List of Nobel laureates in Physiology or Medicine

Last updated

Front side (obverse) of the Nobel Prize Medal for Physiology or Medicine NobelPrize1.jpg
Front side (obverse) of the Nobel Prize Medal for Physiology or Medicine

The Nobel Prize in Physiology or Medicine (Swedish : Nobelpriset i fysiologi eller medicin) is awarded annually by the Swedish Karolinska Institute to scientists in the various fields of physiology or medicine. It is one of the five Nobel Prizes established by the 1895 will of Alfred Nobel (who died in 1896), awarded for outstanding contributions in chemistry, physics, literature, peace, and physiology or medicine. [1] As dictated by Nobel's will, the award is administered by the Nobel Foundation and awarded by a committee that consists of five members and an executive secretary elected by the Karolinska Institute. [2] [3] While commonly referred to as the Nobel Prize in Medicine, Nobel specifically stated that the prize be awarded for "physiology or medicine" in his will. Because of this, the prize can be awarded in a broader range of fields. [3] The first Nobel Prize in Physiology or Medicine was awarded in 1901 to Emil Adolf von Behring, of Germany. Each recipient receives a medal, a diploma and a monetary award that has varied throughout the years. [4] In 1901, von Behring received 150,782 SEK, which was equal to 7,731,004 SEK in December 2008. The award is presented in Stockholm at an annual ceremony on 10 December, the anniversary of Nobel's death. [5]

Contents

Laureates have won the Nobel Prize in a wide range of fields that relate to physiology or medicine. As of 2009, 8 Prizes have been awarded for contributions in the field of signal transduction by G proteins and second messengers, 13 have been awarded for contributions in the field of neurobiology and 13 have been awarded for contributions in intermediary metabolism. [3] In 1939 Gerhard Domagk, a German, was not allowed by his government to accept the prize. He later received a medal and diploma, but not the money. [6] As of 2024, the prize has been awarded to 229 individuals, thirteen of them were women (Gerty Cori being the first to be awarded in 1947). [7]

There have been nine years in which the Nobel Prize in Physiology or Medicine was not awarded (1915–1918, 1921, 1925, 1940–1942). There were also five years for which the Nobel Prize in Physiology or Medicine was delayed for one year. The Prize was not awarded in 1914, as the Nobel Committee for Physiology or Medicine decided that none of that year's nominations met the necessary criteria, but was awarded to Robert Bárány in 1915 and counted as the 1914 prize. [8] This precedent was followed for the 1922 prize awarded to Archibald Hill and Otto Fritz Meyerhof in 1923, [9] the 1926 prize awarded to Johannes Fibiger in 1927, [10] the 1938 prize awarded to Corneille Heymans in 1939, [11] and the 1943 prize awarded to Henrik Dam and Edward Adelbert Doisy in 1944. [12]

