List of Archaea genera

Last updated

This article lists the genera of the Archaea. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) [1] and National Center for Biotechnology Information (NCBI). [2] However, in the List provided below, GTDB has precedence unless otherwise noted.

Contents

Phylogeny

National Center for Biotechnology Information (NCBI) taxonomy was initially used to decorate the genome tree via tax2tree. The 16S rRNA-based Greengenes taxonomy is used to supplement the taxonomy particularly in regions of the tree with no cultured representatives. List of Prokaryotic names with Standing in Nomenclature (LPSN) is used as the primary taxonomic authority for establishing naming priorities. Taxonomic ranks are normalised using phylorank and the taxonomy manually curated to remove polyphyletic groups. [3] [4] [5] [6] [7]

Cladogram was taken from the GTDB release 10-RS226 (16th April 2025). [8] [9] [10] The position of clades with a "question mark" are based on the additional phylogeny of the 16S rRNA-based LTP_12_2021 by The All-Species Living Tree Project. [11] [12] [13]

Archaea

Phylum Microcaldota

Class Microcaldia

Order Microcaldales

  • Family Microcaldaceae Sakai et al. 2023

Phylum "Altarchaeota"

Class "Altarchaeia"

Order "Altarchaeales"

  • Family "Altarchaeaceae" corrig. Probst et al. 2014

Phylum "Iainarchaeota"

Class "Iainarchaeia"

Order "Forterreales"

  • Family "Forterreaceae" Pallen, Rodriguez-R & Alikhan 2022
    • "Candidatus Forterrea" Probst & Banfield 2017

Order "Iainarchaeales"

  • Family "Iainarchaeaceae" Rinke et al. 2020

Phylum "Micrarchaeota"

Phylogeny of Micrarchaeota [8] [9] [10]
"Norongarragalinales"

"Ca. Norongarragalina"

"Micrarchaeales"
"Micrarchaeaceae"
"Anstonellales"
"Anstonellaceae"

"Ca. Anstonella"

"Bilamarchaeaceae"

"Ca. Bilamarchaeum"

"Gugararchaeales"

"Ca. Gugararchaeum"

"Burarchaeales"

"Ca. Burarchaeum"

"Fermentimicrarchaeales"

"Ca. Fermentimicrarchaeum"

Class "Micrarchaeia"

Order "Norongarragalinales"

  • Family "Norongarragalinaceae" Vázquez-Campos et al. 2021

Order "Micrarchaeales"

  • Family "Micrarchaeaceae" Vázquez-Campos et al. 2021

Order "Anstonellales"

  • Family "Anstonellaceae" Vázquez-Campos et al. 2021
    • "Candidatus Anstonella" Vázquez-Campos et al. 2021
  • Family "Bilamarchaeaceae" Vázquez-Campos et al. 2021

Order "Gugararchaeales"

  • Family "Gugararchaeaceae" Vázquez-Campos et al. 2021

Order "Burarchaeales"

  • Family "Burarchaeaceae" Vázquez-Campos et al. 2021

Order "Fermentimicrarchaeales"

  • Family "Fermentimicrarchaeaceae" Kadnikov et al. 2020 (Sv326)

Phylum "Undinarchaeota"

Class "Undinarchaeia"

Order "Undinarchaeales"

  • Family "Naiadarchaeaceae" Dombrowski et al. 2020
  • Family "Undinarchaeaceae" Dombrowski et al. 2020

Phylum "Huberarchaeaota"

Class "Huberarchaeia"

Order "Huberarchaeales"

  • Family "Huberarchaeaceae" Rinke et al. 2020

Phylum "Aenigmatarchaeota"

Class "Aenigmatarchaeia"

Order "Aenigmatarchaeales"

  • Family "Aenigmatarchaeaceae" corrig. Rinke et al. 2020

Phylum "Nanohalarchaeota"

Phylogeny of Nanohalobiales [8] [9] [10]
"Nanohalalkaliarchaeaceae"

"Candidatus Nanohalalkaliarchaeum"

"Nanoanaerosalinaceae"

"Candidatus Asbonarchaeum"

"Candidatus Nanoanaerosalina"

"Nanohalobiaceae"

"Candidatus Nanosalinicola"

"Candidatus Nanohalococcus"

"Candidatus Nanohalobium"

