Snow leopard

Last updated

Snow leopard
Irbis4.JPG
CITES Appendix I (CITES) [1]
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Pantherinae
Genus: Panthera
Species:
P. uncia
Binomial name
Panthera uncia
(Schreber, 1775)
SnowLeopard distribution.jpg
Distribution of the snow leopard, 2017 [1]
Synonyms

The snow leopard (Panthera uncia), also known as the ounce, is a felid in the genus Panthera native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is estimated to number fewer than 10,000 mature individuals and is expected to decline about 10% by 2040. It is threatened by poaching and habitat destruction following infrastructural developments. It inhabits alpine and subalpine zones at elevations of 3,000–4,500 m (9,800–14,800 ft), ranging from eastern Afghanistan, the Himalayas and the Tibetan Plateau to southern Siberia, Mongolia and western China. In the northern part of its range, it also lives at lower elevations.

Contents

Taxonomically, the snow leopard was long classified in the monotypic genus Uncia. Since phylogenetic studies revealed the relationships among Panthera species, it has been considered a member of that genus. Two subspecies were described based on morphological differences, but genetic differences between the two have not been confirmed. It is therefore regarded as a monotypic species.

Naming and etymology

Illustration of an 'Ounce' Ounce1.jpg
Illustration of an 'Ounce'

Both the Latin name uncia and the English word ounce are derived from the Old French once, which was also used for the Eurasian lynx (Lynx lynx). Once is thought to have evolved from an earlier variant of lynx by false splitting; lonce was interpreted as l'once, in which l' is the elided form of the French definite article la ('the'), leaving once to be perceived as the animal's name. [2] The word panther derives from the classical Latin panthēra, itself from the ancient Greek πάνθηρ pánthēr, which was used for spotted cats. [3]

Taxonomy and evolution

Snow leopard skull in the collection of the Museum Wiesbaden Panthera uncia 02 MWNH 355.jpg
Snow leopard skull in the collection of the Museum Wiesbaden

Felis uncia was the scientific name used by Johann Christian Daniel von Schreber in 1777 who described a snow leopard based on an earlier description by Georges-Louis Leclerc, Comte de Buffon, assuming that the cat occurred along the Barbary Coast, in Persia, East India and China. [4] The genus name Uncia was proposed by John Edward Gray in 1854 for Asian cats with a long and thick tail. [5] Felis irbis proposed by Christian Gottfried Ehrenberg in 1830 was a skin of a female snow leopard collected in the Altai Mountains. He also clarified that several leopard (P. pardus) skins were previously misidentified as snow leopard skins. [6] Felis uncioides proposed by Thomas Horsfield in 1855 was a snow leopard skin from Nepal in the collection of the Museum of the East India Company. [7]

Uncia uncia was used by Reginald Innes Pocock in 1930 when he reviewed skins and skulls of Panthera species from Asia. He also described morphological differences between snow leopard and leopard skins. [8] Panthera baikalensis-romanii proposed by a Russian scientist in 2000 was a dark brown snow leopard skin from the Petrovsk-Zabaykalsky District in southern Transbaikal. [9]

The snow leopard was long classified in the monotypic genus Uncia. [10] It was subordinated to the genus Panthera based on results of phylogenetic studies. [11] [12] [13] [14]

Until spring 2017, there was no evidence available for the recognition of subspecies. Results of a phylogeographic analysis indicate that three subspecies should be recognised: [15]

This view has been both contested and supported by different researchers. [16] [17] [18] [19]

Additionally, an extinct subspecies Panthera uncia pyrenaica was described in 2022 based on material found in France. [20]

Evolution

Two cladograms proposed for Panthera. The upper cladogram is based on two studies published in 2006 and 2009, the lower one is based on studies published in 2010 and 2011. Two cladograms for Panthera.svg
Two cladograms proposed for Panthera. The upper cladogram is based on two studies published in 2006 and 2009, the lower one is based on studies published in 2010 and 2011.