Laureates

YearImageLaureate [A] Nationality [B] Rationale [C] Ref
1901 EmilVonBehring.jpg Emil von Behring
(1854–1917)
Flag of the German Empire.svg  Germany "for his work on serum therapy, especially its application against diphtheria, by which he has opened a new road in the domain of medical science and thereby placed in the hands of the physician a victorious weapon against illness and deaths" [13]
1902 Ronald Ross.jpg Sir Ronald Ross
(1857–1932)
Flag of the United Kingdom.svg  United Kingdom "for his work on malaria, by which he has shown how it enters the organism and thereby has laid the foundation for successful research on this disease and methods of combating it" [14]
1903 Niels ryberg.jpg Niels Ryberg Finsen
(1860–1904)
Flag of Denmark.svg  Denmark "[for] his contribution to the treatment of diseases, especially lupus vulgaris, with concentrated light radiation, whereby he has opened a new avenue for medical science" [15]
1904 Ivan Pavlov nobel.jpg Ivan Pavlov
(1849–1936)
Flag of Russia.svg  Russia "in recognition of his work on the physiology of digestion, through which knowledge on vital aspects of the subject has been transformed and enlarged" [16]
1905 RobertKoch cropped.jpg Robert Koch
(1843–1910)
Flag of the German Empire.svg  Germany "for his investigations and discoveries in relation to tuberculosis" [17]
1906 Camillo Golgi nobel.jpg Camillo Golgi
(1843–1926)
Flag of Italy (1861-1946) crowned.svg  Italy "in recognition of their work on the structure of the nervous system" [18]
Cajal-Restored.jpg Santiago Ramón y Cajal
(1852–1934)
Flag of Spain (1785-1873 and 1875-1931).svg  Spain
1907 Charles Laveran nobel.jpg Charles Louis Alphonse Laveran
(1845–1922)
Flag of France (1794-1815, 1830-1958).svg  France "in recognition of his work on the role played by protozoa in causing diseases" [19]
1908 Ilya Mechnikov nobel.jpg Élie Metchnikoff
(1845–1916)
Flag of Russia.svg  Russia "in recognition of their work on immunity" [20]
Paul Ehrlich (1926-27 Microbe Hunters).jpg Paul Ehrlich
(1854–1915)
Flag of the German Empire.svg  Germany
1909 Emil Theodor Kocher nobel.jpg Emil Theodor Kocher
(1841–1917)
Flag of Switzerland (Pantone).svg   Switzerland "for his work on the physiology, pathology and surgery of the thyroid gland" [21]
1910 Kossel, Albrecht (1853-1927).jpg Albrecht Kossel
(1853–1927)
Flag of the German Empire.svg  Germany "in recognition of the contributions to our knowledge of cell chemistry made through his work on proteins, including the nucleic substances" [22]
1911 Allvar Gullstrand.jpg Allvar Gullstrand
(1862–1930)
Flag of Sweden.svg  Sweden "for his work on the dioptrics of the eye" [23]
1912 Alexis Carrel 02.jpg Alexis Carrel
(1873–1944)
Flag of France.svg  France "[for] his work on vascular suture and the transplantation of blood vessels and organs" [24]
1913 Charles Robert Richet nobel.jpg Charles Richet
(1850–1935)
Flag of France.svg  France "[for] his work on anaphylaxis" [25]
1914 Robert Barany (3x4 cropped).jpg Robert Bárány
(1876–1936)
Flag of Austria-Hungary (1867-1918).svg  Austria-Hungary "for his work on the physiology and pathology of the vestibular apparatus" [8]
1915
1916
1917
1918
1919 Jules Bordet nobel.jpg Jules Bordet
(1870–1961)
Flag of Belgium (civil).svg  Belgium "for his discoveries relating to immunity" [26]
1920 August Krogh Bain 32006.jpg August Krogh
(1874–1949)
Flag of Denmark.svg  Denmark "for his discovery of the capillary motor regulating mechanism" [27]
1921
1922 Archibald Vivian Hill.jpg Archibald Hill
(1886–1977)
Flag of the United Kingdom.svg  United Kingdom "for his discovery relating to the production of heat in the muscle" [9]
Otto Fritz Meyerhof.jpg Otto Fritz Meyerhof
(1884–1951)
Flag of Germany (3-2 aspect ratio).svg  Germany "for his discovery of the fixed relationship between the consumption of oxygen and the metabolism of lactic acid in the muscle" [9]
1923 Fredrick banting.jpg Sir Frederick Banting
(1891–1941)
Canadian Red Ensign (1921-1957).svg  Canada "for the discovery of insulin" [28]
J.J.R. Macleod ca. 1928.png John Macleod
(1876–1935)
Flag of the United Kingdom.svg  United Kingdom
1924 Willem Einthoven.jpg Willem Einthoven
(1860–1927)
Flag of the Netherlands.svg  Netherlands "for the discovery of the mechanism of the electrocardiogram" [29]
1925
1926 J Fibiger.jpg Johannes Fibiger
(1867–1928)
Flag of Denmark.svg  Denmark "for his discovery of the Spiroptera carcinoma" [10]
1927 Julius Wagner-Jauregg ohne Unterschrift.jpg Julius Wagner-Jauregg
(1857–1940)
Flag of Austria.svg  Austria "for his discovery of the therapeutic value of malaria inoculation in the treatment of dementia paralytica" [30]
1928 Jules Nicole (cropped).jpg Charles Nicolle
(1866–1936)
Flag of France.svg  France "for his work on typhus" [31]
1929 Christiaan Eijkman.jpg Christiaan Eijkman
(1868–1930)
Flag of the Netherlands.svg  Netherlands "for his discovery of the antineuritic vitamin" [32]
Frederick Gowland Hopkins nobel.jpg Sir Frederick Gowland Hopkins
(1861–1947)
Flag of the United Kingdom.svg  United Kingdom "for his discovery of the growth-stimulating vitamins" [32]
1930 Karl Landsteiner nobel.jpg Karl Landsteiner
(1868–1943)
Flag of Austria.svg  Austria
Flag of the United States (1912-1959).svg  United States
"for his discovery of human blood groups" [33]
1931 Otto Warburg.jpg Otto Heinrich Warburg
(1883–1970)
Flag of Germany (3-2 aspect ratio).svg  Germany "for his discovery of the nature and mode of action of the respiratory enzyme" [34]
1932 Prof. Charles Scott Sherrington.jpg Sir Charles Scott Sherrington
(1857–1952)
Flag of the United Kingdom.svg  United Kingdom "for their discoveries regarding the functions of neurons" [35]
Edgar Douglas Adrian nobel.jpg Edgar Adrian
(1889–1977)
1933 Thomas Hunt Morgan.jpg Thomas Hunt Morgan
(1866–1945)
Flag of the United States (1912-1959).svg  United States "for his discoveries concerning the role played by the chromosome in heredity" [36]