"Candidatus Nanohalovita"

"Candidatus Nanosalina"

Class "Nanohalarchaeia"

Order "Nanohalarchaeales"

  • Family "Nanohalarchaeaceae"

Order "Nanohydrothermales"

  • Family "Nanohydrothermaceae" Xie et al. 2022

Order "Nucleotidisoterales"

  • Family "Nanosalenecaceae" Xie et al. 2022
  • Family "Nucleotidisoteraceae" Xie et al. 2022
  • Family "Nucleotidivindicaceae" Xie et al. 2022

Class "Nanohalobiia"

Order "Nanohalobiales"

Phylum Nanobdellota

Phylogeny of Nanobdellota [8] [9] [10]
Nanobdellales
"Nanoarchaeaceae"

" Nanoarchaeum "

Nanobdellaceae
"Tiddalikarchaeales"
"Tiddalikarchaeaceae"

"Ca. Tiddalikarchaeum"

"Jingweiarchaeales"
"Jingweiarchaeaceae"

"Ca. Jingweiarchaeum"

"JAPDLS01"
"Haiyanarchaeaceae"

"Ca. Haiyanarchaeum"

"Parvarchaeales"
"Parvarchaeaceae"

Class Nanobdellia

Nanobdellia incertae sedis

Order JAPDLS01

  • Family "Haiyanarchaeaceae" Rao et al. 2023

Order "Jingweiarchaeales"

  • Family "Jingweiarchaeaceae" Rao et al. 2023

Order Nanobdellales

  • Family "Nanoarchaeaceae" Huber et al. 2011
  • Family Nanobdellaceae Kato et al. 2022 ["Nanopusillaceae" Huber et al. 2011]
    • Nanobdella Kato et al. 2022 ["Candidatus Nanobsidianus" Castelle et al. 2015; "Candidatus Nanopusillus" Wurch et al. 2016]
    • "Candidatus Nanoclepta" St. John et al. 2019

Order "Tiddalikarchaeales"

  • Family "Tiddalikarchaeaceae" Vázquez-Campos et al. 2021

Order "Parvarchaeales"

Phylum Promethearchaeota

Phylogeny of Promethearchaeota [8] [9] [10]

Promethearchaeota incertae sedis

Class "Asgardarchaeia"

Order "Asgardarchaeales"

  • Family "Asgardarchaeaceae" Eme et al. 2023 ex Tamarit et al. 2024

Class "Atabeyarchaeia"

Order "Atabeyarchaeales"

  • Family "Asgardarchaeaceae" Eme et al. 2023 ex Tamarit et al. 2024

Class "Sifarchaeia"

Order "Borrarchaeales"

  • Family "Borrarchaeaceae" Liu et al. 2021

Order "Sifarchaeales"

  • Family "Sifarchaeaceae" Sun et al. 2021
    • "Candidatus Sifarchaeum" corrig. Farag, Zhao & Biddle 2020

Class "Wukongarchaeia"

Order "Wukongarchaeales"

  • Family "Wukongarchaeaceae" Liu et al. 2021

Class "Njordarchaeia"

Order "Njordarchaeales"

Class "Heimdallarchaeia"

Order "Hodarchaeales"

  • Family "Hodarchaeaceae" Liu et al. 2021

Order "Gerdarchaeales"

Order "Heimdallarchaeales"

  • Family "Kariarchaeaceae" Liu et al. 2021
  • Family "Heimdallarchaeaceae" ?Wu et al. 2022

Class "Odinarchaeia"

Order "Odinarchaeales"

  • Family "Odinarchaeaceae" Tamarit et al. 2022

Class "Jordarchaeia"

Order "Jordarchaeales"

  • Family "Freyrarchaeaceae" Xie et al. 2022
  • Family "Jordarchaeaceae" Sun et al. 2021

Class "Baldrarchaeia"

Order "Baldrarchaeales"

  • Family "Baldrarchaeaceae" corrig. Liu et al. 2021

Class "Thorarchaeia"

Order "Thorarchaeales"

Class "Hermodarchaeia"

Order "Hermodarchaeales"

  • Family "Hermodarchaeaceae" corrig. Liu et al. 2021

Class Promethearchaeia

Order "Helarchaeales"