Based on phylogenetic analysis of DNA sequence sampled across the living Felidae, the snow leopard forms a sister group with the tiger (P. tigris). Genetic divergence time of this group is estimated at 4.62 to 1.82 million years ago. [11] [21] The snow leopard and the tiger probably diverged between 3.7 to 2.7 million years ago. [12] Panthera originates most likely in northern Central Asia. Panthera blytheae excavated in western Tibet's Ngari Prefecture is the oldest known Panthera species and exhibits skull characteristics similar to the snow leopard. [23]

The mitochondrial genomes of the snow leopard, the leopard and the lion (P. leo) are more similar to each other than their nuclear genomes, indicating that their ancestors hybridised at some point in their evolution. [24]

Characteristics

Snow leopard portrait.jpg
Head of a male snow leopard
Panthera uncia (33172899150).jpg
Snow leopard showing its canine teeth
Leopard des neiges 14081.jpg
The thickly furred tail of a snow leopard

The snow leopard's fur is whitish to grey with black spots on head and neck, with larger rosettes on the back, flanks and bushy tail. The belly is whitish. Its eyes are pale green or grey in color. Its muzzle is short and its forehead domed. Its nasal cavities are large. The fur is thick with hairs between 5 and 12 cm (2.0 and 4.7 in) long. Its body is stocky, short-legged, and slightly smaller than the other cats of the genus Panthera, reaching a shoulder height of 56 cm (22 in), and ranging in head to body size from 75 to 150 cm (30 to 59 in). Its tail is 80 to 105 cm (31 to 41 in) long. [25] It weighs between 22 and 55 kg (49 and 121 lb), with an occasional large male reaching 75 kg (165 lb), and small female of under 25 kg (55 lb). [26] Its canine teeth are 28.6 mm (1.13 in) long and are more slender than those of the other Panthera species. [27] In relation to the length of its skull and width of its palate, it has large nasal openings, which allow for increasing the volume of air inhaled with each breath, and at the same time for warming and humidifying cold dry air. [28] It is not especially adapted to high-altitude hypoxia. [29]

The snow leopard shows several adaptations for living in a cold, mountainous environment. Its small rounded ears help to minimize heat loss. Its broad paws well distribute the body weight for walking on snow, and have fur on their undersides to increase the grip on steep and unstable surfaces; it also helps to minimize heat loss. Its long and flexible tail helps to maintain balance in the rocky terrain. The tail is very thick due to fat storage, and is covered in a thick layer of fur, which allows the cat to use it like a blanket to protect its face when asleep. [30]

The snow leopard differs from the other Panthera species by a shorter muzzle, an elevated forehead, a vertical chin and a less developed posterior process of the lower jaw. [8] It cannot roar despite its partly ossified hyoid bone, as its 9 mm (0.35 in) short vocal folds provide little resistance to airflow. [31] [32]

Distribution and habitat

The snow leopard is distributed from the west of Lake Baikal through southern Siberia, in the Kunlun Mountains, Altai Mountains, Sayan and Tannu-Ola Mountains, in the Tian Shan, through Tajikistan, Kyrgyzstan, Uzbekistan and Kazakhstan to the Hindu Kush in eastern Afghanistan, Karakoram in northern Pakistan, in the Pamir Mountains, the Tibetan Plateau and in the high elevations of the Himalayas in India, Nepal and Bhutan. In Mongolia, it inhabits the Mongolian and Gobi Altai Mountains and the Khangai Mountains. In Tibet, it occurs up to the Altyn-Tagh in the north. [33] [34] It inhabits alpine and subalpine zones at elevations from 3,000 to 4,500 m (9,800 to 14,800 ft), but also lives at lower elevations in the northern part of its range. [35] Potential snow leopard habitat in the Indian Himalayas is estimated at less than 90,000 km2 (35,000 sq mi) in Jammu and Kashmir, Ladakh, Uttarakhand, Himachal Pradesh, Sikkim and Arunachal Pradesh, of which about 34,000 km2 (13,000 sq mi) is considered good habitat, and 14.4% is protected. In the beginning of the 1990s, the Indian snow leopard population was estimated at 200–600 individuals living across about 25 protected areas. [34]