1934 George Whipple nobel.jpg George Whipple
(1878–1976)
Flag of the United States (1912-1959).svg  United States "for their discoveries concerning liver therapy in cases of anaemia" [37]
George Minot nobel.jpg George Minot
(1885–1950)
William P Murphy.jpg William P. Murphy
(1892–1987)
1935 Hans Spemann nobel.jpg Hans Spemann
(1869–1941)
Flag of Germany (1935-1945).svg  Germany was awarded a Nobel Prize in Physiology or Medicine in 1935 for his student Hilde Mangold's discovery of the effect now known as embryonic induction, an influence, exercised by various parts of the embryo, that directs the development of groups of cells into particular tissues and organs, the start of artificial cloning of organisms. Spemann added his name as an author to Hilde Mangold's dissertation (although she objected). [38] [39]
1936 Henry Dale nobel.jpg Sir Henry Hallett Dale
(1875–1968)
Flag of the United Kingdom.svg  United Kingdom "for their discoveries relating to chemical transmission of nerve impulses" [40]
Otto Loewi nobel.jpg Otto Loewi
(1873–1961)
Flag of Germany (1935-1945).svg  Germany
1937 Albert Szent-Gyorgyi cropped.jpg Albert Szent-Györgyi
(1893–1986)
Flag of Hungary (1918-1919).svg  Hungary "for his discoveries in connection with the biological combustion processes, with special reference to vitamin C and the catalysis of fumaric acid" [41]
1938 Corneille Heymans nobel.jpg Corneille Heymans
(1892–1968)
Flag of Belgium (civil).svg  Belgium "for the discovery of the role played by the sinus and aortic mechanisms in the regulation of respiration" [11]
1939 Gerhard Domagk nobel.jpg Gerhard Domagk
(1895–1964)
Flag of Germany (1935-1945).svg  Nazi Germany "for the discovery of the antibacterial effects of prontosil" [42]
1940
1941
1942
1943 Henrik Dam nobel.jpg Henrik Dam
(1895–1976)
Flag of Denmark.svg  Denmark "for his discovery of vitamin K" [12]
Edward A Doisy.jpg Edward Adelbert Doisy
(1893–1986)
Flag of the United States (1912-1959).svg  United States "for his discovery of the chemical nature of vitamin K" [12]
1944 Joseph Erlanger nobel.jpg Joseph Erlanger
(1874–1965)
Flag of the United States (1912-1959).svg  United States "for their discoveries relating to the highly differentiated functions of single nerve fibres" [43]
Herbert Spencer Gasser nobel.jpg Herbert Spencer Gasser
(1888–1963)
1945 Alexander Fleming 1945.jpg Sir Alexander Fleming
(1881–1955)
Flag of the United Kingdom.svg  United Kingdom "for the discovery of penicillin and its curative effect in various infectious diseases" [44]
Ernst Boris Chain 1945.jpg Sir Ernst Boris Chain
(1906–1979)
Howard Walter Florey 1945.jpg Howard Walter Florey
(1898–1968)
Flag of Australia (converted).svg  Australia
1946 HJ Muller 1952.jpg Hermann Joseph Muller
(1890–1967)
Flag of the United States (1912-1959).svg  United States "for the discovery of the production of mutations by means of X-ray irradiation" [45]
1947 Carl Ferdinand Cori.jpg Carl Ferdinand Cori
(1896–1984)
Flag of the Czech Republic.svg  Czechoslovakia
Flag of the United States (1912-1959).svg  United States
"for their discovery of the course of the catalytic conversion of glycogen" [46]
Gerty Theresa Cori.jpg Gerty Theresa Cori, née Radnitz
(1896–1957)
Bernado Houssay.JPG Bernardo Alberto Houssay
(1887–1971)
Flag of Argentina.svg  Argentina "for his discovery of the part played by the hormone of the anterior pituitary lobe in the metabolism of sugar" [46]
1948 Paul Hermann Muller nobel.jpg Paul Hermann Müller
(1899–1965)
Flag of Switzerland (Pantone).svg   Switzerland "for his discovery of the high efficiency of DDT as a contact poison against several arthropods" [47]
1949 Walter Hess.jpg Walter Rudolf Hess
(1881–1973)
Flag of Switzerland (Pantone).svg   Switzerland "for his discovery of the functional organization of the interbrain as a coordinator of the activities of the internal organs" [48]
Moniz.jpg António Caetano Egas Moniz
(1874–1955)
Flag of Portugal.svg  Portugal "for his discovery of the therapeutic value of leucotomy (lobotomy) in certain psychoses" [48]
1950 Philip Showalter Hench.jpg Philip Showalter Hench
(1896–1965)
Flag of the United States (1912-1959).svg  United States "for their discoveries relating to the hormones of the adrenal cortex, their structure and biological effects" [49]
Edward Calvin Kendall nobel.jpg Edward Calvin Kendall
(1886–1972)
Thadeus Reichstein ETH-Bib Portr 10137.jpg Tadeusz Reichstein
(1897–1996)
Flag of Poland (1928-1980).svg  Poland
Flag of Switzerland (Pantone).svg   Switzerland
1951 Max Theiler nobel.jpg Max Theiler
(1899–1972)
Flag of South Africa (1928-1982).svg  South Africa
Flag of the United States (1912-1959).svg  United States
"for his discoveries concerning yellow fever and how to combat it" [50]
1952 Selman Waksman NYWTScrop.jpg Selman Abraham Waksman
(1888–1973)
Flag of the United States (1912-1959).svg  United States "for his discovery of streptomycin, the first antibiotic effective against tuberculosis" [51]
1953 Hans Adolf Krebs.jpg Sir Hans Adolf Krebs
(1900–1981)
Flag of Germany.svg  West Germany
Flag of the United Kingdom.svg  United Kingdom
"for his discovery of the citric acid cycle" [52]
Portrait of Fritz Albert Lipmann (1899-1986), Biochemist (2551001689).jpg Fritz Albert Lipmann
(1899–1986)
Flag of the United States (1912-1959).svg  United States "for his discovery of co-enzyme A and its importance for intermediary metabolism" [52]
1954 John Franklin Enders
(1897–1985)
Flag of the United States (1912-1959).svg  United States "for their discovery of the ability of poliomyelitis viruses to grow in cultures of various types of tissue" [53]
Frederick Chapman Robbins nobel.jpg Frederick Chapman Robbins
(1916–2003)
Thomas Huckle Weller.jpg Thomas Huckle Weller
(1915–2008)
1955 Hugo Theorell nobel.jpg Axel Hugo Theodor Theorell
(1903–1982)