Order Promethearchaeales

In GTDB as order CR-4

Phylogeny of Promethearchaeales [8] [9] [10]
"Sigynarchaeaceae"

"Ca. Sigynarchaeum"

Promethearchaeaceae

Phylum "Korarchaeota"

Class "Korarchaeia"

Order "Korarchaeales"

Phylum Thermoproteota

Phylogeny of Thermoproteota [8] [9] [10]

Thermoproteota Incertae sedis

Class "Bathyarchaeia"

Order "Bifangarchaeales"

  • Family "Bifangicolaceae" Hou et al. 2023
    •  ?"Candidatus Bifangicola" Hou et al. 2023 [3300025546_26]
  • Family "Bifangarchaeaceae" Hou et al. 2023
    • "Candidatus Bifangarchaeum" corrig. Hou et al. 2023 [B24: DTBU01 sp011374155]
  • Family "Bifangistellaceae" Hou et al. 2023
    • "Candidatus Bifangistella" Hou et al. 2023 [DRYS01: DRYS01 sp011367275]

Order "Xuanwuarculales"

  • Family "Mazuisomaceae" Hou et al. 2023
  • Family "Mazuicellaceae" Hou et al. 2023
  • Family "Mazuousiaecae" Hou et al. 2023
  • Family "Xuanwuarculaceae" Hou et al. 2023
    • "Candidatus Xuanwuarcula" Hou et al. 2023 [RBG-16-48-13 sp001775995]

Order "Hecatellales"

Phylogeny of "Hecatellaceae" [8] [9] [10]
  • Family "Hecatellaceae" Rinke et al. 2020

Order "Houtuarculales"

Phylogeny of Houtuarculales [8] [9] [10]
"Crenohoutuithermaceae"

"Ca. Crenohoutuithermus"

"Fontihoutuithermaceae"

"Ca. Fontihoutuithermus"

"Aphrohoutuibellaceae"
"Houtuousiaceae"
  • Family "Houtuithermaceae" Hou et al. 2023
  • Family "Marihoutuimonadaceae" Hou et al. 2023
  • Family "Crenohoutuithermaceae" Hou et al. 2023
  • Family "Fontihoutuithermaceae" Hou et al. 2023
  • Family "Aphrohoutuibellaceae" Hou et al. 2023
  • Family "Houtuousiaceae" Hou et al. 2023

Order "Wuzhiqiibiales"

Phylogeny of "Wuzhiqiibiales" [8] [9] [10]
"Wuzhiqiimorphaceae"

"Ca. Wuzhiqiimorpha"

"Wuzhiqiimonadaceae"

"Ca. Wuzhiqiimonas"

"Wuzhiqiibiaceae"

"Ca. Wuzhiqiibium"

  • Family "Wuzhiqiithermaceae" Hou et al. 2023
  • Family "Wuzhiqiitorraceae" Hou et al. 2023
  • Family "Wuzhiqiimorphaceae" Hou et al. 2023
    • "Candidatus Wuzhiqiimorpha" Hou et al. 2023 [PIYN01: PIYN01 sp003601775]
  • Family "Wuzhiqiimonadaceae" Hou et al. 2023
  • Family "Wuzhiqiibiaceae" Hou et al. 2023

Order "Zhuquarculales"

  • Family "Zhuquarculaceae" Hou et al. 2023
  • Family "Zhuquomorphaceae" Hou et al. 2023

Order "Bathyarchaeales"

Phylogeny of "Bathyarchaeales" [8] [9] [10]
"Pontizhurongicolaceae"

"Ca. Pontizhurongicola"

"Hanbarchaeaceae"
"Marizhurongicellaceae"
"Hanbopyraceae"

"Ca. Hanbopyrus"

"Jingweiiplasmataceae"

"Ca. Jingweiiplasma"

"Bathyarchaeaceae"
"Bathycorpusculaceae"

Class Nitrososphaeria_A

Order "Caldarchaeales"

Phylogeny of "Caldarchaeales" [8] [9] [10]
"Calditenuaceae"

"Ca. Calditenuis"

"Caldarchaeaceae"
"Wolframiiraptoraceae"
  • Family "Calditenuaceae" Balbay et al. 2023
  • Family "Caldarchaeaceae" Rinke et al. 2020
  • Family "Wolframiiraptoraceae" Buessecker et al. 2022