In summer, the snow leopard usually lives above the tree line on alpine meadows and in rocky regions at elevations from 2,700 to 6,000 m (8,900 to 19,700 ft). In winter, it descends to elevations around 1,200 to 2,000 m (3,900 to 6,600 ft). It prefers rocky, broken terrain, and can move in 85 cm (33 in) deep snow, but prefers to use existing trails made by other animals. [26]

Snow leopards were recorded by camera traps at 16 locations in northeastern Afghanistan's isolated Wakhan Corridor. [36]

Behavior and ecology

Walking in the snow Uncia uncia.jpg
Walking in the snow

The snow leopard's vocalizations include meowing, grunting, prusten and moaning. It can purr when exhaling. [25]

It is solitary and active mostly at dawn until early morning, and again in afternoons and early evenings. It mostly rests near cliffs and ridges that provide vantage points and shade. In Nepal's Shey Phoksundo National Park, the home ranges of five adult radio-collared snow leopards overlapped largely, though they rarely met. Their individual home ranges ranged in size from 12 to 39 km2 (4.6 to 15.1 sq mi). Males moved between 0.5 and 5.45 km (0.31 and 3.39 mi) per day, and females between 0.2 and 2.25 km (0.12 and 1.40 mi), measured in straight lines between survey points. Since they often zigzagged in the precipitous terrain, they actually moved up to 7 km (4.3 mi) in a single night. [37] Up to 10 individuals inhabit an area of 100 km2 (40 sq mi); in habitats with sparse prey, an area of 1,000 km2 (400 sq mi) supports only five individuals. [38]

A study in the Gobi Desert lasting from 2008 to 2014 revealed that adult male snow leopards used a mean home range of 144–270 km2 (56–104 sq mi), while adult females ranged in areas of 83–165 km2 (32–64 sq mi). Their home ranges overlapped less than 20%. These results indicate that about 40% of the 170 protected areas in snow leopard range countries are smaller than the home range of a single male snow leopard. [39]

Snow leopards leave scent marks to indicate their territories and common travel routes. They scrape the ground with the hind feet before depositing urine or feces, but also spray urine onto rocks. [26] Their urine contains many characteristic low molecular weight compounds with diverse functional groups including pentanol, hexanol, heptanol, 3-octanone, nonanal and indole, which possibly play a role in chemical communication. [40]

Hunting and diet

Wild Snow Leopard Goes Grocery Shopping.png
Snow leopard with a gray marmot in Kyrgyzstan

In Hemis National Park, a snow leopard was observed while approaching prey from above, using rocky cliffs for cover; at a distance of about 40 m (130 ft) from the prey, it walked rapidly for about 15 m (49 ft), ran the last 25 m (82 ft) and killed the prey with a neck bite. While squatting on its haunches, it ripped out clumps of hair from the abdomen and then opened it to first feed on the viscera. [41] The snow leopard is a carnivore and actively hunts its prey. Its preferred wild prey species are Himalayan blue sheep (Pseudois nayaur), Himalayan tahr (Hemitragus jemlahicus), argali (Ovis ammon), markhor (Capra falconeri) and wild goat (C. aegagrus). It also preys on domestic livestock. [42] It prefers prey ranging in weight from 36 to 76 kg (79 to 168 lb), but also hunts smaller mammals such as Himalayan marmot (Marmota himalayana), pika and vole species. Its diet depends on prey availability and varies across its range and season. In the Himalayas, it preys mostly on Himalayan blue sheep, Siberian ibex (C. sibirica), white-bellied musk deer (Moschus leucogaster) and wild boar (Sus scrofa). In the Karakoram, Tian Shan, Altai and Mongolia's Tost Mountains, its main prey consists of Siberian ibex, Thorold's deer (Cervus albirostris), Siberian roe deer (Capreolus pygargus) and argali. [43] [44] Snow leopard feces collected in northern Pakistan also contained remains of rhesus macaque (Macaca mulatta), masked palm civet (Paguma larvata), Cape hare (Lepus capensis), house mouse (Mus musculus), Kashmir field mouse (Apodemus rusiges), grey dwarf hamster (Cricetulus migratorius) and Turkestan rat (Rattus pyctoris). [45] In 2017, a snow leopard was photographed carrying a freshly killed woolly flying squirrel (Eupetaurus cinereus) near Gangotri National Park. [46] In Mongolia, domestic sheep comprises less than 20% of snow leopard diet, although wild prey has been reduced and interactions with people are common. [44]