Flag of Sweden.svg  Sweden "for his discoveries concerning the nature and mode of action of oxidation enzymes" [54]
1956 Andre Frederic Cournand nobel.jpg André Frédéric Cournand
(1895–1988)
Flag of France.svg  France
Flag of the United States (1912-1959).svg  United States
"for their discoveries concerning heart catheterization and pathological changes in the circulatory system" [55]
Werner Forssmann nobel.jpg Werner Forssmann
(1904–1979)
Flag of Germany.svg  West Germany
Dickinson W. Richards nobel.jpg Dickinson W. Richards
(1895–1973)
Flag of the United States (1912-1959).svg  United States
1957 Daniel Bovet nobel.jpg Daniel Bovet
(1907–1992)
Flag of Switzerland (Pantone).svg   Switzerland
Flag of Italy.svg  Italy
"for his discoveries relating to synthetic compounds that inhibit the action of certain body substances, and especially their action on the vascular system and the skeletal muscles" [56]
1958 George Wells Beadle.jpg George Wells Beadle
(1903–1989)
Flag of the United States (1912-1959).svg  United States "for their discovery that genes act by regulating definite chemical events" [57]
Edward Lawrie Tatum nobel.jpg Edward Lawrie Tatum
(1909–1975)
Joshua Lederberg crop.jpg Joshua Lederberg
(1925–2008)
"for his discoveries concerning genetic recombination and the organization of the genetic material of bacteria" [57]
1959 Arthur Kornberg (cropped).jpg Arthur Kornberg
(1918–2007)
Flag of the United States (1959-1960).svg  United States "for their discovery of the mechanisms in the biological synthesis of ribonucleic acid and deoxyribonucleic acid" [58]
Severo Ochoa nobel.jpg Severo Ochoa
(1905–1993)
Flag of Spain (1945-1977).svg  Spain
Flag of the United States (1959-1960).svg  United States
1960 Frank Macfarlane Burnet nobel.jpg Sir Frank Macfarlane Burnet
(1899–1985)
Flag of Australia (converted).svg  Australia "for discovery of acquired immunological tolerance" [59]
Peter Brian Medawar.jpg Sir Peter Brian Medawar
(1915–1987)
Flag of the United Kingdom.svg  United Kingdom
1961 Georg von Bekesy nobel.jpg Georg von Békésy
(1899–1972)
Flag of Hungary.svg  Hungary
Flag of the United States.svg  United States
"for his discoveries of the physical mechanism of stimulation within the cochlea" [60]
1962 Francis Crick (3x4 cropped).jpg Francis Crick
(1916–2004)
Flag of the United Kingdom.svg  United Kingdom "for their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material" [61] [62]
James Dewey Watson (cropped).jpg James Dewey Watson
(b. 1928)
Flag of the United States.svg  United States
Maurice Wilkins nobel.jpg Maurice Hugh Frederick Wilkins
(1916–2004)
Flag of New Zealand.svg  New Zealand
Flag of the United Kingdom.svg  United Kingdom
1963 Sir John Eccles Wellcome L0026812.jpg Sir John Carew Eccles
(1903–1997)
Flag of Australia (converted).svg  Australia "for their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane" [63]
Alan Lloyd Hodgkin nobel.jpg Sir Alan Lloyd Hodgkin
(1914–1998)
Flag of the United Kingdom.svg  United Kingdom
Andrew Fielding Huxley nobel.jpg Sir Andrew Fielding Huxley
(1917–2012)
1964 Konrad Bloch (head shoulders 3x4 cropped) (cropped).JPG Konrad Bloch
(1912–2000)
Flag of the United States.svg  United States "for their discoveries concerning the mechanism and regulation of the cholesterol and fatty acid metabolism" [64]
Feodor Lynen with pipette.jpg Feodor Lynen
(1911–1979)
Flag of Germany.svg  West Germany
1965 Francois Jacob nobel.jpg François Jacob
(1920–2013)
Flag of France.svg  France "for their discoveries concerning genetic control of enzyme and virus synthesis" [65]
Andre Lwoff nobel.jpg André Lwoff
(1902–1994)
Jacques Monod nobel (cropped).jpg Jacques Monod
(1910–1976)
1966 Peyton Rous nobel.jpg Peyton Rous
(1879–1970)
Flag of the United States.svg  United States "for his discovery of tumour-inducing viruses" [66]
Charles Brenton Huggins nobel.jpg Charles Brenton Huggins
(1901–1997)
Flag of Canada (Pantone).svg  Canada
Flag of the United States.svg  United States
"for his discoveries concerning hormonal treatment of prostatic cancer" [66]
1967 Ragnar Granit2.jpg Ragnar Granit
(1900–1991)
Flag of Finland.svg  Finland
Flag of Sweden.svg  Sweden
"for their discoveries concerning the primary physiological and chemical visual processes in the eye" [67]
Haldan Keffer Hartline nobel.jpg Haldan Keffer Hartline
(1903–1983)
Flag of the United States.svg  United States
George Wald nobel.jpg George Wald
(1906–1997)
1968 Robert W. Holley nobel.jpg Robert W. Holley
(1922–1993)
Flag of the United States.svg  United States "for their interpretation of the genetic code and its function in protein synthesis" [68]
Har Gobind Khorana nobel.jpg Har Gobind Khorana
(1922–2011)
Flag of India.svg  India
Flag of the United States.svg  United States
Marshall Nirenberg 2003.jpg Marshall W. Nirenberg
(1927–2010)
Flag of the United States.svg  United States
1969 Max Delbruck (cropped).jpg Max Delbrück
(1906–1981)
Flag of the United States.svg  United States "for their discoveries concerning the replication mechanism and the genetic structure of viruses" [69]
Alfred Hershey (cropped).jpg Alfred Hershey
(1908–1997)
Salvador E. Luria ca.1969 (cropped).jpg Salvador E. Luria
(1912–1991)
Flag of Italy.svg  Italy
Flag of the United States.svg  United States
1970 Axelrod01.jpg Julius Axelrod
(1912–2004)
Flag of the United States.svg  United States "for their discoveries concerning the humoral transmitters in the nerve terminals and the mechanism for their storage, release and inactivation" [70]
Ulf von Euler.jpg Ulf von Euler
(1905–1983)
Flag of Sweden.svg  Sweden
Sir Bernard Katz.jpg Sir Bernard Katz
(1911–2003)
Flag of Germany.svg  West Germany
Flag of the United Kingdom.svg  United Kingdom
1971 Earl Wilbur Sutherland Jr.jpg Earl W. Sutherland Jr.
(1915–1974)
Flag of the United States.svg  United States "for his discoveries concerning the mechanisms of the action of hormones" [71]