Class Nitrososphaeria

Nitrososphaeria Incertae sedis

Order "Nitrosomirales"

Order "Geothermarchaeales"

Order PSMU01

Order "Conexivisphaerales"

  • Family "Conexivisphaeraceae" Kato et al. 2021 (THSC Group)

Order Nitrososphaerales

Phylogeny of Nitrososphaerales [8] [9] [10]

Class "Methanomethylicia"

Methanomethylicia incertae sedis

  • Order "Methanohydrogenicales"
    • Family "Methanohydrogenicaceae" Berghuis et al. 2019
  • Order "Methanomethylarchaeales"
    • Family "Methanomethylarchaeceae" corrig. Hua et al. 2019
  • Order "Methanomethylovorales"
    • Family "Methanomethylovoraceae" Hua et al. 2019

Order "Nezhaarchaeales"

Order "Culexarchaeles"

  • Family "Culexarchaeaceae" Kohtz et al. 2022 ["Methanomediaceae" Berghuis et al. 2019]
    • "Candidatus Culexarchaeum" Kohtz et al. 2022 ["Candidatus Methanomedium" Berghuis et al. 2019]
  • Family "Culexmicrobiaceae" Kohtz et al. 2022

Order "Methanomethylicales"

Class Thermoproteia

Order "Gearchaeales"

  • Family "Gearchaeaceae" corrig.

Order Thermofilales

Order Thermoproteales

Phylogeny of Thermoproteales [8] [9] [10]

Class "Sulfolobia"

Order "Marsarchaeales"

Order Sulfolobales

Phylogeny of Sulfolobales [8] [9] [10]

Phylum "Hadarchaeota"

Class "Persephonarchaeia"

Class "Hadarchaeia"

Order "Hadarchaeales"

  • Family "Cerberiarchaeaceae" Benito Merino et al. 2024
  • Family "Hadarchaeaceae" Chuvochina et al. 2019 ex Chuvochina et al. 2023

Phylum Methanobacteriota_B

Phylogeny of Methanobacteriota_B [8] [9] [10]

Class "Theionarchaeia"

Class Thermococci

Order "Methanofastidiosales"

  • Family "Methanofastidiosaceae" Rinke et al. 2020

Order Thermococcales

Phylum "Hydrothermarchaeota"

Class "Hydrothermarchaeia"

Order "Hydrothermarchaeales"

  • Family BMS3B
  • Family "Hydrothermarchaeaceae" Chuvochina et al. 2019

Phylum Methanobacteriota

Phylogeny of Methanobacteriota [8] [9] [10]

Class "Methanococcia"

Order Methanococcales

Class Methanopyria

Order Methanopyrales

Class Methanobacteriia

Order Methanobacteriales

Phylum Thermoplasmatota

Phylogeny of Thermoplasmatota [8] [9] [10]

Class "Izemarchaea"

Order "Thermoprofundales"

  • (MBG-D, E2)

Class "Poseidoniia"

Order MGIII

Order "Poseidoniales"

  • Family "Poseidoniaceae" corrig. Rinke et al. 2019
  • Family "Thalassarchaeaceae" corrig. Rinke et al. 2019

Class Thermoplasmata

Order "Lunaplasmatales"

  • Family "Lunaplasmataceae" Zinke et al. 2021

Order "Proteinoplasmatales"

  • (SG8-5)

Order "Yaplasmatales"

  • (RBG-16-68-12)

Order "Gimiplasmatales"

  • Family "Gimiplasmataceae" Hu et al. 2020

Order "Sysuiplasmatales"

  • Family "Sysuiplasmataceae" Yuan et al. 2021

Order Methanomassiliicoccales

Phylogeny of Methanomassiliicoccales [8] [9] [10]

Order "Halarchaeoplasmatales"

Phylogeny of Halarchaeoplasmatales [8] [9] [10]

"Candidatus Haladaptiplasma"

"Candidatus Halarchaeoplasma"

"Candidatus Natronoplasma"

"Candidatus Saliniplasma"

Order "Lutacidiplasmatales"

  • Family "Lutacidiplasmataceae" Sheridan et al. 2022

Order "Aciduliprofundales"

  • Family "Aciduliprofundaceae" Rinke et al. 2020

Order Thermoplasmatales

Phylogeny of Thermoplasmatales [8] [9] [10]

