct family of dinosaurs

Metriacanthosauridae is an extinct family of allosauroid theropod dinosaurs that lived from the Middle Jurassic to the Early Cretaceous. When broken down into its Greek roots, it means "moderately-spined lizards". The family is split into two subgroups: Metriacanthosaurinae, which includes dinosaurs closely related to Metriacanthosaurus, and another group composed of the close relatives of Yangchuanosaurus. Metriacanthosaurids are considered carnosaurs, belonging to the Allosauroidea superfamily. The group includes species of large range in body size. Of their physical traits, most notable are their neural spines. The records of the group are mostly confined to Asia, though Metriacanthosaurus is known from Europe. Metriacanthosauridae is used as a senior synonym of Sinraptoridae.

<span class="mw-page-title-main">Allosauroidea</span> Extinct superfamily of Dinosaurs

Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among the apex predators that were active during the Middle Jurassic to Late Cretaceous periods. The most famous and best understood allosauroid is the North American genus Allosaurus.

<i>Lourinhanosaurus</i> Extinct genus of dinosaurs

Lourinhanosaurus was a genus of carnivorous theropod dinosaur that lived during the Late Jurassic Period (Kimmeridgian/Tithonian) in Portugal. It is one of many large predators discovered at the Lourinhã Formation and probably competed with coeval Torvosaurus gurneyi, Allosaurus europaeus, and Ceratosaurus.

<span class="mw-page-title-main">Avetheropoda</span> Clade of dinosaurs

Avetheropoda, or "bird theropods", is a clade that includes Carnosaurs and coelurosaurs to the exclusion of other Dinosaurs.

<span class="mw-page-title-main">Neovenatoridae</span> Extinct family of dinosaurs

Neovenatoridae is a proposed clade of carcharodontosaurian dinosaurs uniting some primitive members of the group such as Neovenator with the Megaraptora, a group of theropods with controversial affinities. Other studies recover megaraptorans as basal coelurosaurs unrelated to carcharodontosaurs. Other theropods with uncertain affinities such as Gualicho, Chilantaisaurus and Deltadromeus are also sometimes included.

<span class="mw-page-title-main">Megaraptora</span> Extinct clade of dinosaurs

Megaraptora is a clade of carnivorous theropod dinosaurs. Its derived members, the Megaraptoridae are noted for their large hand claws and powerfully-built forelimbs, which are usually reduced in size in other large theropods.

<span class="mw-page-title-main">Piatnitzkysauridae</span> Extinct family of dinosaurs

Piatnitzkysauridae is an extinct family of megalosauroid or basal allosauroid dinosaurs. It only consists of three to four known dinosaur genera: Condorraptor, Marshosaurus, Piatnitzkysaurus and possibly Xuanhanosaurus. The most complete and well known member of this family is Piatnitzkysaurus, which also gives the family its name.

<i>Sciurumimus</i> Extinct species of reptile

Sciurumimus is an extinct genus of tetanuran theropod from the Late Jurassic Torleite Formation of Germany. It is known from a single juvenile specimen representing the type species, Sciurumimus albersdoerferi, which was found in a limestone quarry close to Painten in Lower Bavaria. The specimen was preserved with traces of feather-like filaments.

References

  1. Rauhut, Oliver W. M.; Pol, Diego (2019-12-11). "Probable basal allosauroid from the early Middle Jurassic Cañadón Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs". Scientific Reports. 9 (1): 18826. Bibcode:2019NatSR...918826R. doi:10.1038/s41598-019-53672-7. ISSN   2045-2322. PMC   6906444 . PMID   31827108.
  2. 1 2 3 Carrano, M. T.; Benson, R. B. J.; Sampson, S. D. (2012). "The phylogeny of Tetanurae (Dinosauria: Theropoda)". Journal of Systematic Palaeontology. 10 (2): 211–300. doi:10.1080/14772019.2011.630927.
  3. Hendrickx, C.; Hartman, S.A.; Mateus, O. (2015). "An overview of non-avian theropod discoveries and classification". PalArch's Journal of Vertebrate Palaeontology. 12 (1): 1–73.
  4. Zanno, L. E.; Makovicky, P. J. (2013). "Neovenatorid theropods are apex predators in the Late Cretaceous of North America". Nature Communications. 4: 2827. Bibcode:2013NatCo...4.2827Z. doi: 10.1038/ncomms3827 . PMID   24264527.