1972 Professor Gerald M. Edelman (cropped).jpg Gerald M. Edelman
(1929–2014)
Flag of the United States.svg  United States "for their discoveries concerning the chemical structure of antibodies" [72]
Rodney Robert Porter.jpg Rodney R. Porter
(1917–1985)
Flag of the United Kingdom.svg  United Kingdom
1973 Karl von Frisch - Atelier Veritas, c. 1926.jpg Karl von Frisch
(1886–1982)
Flag of Austria.svg  Austria "for their discoveries concerning organization and elicitation of individual and social behaviour patterns" [73]
Konrad Lorenz (3x4 cropped).JPG Konrad Lorenz
(1903–1989)
Nikolaas Tinbergen 1978.jpg Nikolaas Tinbergen
(1907–1988)
Flag of the Netherlands.svg  Netherlands
1974 Albert Claude 1974.jpg Albert Claude
(1899–1983)
Flag of Belgium (civil).svg  Belgium
Flag of the United States.svg  United States
"for their discoveries concerning the structural and functional organization of the cell" [74]
Christian de Duve cropped (3x4 cropped).jpg Christian de Duve
(1917–2013)
Flag of Belgium (civil).svg  Belgium
George Emil Palade 2021 stamp of Romania (cropped).jpg George E. Palade
(1912–2008)
Flag of Romania (1965-1989).svg  Romania
Flag of the United States.svg  United States
1975 Dr. David Baltimore2.jpg David Baltimore
(b. 1938)
Flag of the United States.svg  United States "for their discoveries concerning the interaction between tumour viruses and the genetic material of the cell" [75]
Renato Dulbecco crop.jpg Renato Dulbecco
(1914–2012)
Flag of Italy.svg  Italy
Flag of the United States.svg  United States
Howard Martin Temin 1975 (cropped).jpg Howard Martin Temin
(1934–1994)
Flag of the United States.svg  United States
1976 Baruch Samuel Blumberg by Tom Trower (NASA) (3x4 cropped).jpg Baruch Samuel Blumberg
(1925–2011)
Flag of the United States.svg  United States "for their discoveries concerning new mechanisms for the origin and dissemination of infectious diseases" [76]
Daniel Carleton Gajdusek 2bw cropped.jpg D. Carleton Gajdusek
(1923–2008)
1977 Roger Guillemin.jpg Roger Guillemin
(1924–2024)
Flag of France (lighter variant).svg  France
Flag of the United States.svg  United States
"for their discoveries concerning the peptide hormone production of the brain" [77]
Schally portrait.jpg Andrew V. Schally
(1926–2024)
Flag of the United States.svg  United States
Rosalyn Yalow (3x4 cropped).jpg Rosalyn Yalow
(1921–2011)
"for the development of radioimmunoassays of peptide hormones" [77]
1978 Werner Arber 2008.jpg Werner Arber
(b. 1929)
Flag of Switzerland (Pantone).svg   Switzerland "for the discovery of restriction enzymes and their application to problems of molecular genetics" [78]
37. Tagung 1987 Mediziner Daniel Nathans lachelnd - W134Nr.125411c - Willy Pragher (cropped).jpg Daniel Nathans
(1928–1999)
Flag of the United States.svg  United States
Hamilton Smith (3x4 cropped).jpg Hamilton O. Smith
(b. 1931)
1979 A. M. Cormack.jpg Allan M. Cormack
(1924–1998)
Flag of South Africa (1928-1982).svg  South Africa
Flag of the United States.svg  United States
"for the development of computer assisted tomography" [79]
  Godfrey Hounsfield.jpg Sir Godfrey N. Hounsfield
(1919–2004)
Flag of the United Kingdom.svg  United Kingdom
1980 Baruj Benacerraf 1969.jpg Baruj Benacerraf
(1920–2011)
Flag of Venezuela (1954-2006).svg  Venezuela
Flag of the United States.svg  United States
"for their discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions" [80]
Jean Dausset 1968.jpg Jean Dausset
(1916–2009)
Flag of France (lighter variant).svg  France
No image.svg George D. Snell
(1903–1996)
Flag of the United States.svg  United States
1981 Roger Wolcott Sperry.jpg Roger W. Sperry
(1913–1994)
Flag of the United States.svg  United States "for his discoveries concerning the functional specialization of the cerebral hemispheres" [81]
DHUBEL.jpg David H. Hubel
(1926–2013)
"for their discoveries concerning information processing in the visual system" [81]
Torsten Wiesel-7Nov2006.jpg Torsten N. Wiesel
(b. 1924)
Flag of Sweden.svg  Sweden
1982 Sune Bergstrom 3.jpg Sune K. Bergström
(1916–2004)
Flag of Sweden.svg  Sweden "for their discoveries concerning prostaglandins and related biologically active substances" [82]
Bengt Samuelsson.png Bengt I. Samuelsson
(1934–2024)
John Robert Vane.jpg Sir John R. Vane
(1927–2004)
Flag of the United Kingdom.svg  United Kingdom
1983 Barbara McClintock (1902-1992) shown in her laboratory in 1947 (cropped).jpg Barbara McClintock
(1902–1992)
Flag of the United States.svg  United States "for her discovery of mobile genetic elements" [83]
1984 Niels Kaj Jerne 1950 crop.jpg Niels K. Jerne
(1911–1994)
Flag of Denmark.svg  Denmark "for theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies" [84]
Einweihung des Max-Planck-Institutes fur Immunbiologie (1990) (cropped).jpg Georges J. F. Köhler
(1946–1995)
Flag of Germany.svg  West Germany
Milstein lnp restauracion (3x4 cropped).jpg César Milstein
(1927–2002)
Flag of Argentina.svg  Argentina
1985 Mike Brown 2003.jpg Michael S. Brown
(b. 1941)
Flag of the United States.svg  United States "for their discoveries concerning the regulation of cholesterol metabolism" [85]
Joe Goldstein.JPG Joseph L. Goldstein
(b. 1940)
1986 Stanley Cohen-Biochemist.jpg Stanley Cohen
(1922–2020)
Flag of the United States.svg  United States "for their discoveries of growth factors" [86]
Rita Levi Montalcini.jpg Rita Levi-Montalcini
(1909–2012)
Flag of Italy.svg  Italy
1987 Susumu Tonegawa Photo cropped.jpg Susumu Tonegawa
(b. 1939)
Flag of Japan (1870-1999).svg  Japan "for his discovery of the genetic principle for generation of antibody diversity" [87]
1988 James Black (pharmacologist) (3x4 cropped).jpg Sir James W. Black
(1924–2010)
Flag of the United Kingdom.svg  United Kingdom "for their discoveries of important principles for drug treatment" [88]
Gertrude Elion.jpg Gertrude B. Elion
(1918–1999)
Flag of the United States.svg  United States
George H. Hitchings 1988.jpg George H. Hitchings