Phylum Halobacteriota

Phylogeny of Halobacteriota [8] [9] [10]

Halobacteriota incertae sedis

Order "Methanoflorentales"

  • Family "Methanoflorentaceae" Mondav et al. 2014

Class "Methanonatronarchaeia"

Order "Methanonatronarchaeales"

Class "Methanoliparia"

Order "Methanoliparales"

Class "Archaeoglobia"

Phylogeny of "Archaeoglobia" [8] [9] [10]
"Mnemosynellales"
"Mnemosynellaceae"

"Candidatus Mnemosynella"

Archaeoglobales

Order "Mnemosynellales"

  • Family "Mnemosynellaceae" Adam et al. 2022

Order Archaeoglobales

Class "Syntropharchaeia"

Phylogeny of "Syntropharchaeia" [8] [9] [10]
"Syntropharchaeales"
"Syntropharchaeaceae"

"Candidatus Syntropharchaeum"

"Alkanophagales"
"Alkanophagaceae"

"Candidatus Alkanophaga"

"Methanospiraraceae"

"Candidatus Methanoxibalbensis"

"Candidatus Methanospirare"

Order "Syntropharchaeales"

  • Family "Syntropharchaeaceae" corrig. (ANME-2 cluster)

Order "Alkanophagales"

  • Family "Methanophagaceae" Chadwick et al. 2022
  • Family ?"Veteromethanophagaceae" Vulcano et al. 2022
  • Family "Alkanophagaceae" [B39-G2]
  • Family "Methanospiraraceae" Laso-Pérez et al. 2022

Class Methanocellia

Order Methanocellales

Phylogeny of Methanocellales [8] [9] [10]

Class Methanosarcinia

Order Methanotrichales

Order Methanosarcinales_A

  • Family Methermicoccaceae Cheng et al. 2007

Order Methanosarcinales

Phylogeny of Methanosarcinales [8] [9] [10]

Class Methanomicrobia

Order Methanomicrobiales

Phylogeny of Methanomicrobiales [8] [9] [10]

Class "Hikarchaeia"

Order "Hikarchaeales"

  • Family "Hikarchaeaceae" Martijn et al. 2020 (MG-IV)

Class "Ordosarchaeia"

Order "Ordosarchaeales"

  • Family "Ordosarchaeaceae" Zhao et al. 2024

Class Halobacteria

Order Halorutilales

  • Family Halorutilaceae Durán-Viseras et al. 2023

Order Halobacteriales

Phylogeny of Halobacteriales [8] [9] [10]
Phylogeny of Natrialbaceae [8] [9] [10]
Phylogeny of Haloferacaceae [8] [9] [10]