(1905–1998)
1989 Nci-vol-8172-300 j michael bishop.jpg J. Michael Bishop
(b. 1936)
Flag of the United States.svg  United States "for their discovery of the cellular origin of retroviral oncogenes" [89]
HEVarmus.jpg Harold E. Varmus
(b. 1939)
1990 Joseph E. Murray
(1919–2012)
Flag of the United States.svg  United States "for their discoveries concerning organ and cell transplantation in the treatment of human disease" [90]
Edward Donnall "Don" Thomas.jpg E. Donnall Thomas
(1920–2012)
1991 Erwin neher 2007 lindau.jpg Erwin Neher
(b. 1944)
Flag of Germany.svg  Germany "for their discoveries concerning the function of single ion channels in cells" [91]
Sakmann.jpg Bert Sakmann
(b. 1942)
1992 Edmond Fischer Nobel Laureate 1992.jpg Edmond H. Fischer
(1920–2021)
Flag of Switzerland (Pantone).svg   Switzerland
Flag of the United States.svg  United States
"for their discoveries concerning reversible protein phosphorylation as a biological regulatory mechanism" [92]
Edwin G. Krebs
(1918–2009)
Flag of the United States.svg  United States
1993 Roberts, Richard John (1943).jpg Sir Richard J. Roberts
(b. 1943)
Flag of the United Kingdom.svg  United Kingdom "for their discoveries of split genes" [93]
Phillip A Sharp NIH.jpg Phillip A. Sharp
(b. 1944)
Flag of the United States.svg  United States
1994 Alfred Goodman Gilman (cropped).jpg Alfred G. Gilman
(1941–2015)
Flag of the United States.svg  United States "for their discovery of G-proteins and the role of these proteins in signal transduction in cells" [94]
Rodbell, Martin (1925-1998).jpg Martin Rodbell
(1925–1998)
1995 Ed Lewis 1986.png Edward B. Lewis
(1918–2004)
Flag of the United States.svg  United States "for their discoveries concerning the genetic control of early embryonic development" [95]
Christiane Nusslein-Volhard mg 4383 (cropped).jpg Christiane Nüsslein-Volhard
(b. 1942)
Flag of Germany.svg  Germany
Eric F. Wieschaus.jpg Eric F. Wieschaus
(b. 1947)
Flag of the United States.svg  United States
1996 Peter C. Doherty 2017.jpg Peter C. Doherty
(b. 1940)
Flag of Australia (converted).svg  Australia "for their discoveries concerning the specificity of the cell mediated immune defence" [96]
Rolf Zinkernagel Erudite Conclave medical college trivandrum (3x4 cropped).jpg Rolf M. Zinkernagel
(b. 1944)
Flag of Switzerland (Pantone).svg   Switzerland
1997 Prusiner 1.JPG Stanley B. Prusiner
(b. 1942)
Flag of the United States.svg  United States "for his discovery of Prions - a new biological principle of infection" [97]
1998 Drfurchgott.jpg Robert F. Furchgott
(1916–2009)
Flag of the United States.svg  United States "for their discoveries concerning nitric oxide as a signalling molecule in the cardiovascular system" [98]
Loius Ignarro.jpg Louis J. Ignarro
(b. 1941)
Ferid Murad.jpg Ferid Murad
(1936–2023)
1999 Gunter Blobel 2008 1.JPG Günter Blobel
(1936–2018)
Flag of the United States.svg  United States "for the discovery that proteins have intrinsic signals that govern their transport and localization in the cell" [99]
2000 Arvid Carlsson 2011a (3x4 cropped).jpg Arvid Carlsson
(1923–2018)
Flag of Sweden.svg  Sweden "for their discoveries concerning signal transduction in the nervous system" [100]
Paul Greengard.jpg Paul Greengard
(1925–2019)
Flag of the United States.svg  United States
Eric Kandel 01 (3x4 cropped).JPG Eric Kandel
(b. 1929)
Flag of Austria.svg  Austria
Flag of the United States.svg  United States
2001 No image.svg Leland H. Hartwell
(b. 1939)
Flag of the United States.svg  United States "for their discoveries of key regulators of the cell cycle" [101]
Tim hunt.jpg Sir Tim Hunt
(b. 1943)
Flag of the United Kingdom.svg  United Kingdom
Paul Nurse 2007.jpg Sir Paul M. Nurse
(b. 1949)
2002 Sydney Brenner OIST 2008 (33208371153) (cropped).jpg Sydney Brenner
(1927–2019)
Flag of South Africa.svg  South Africa "for their discoveries concerning 'genetic regulation of organ development and programmed cell death'" [102]
H. Robert Horvitz
(b. 1947)
Flag of the United States.svg  United States
Sulston, John Edward (1942).jpg Sir John E. Sulston
(1942–2018)
Flag of the United Kingdom.svg  United Kingdom
2003 Paul Lauterbur 2003 cropped.jpg Paul Lauterbur
(1929–2007)
Flag of the United States.svg  United States "for their discoveries concerning magnetic resonance imaging" [103]
Peter Mansfield Leipzig.jpg Sir Peter Mansfield
(1933–2017)
Flag of the United Kingdom.svg  United Kingdom
2004 Richard Axel (cropped).jpg Richard Axel
(b. 1946)
Flag of the United States.svg  United States "for their discoveries of odorant receptors and the organization of the olfactory system" [104]
Linda Buck cropped 1 (3x4 cropped).jpg Linda B. Buck
(b. 1947)
2005 Marshall 2008.JPG Barry J. Marshall
(b. 1951)
Flag of Australia (converted).svg  Australia "for their discovery of the bacterium Helicobacter pylori and its role in gastritis and peptic ulcer disease" [105]
Robin Warren (3x4 cropped).jpg J. Robin Warren
(1937–2024)
2006 Andrew Fire, Stanford University.jpg Andrew Z. Fire
(b. 1959)
Flag of the United States.svg  United States "for their discovery of RNA interference - gene silencing by double-stranded RNA" [106]
Craig Mello 01 (3x4 cropped).jpg Craig C. Mello
(b. 1960)
2007 MarioCapecchiFotoThalerTamas2.JPG Mario R. Capecchi
(b. 1937)
Flag of Italy.svg  Italy
Flag of the United States.svg  United States
"for their discoveries of principles for introducing specific gene modifications in mice by the use of embryonic stem cells." [107]
Martin Evans Nobel Prize (3x4 cropped).jpg Sir Martin Evans
(b. 1941)
Flag of the United Kingdom.svg  United Kingdom
Oliver Smithies HD2009 AIC Gold Medal portrait (3x4cropped).JPG Oliver Smithies
(1925–2017)
Flag of the United Kingdom.svg  United Kingdom
Flag of the United States.svg  United States
2008 Harald zur Hausen-press conference Dec 06th, 2008-6.jpg Harald zur Hausen
(1936–2023)
Flag of Germany.svg  Germany "for his discovery of human papilloma viruses causing cervical cancer" [108]
Francoise Barre-Sinoussi-press conference Dec 06th, 2008-1.jpg Françoise Barré-Sinoussi
(b. 1947)
Flag of France (lighter variant).svg  France "for their discovery of human immunodeficiency virus" [108]
LucMontagnier1995 065.jpg Luc Montagnier
(1932–2022)
2009 Elizabeth Blackburn 2009-01.JPG Elizabeth H. Blackburn
(b. 1948)
Flag of Australia (converted).svg  Australia
Flag of the United States.svg  United States
"for the discovery of how chromosomes are protected by telomeres and the enzyme telomerase" [109]
GREIDER Carol 2014 - Less vignetting.jpg Carol W. Greider
(b. 1961)
Flag of the United States.svg  United States
JSzostak.jpg Jack W. Szostak
(b. 1952)
Flag of Canada (Pantone).svg  Canada
Flag of the United States.svg  United States
2010 Robert Geoffrey Edwards (2010).jpg Sir Robert G. Edwards
(1925–2013)
Flag of the United Kingdom.svg  United Kingdom "for the development of in vitro fertilization" [110]
2011 Nobel Prize 2011-Press Conference (3x4 cropped).jpg Bruce A. Beutler
(b. 1957)
Flag of the United States.svg  United States "for their discoveries concerning the activation of innate immunity" [111]
Nobel Prize 2011-Press Conference KI-DSC 7584.jpg Jules A. Hoffmann
(b. 1941)
Flag of France (lighter variant).svg  France
RMSt.jpg Ralph M. Steinman
(1943–2011)
Flag of Canada (Pantone).svg  Canada "for his discovery of the dendritic cell and its role in adaptive immunity" (awarded posthumously) [112] [113] [111]
2012 John Gurdon Cambridge 2012.JPG Sir John B. Gurdon
(b. 1933)
Flag of the United Kingdom.svg  United Kingdom "for the discovery that mature cells can be reprogrammed to become pluripotent" [114]
Shinya Yamanaka 3x4.jpg Shinya Yamanaka
(b. 1962)
Flag of Japan.svg  Japan
2013 James Edward Rothman.jpg James E. Rothman
(b. 1950)
Flag of the United States.svg  United States "for their discoveries of machinery regulating vesicle traffic, a major transport system in our cells" [115]
Randy Schekman 8 February 2012.jpg Randy W. Schekman
(b. 1948)
Thomas C Sudhof 2013 (3x4 cropped).jpg Thomas C. Südhof
(b. 1955)
Flag of Germany.svg  Germany
Flag of the United States.svg  United States
2014 John O'Keefe (neuroscientist) 2014b.jpg John O'Keefe
(b. 1939)
Flag of the United States.svg  United States
Flag of the United Kingdom.svg  United Kingdom
"for their discoveries of cells that constitute a positioning system in the brain" [116]
May-Britt Moser 2014.jpg May-Britt Moser
(b. 1963)
Flag of Norway.svg  Norway
Edvard Moser (3x4 cropped).jpg Edvard I. Moser
(b. 1962)
2015 William C. Campbell 4983-1-2015.jpg William C. Campbell
(b. 1930)
Flag of Ireland.svg  Ireland
Flag of the United States.svg  United States
"for their discoveries concerning a novel therapy against infections caused by roundworm parasites" [117]
Satoshi Omura 5086-1-2015.jpg Satoshi Ōmura
(b. 1935)
Flag of Japan.svg  Japan
Tu Youyou 5012-1-2015.jpg Tu Youyou
(b. 1930)
Flag of the People's Republic of China.svg  China "for her discoveries concerning a novel therapy against malaria" [117]
2016 Nobel Laureates 1042 (30647248184).jpg Yoshinori Ohsumi
(b. 1945)
Flag of Japan.svg  Japan "for his discoveries of mechanisms for autophagy" [118]
2017 Jeffrey C. Hall EM1B8737 (38162359274).jpg Jeffrey C. Hall
(b. 1945)
Flag of the United States.svg  United States "for their discoveries of molecular mechanisms controlling the circadian rhythm" [119]
Michael Rosbash EM1B8756 (38847326642).jpg Michael Rosbash
(b. 1944)
Michael W. Young D81 4345 (38162439194).jpg Michael W. Young
(b. 1949)
2018 James P. Allison (2015).JPG James P. Allison
(b. 1948)
Flag of the United States.svg  United States "for their discovery of cancer therapy by inhibition of negative immune regulation" [120]
Tasuku Honjo 201311.jpg Tasuku Honjo
(b. 1942)
Flag of Japan.svg  Japan
2019 William G. Kaelin Jr. UNIST CGI 2019 (cropped).jpg William Kaelin Jr.
(b. 1957)
Flag of the United States.svg  United States "for their discoveries of how cells sense and adapt to oxygen availability" [121]
Peter J. Ratcliffe (cropped).jpg Peter J. Ratcliffe
(b. 1954)
Flag of the United Kingdom.svg  United Kingdom
Gregg L. Semenza (cropped).jpg Gregg L. Semenza
(b. 1956)
Flag of the United States.svg  United States
2020 Harvey J Alter 2020 (3x4 cropped).jpg Harvey J. Alter
(b. 1935)
Flag of the United States.svg  United States "for the discovery of Hepatitis C virus" [122]
Prof Michael Houghton.jpg Michael Houghton
(b. 1949)
Flag of the United Kingdom.svg  United Kingdom
Charles M. Rice (3x4 close cropped).jpg Charles M. Rice
(b. 1952)
Flag of the United States.svg  United States
2021 Dr David Julius by christopher michel in 2022 (3x4 cropped).jpg David Julius
(b. 1955)
Flag of the United States.svg  United States "for the discovery of receptors for temperature and touch" [123]
Ardem Patapoutian by C Michel 67.jpg Ardem Patapoutian
(b. 1967)
 Flag of Lebanon.svg  Lebanon
Flag of the United States.svg  United States
2022 Professor Svante Paabo ForMemRS (cropped).jpg Svante Pääbo
(b. 1955)
Flag of Sweden.svg  Sweden "for his discoveries concerning the genomes of extinct hominins and human evolution" [124]
2023 Katalin Kariko by Christopher Michel in 2024 (3x4 close cropped).jpg Katalin Karikó
(b. 1955)
Flag of Hungary.svg  Hungary
Flag of the United States.svg  United States
"for their discoveries concerning nucleoside base modifications that enabled the development of effective mRNA vaccines against COVID-19" [125]
Drew Weissman Life Science medalist (close cropped).jpg Drew Weissman
(b. 1959)
Flag of the United States.svg  United States
2024 Victor Ambros 3x4.jpg Victor Ambros
(b. 1953)
Flag of the United States.svg  United States "for the discovery of microRNA and its role in post-transcriptional gene regulation" [126]
Gruber Prize Laureate Gary Ruvkun (cropped).jpg Gary Ruvkun
(b. 1952)