See also

References

  1. Domain Archaea in LPSN ; Parte, Aidan C.; Sardà Carbasse, Joaquim; Meier-Kolthoff, Jan P.; Reimer, Lorenz C.; Göker, Markus (1 November 2020). "List of Prokaryotic names with Standing in Nomenclature (LPSN) moves to the DSMZ". International Journal of Systematic and Evolutionary Microbiology. 70 (11): 5607–5612. doi: 10.1099/ijsem.0.004332 .
  2. Sayers; et al. "Archaea". National Center for Biotechnology Information (NCBI) taxonomy database. Retrieved 2021-03-20.
  3. Parks, DH; Chuvochina, M; Waite, DW; Rinke, C; Skarshewski, A; Chaumeil, PA; Hugenholtz, P (November 2018). "A standardized bacterial taxonomy based on genome phylogeny substantially revises the tree of life". Nature Biotechnology. 36 (10): 996–1004. bioRxiv   10.1101/256800 . doi:10.1038/nbt.4229. PMID   30148503. S2CID   52093100.
  4. Parks, DH; Chuvochina, M; Chaumeil, PA; Rinke, C; Mussig, AJ; Hugenholtz, P (September 2020). "A complete domain-to-species taxonomy for Bacteria and Archaea". Nature Biotechnology. 38 (9): 1079–1086. bioRxiv   10.1101/771964 . doi:10.1038/s41587-020-0501-8. PMID   32341564. S2CID   216560589.
  5. Chaumeil, PA; Mussig, AJ; Hugenholtz, P; Parks, DH (15 November 2019). "GTDB-Tk: a toolkit to classify genomes with the Genome Taxonomy Database". Bioinformatics. 36 (6): 1925–1927. doi: 10.1093/bioinformatics/btz848 . PMC   7703759 . PMID   31730192.
  6. Almeida, Alexandre; Nayfach, Stephen; Boland, Miguel; Strozzi, Francesco; Beracochea, Martin; Shi, Zhou Jason; Pollard, Katherine S.; Sakharova, Ekaterina; Parks, Donovan H.; Hugenholtz, Philip; Segata, Nicola; Kyrpides, Nikos C.; Finn, Robert D. (20 July 2020). "A unified catalog of 204,938 reference genomes from the human gut microbiome". Nature Biotechnology. 39 (1): 105–114. doi: 10.1038/s41587-020-0603-3 . PMC   7801254 . PMID   32690973.
  7. Nayfach, Stephen; et al. (9 November 2020). "A genomic catalog of Earth's microbiomes". Nature Biotechnology. 39 (4): 499–509. doi: 10.1038/s41587-020-0718-6 . PMC   8041624 . PMID   33169036.
  8. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 "GTDB release 10-RS226". Genome Taxonomy Database . Retrieved 1 May 2025.
  9. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 "ar53_r226.sp_label". Genome Taxonomy Database . Retrieved 1 May 2025.
  10. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 "Taxon History". Genome Taxonomy Database . Retrieved 1 May 2025.
  11. "The LTP" . Retrieved 23 February 2021.
  12. "LTP_all tree in newick format" . Retrieved 23 February 2021.
  13. "LTP_12_2021 Release Notes" (PDF). Retrieved 23 February 2021.
  14. 1 2 3 Xie, Ruize; Wang, Yinzhao; Huang, Danyue; Hou, Jialin; Li, Liuyang; Hu, Haining; Zhao, Xiaoxiao; Wang, Fengping (April 2022). "Expanding Asgard members in the domain of Archaea sheds new light on the origin of eukaryotes". Science China Life Sciences. 65 (4): 818–829. doi:10.1007/s11427-021-1969-6. ISSN   1674-7305. PMID   34378142.
  15. "Njordarchaeota, a new candidate for a sister group to eukaryotes". EurekAlert!. Oct 20, 2021.
  16. Order "Candidatus Njordarchaeales" in LPSN ; Parte, Aidan C.; Sardà Carbasse, Joaquim; Meier-Kolthoff, Jan P.; Reimer, Lorenz C.; Göker, Markus (1 November 2020). "List of Prokaryotic names with Standing in Nomenclature (LPSN) moves to the DSMZ". International Journal of Systematic and Evolutionary Microbiology. 70 (11): 5607–5612. doi: 10.1099/ijsem.0.004332 .
  17. 1 2 Wu, Fabai; Speth, Daan R.; Philosof, Alon; Crémière, Antoine; Narayanan, Aditi; Barco, Roman A.; Connon, Stephanie A.; Amend, Jan P.; Antoshechkin, Igor A.; Orphan, Victoria J. (February 2022). "Unique mobile elements and scalable gene flow at the prokaryote-eukaryote boundary revealed by circularized Asgard archaea genomes". Nature Microbiology. 7 (2): 200–212. doi:10.1038/s41564-021-01039-y. ISSN   2058-5276. PMC   8813620 . PMID   35027677.
  18. Order "Candidatus Freyrarchaeales" in LPSN ; Parte, Aidan C.; Sardà Carbasse, Joaquim; Meier-Kolthoff, Jan P.; Reimer, Lorenz C.; Göker, Markus (1 November 2020). "List of Prokaryotic names with Standing in Nomenclature (LPSN) moves to the DSMZ". International Journal of Systematic and Evolutionary Microbiology. 70 (11): 5607–5612. doi: 10.1099/ijsem.0.004332 .
  19. Family "Candidatus Sigynarchaeaceae" in LPSN ; Parte, Aidan C.; Sardà Carbasse, Joaquim; Meier-Kolthoff, Jan P.; Reimer, Lorenz C.; Göker, Markus (1 November 2020). "List of Prokaryotic names with Standing in Nomenclature (LPSN) moves to the DSMZ". International Journal of Systematic and Evolutionary Microbiology. 70 (11): 5607–5612. doi: 10.1099/ijsem.0.004332 .