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<span class="mw-page-title-main">Nobel Prize</span> Prizes established by Alfred Nobel in 1895

The Nobel Prizes are five separate prizes awarded to those who, during the preceding year, have conferred the greatest benefit to humankind, as established by the 1895 will of Swedish chemist, engineer, and industrialist Alfred Nobel, in the year before he died. Prizes were first awarded in 1901 by the Nobel Foundation. Nobel's will indicated that the awards should be granted in the fields of Physics, Chemistry, Physiology or Medicine, Literature, and Peace. A sixth prize for Economic Sciences, endowed by Sweden's central bank, Sveriges Riksbank, and first presented in 1969, is also frequently included, as it is also administered by the Nobel Foundation. The Nobel Prizes are widely regarded as the most prestigious awards available in their respective fields.

<span class="mw-page-title-main">Nobel Prize in Physics</span> One of the five Nobel Prizes established in 1895 by Alfred Nobel

The Nobel Prize in Physics is an annual award given by the Royal Swedish Academy of Sciences for those who have made the most outstanding contributions to mankind in the field of physics. It is one of the five Nobel Prizes established by the will of Alfred Nobel in 1895 and awarded since 1901, the others being the Nobel Prize in Chemistry, Nobel Prize in Literature, Nobel Peace Prize, and Nobel Prize in Physiology or Medicine. Physics is traditionally the first award presented in the Nobel Prize ceremony.

<span class="mw-page-title-main">Nobel Prize in Physiology or Medicine</span> One of five Nobel Prizes established in 1895 by Alfred Nobel

The Nobel Prize in Physiology or Medicine is awarded yearly by the Nobel Assembly at the Karolinska Institute for outstanding discoveries in physiology or medicine. The Nobel Prize is not a single prize, but five separate prizes that, according to Alfred Nobel's 1895 will, are awarded "to those who, during the preceding year, have conferred the greatest benefit to humankind". Nobel Prizes are awarded in the fields of Physics, Medicine or Physiology, Chemistry, Literature, and Peace.

<span class="mw-page-title-main">Nobel Foundation</span> Private institution managing the finances and administration of the Nobel Prizes

The Nobel Foundation is a private institution founded on 29 June 1900 to manage the finances and administration of the Nobel Prizes. The foundation is based on the last will of Alfred Nobel, the inventor of dynamite.

<span class="mw-page-title-main">Nobel Memorial Prize in Economic Sciences</span> Award established in 1968 by Sveriges riksbank in memory of Alfred Nobel

The Nobel Memorial Prize in Economic Sciences, officially the Sveriges Riksbank Prize in Economic Sciences in Memory of Alfred Nobel, is an economics award funded by Sveriges Riksbank and administered by the Nobel Foundation.

<span class="mw-page-title-main">Nobel Prize in Chemistry</span> One of the five Nobel Prizes established in 1895 by Alfred Nobel

The Nobel Prize in Chemistry is awarded annually by the Royal Swedish Academy of Sciences to scientists in the various fields of chemistry. It is one of the five Nobel Prizes established by the will of Alfred Nobel in 1895, awarded for outstanding contributions in chemistry, physics, literature, peace, and physiology or medicine. This award is administered by the Nobel Foundation, and awarded by the Royal Swedish Academy of Sciences on proposal of the Nobel Committee for Chemistry which consists of five members elected by the Academy. The award is presented in Stockholm at an annual ceremony on 10 December, the anniversary of Nobel's death.

References

Notes

^ A. The form and spelling of the names in the name column is according to nobelprize.org, the official website of the Nobel Foundation. Alternative spellings and name forms, where they exist, are given at the articles linked from this column. Where available, an image of each Nobel laureate is provided. For the official pictures provided by the Nobel Foundation, see the pages for each Nobel laureate at nobelprize.org.

^ B. The information in the country column is according to nobelprize.org, the official website of the Nobel Foundation. This information may not necessarily reflect the recipient's birthplace or citizenship, and instead reflect their academic or scientific affiliation at the time of receiving the award.

^ C. The citation for each award is quoted (not always in full) from nobelprize.org, the official website of the Nobel Foundation. The links in this column are to articles (or sections of articles) on the history and  areas of physiology and medicine for which the awards were presented. The links are intended only as a guide and explanation. For a full account of the work done by each Nobel laureate, please see the biography articles linked from the name column.

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