Snow leopards actively pursue prey down steep mountainsides, using the momentum of their initial leap to chase animals for up to 300 m (980 ft). They drag the prey to a safe location and consume all edible parts of the carcass. They can survive on a single Himalayan blue sheep for two weeks before hunting again, and one adult individual apparently needs 20–30 adult blue sheep per year. [1] [26] Snow leopards have been recorded to hunt successfully in pairs, especially mating pairs. [47]

The snow leopard is capable of killing most animals in its range, with the probable exception of the adult male yak. It also eats a significant amount of vegetation, including grass and twigs. It has not been reported to attack humans, is easily driven away from livestock and readily abandons kills, often without defending itself. [26]

Reproduction and life cycle

SnowCubs01.jpg
Cubs at the Cat Survival Trust in Welwyn
Schneeleoparden Kailash und Dshamilja frontal.jpg
A female snow leopard with her cub in Zurich Zoo

Snow leopards become sexually mature at two to three years, and normally live for 15–18 years in the wild. In captivity they can live for up to 25 years. Oestrus typically lasts from five to eight days, and males tend not to seek out another partner after mating, probably because the short mating season does not allow sufficient time. Paired snow leopards mate in the usual felid posture, from 12 to 36 times a day. They are unusual among large cats in that they have a well-defined birth peak. They usually mate in late winter, marked by a noticeable increase in marking and calling. Females have a gestation period of 90–100 days, and the cubs are born between April and June. [26] A litter usually consists of two to three cubs, in exceptional cases there can be up to seven. [33]

The female gives birth in a rocky den or crevice lined with fur shed from her underside. The cubs are blind and helpless at birth, although already with a thick coat of fur, and weigh from 320 to 567 g (11.3 to 20.0 oz). Their eyes open at around seven days, and the cubs can walk at five weeks and are fully weaned by 10 weeks. The cubs leave the den when they are around two to four months of age. [26] Three radio-collared snow leopards in Mongolia's Tost Mountains gave birth between late April and late June. Two female cubs started to part from their mothers at the age of 20 to 21 months, but reunited with them several times for a few days over a period of 4–7 months. One male cub separated from its mother at the age of about 22 months, but stayed in her vicinity for a month and moved out of his natal range at 23 months of age. [48]

The snow leopard has a generation length of eight years. [49]

Threats

The major threat to snow leopard populations is poaching and illegal trade of skins and body parts. [1] Between 1999 and 2002, three live snow leopard cubs and 16 skins were confiscated, 330 traps were destroyed and 110 poachers were arrested in Kyrgyzstan. Undercover operations in the country revealed an illegal trade network with links to Russia and China via Kazakhstan. The major skin trade center in the region is the city of Kashgar in Xinjiang. [50] In Tibet and Mongolia, skins are used for traditional dresses, and meat in traditional Tibetan medicine to cure kidney problems; bones are used in traditional Chinese and Mongolian medicine for treating rheumatism, injuries and pain of human bones and tendons. Between 1996 and 2002, 37 skins were found in wildlife markets and tourist shops in Mongolia. [51] Between 2003 and 2016, 710 skins were traded, of which 288 skins were confiscated. In China, 103 to 236 animals are poached every year, in Mongolia between 34 and 53, in Pakistan between 23 and 53, in India from 21 to 45, and in Tajikistan 20 to 25. In 2016, a survey of Chinese websites revealed 15 advertisements for 44 snow leopard products; the dealers offered skins, canine teeth, claws and a tongue. [52] Nine snow leopard skins were found during a market survey in September 2014 in Afghanistan. [53]

Greenhouse gas emissions will likely cause a shift of the treeline in the Himalayas and a shrinking of the alpine zone, which may reduce snow leopard habitat by 30%. [54]

Where snow leopards prey on domestic livestock, they are subject to conflict with humans. [1] The loss of natural prey due to overgrazing by livestock, poaching, and defense of livestock are the major drivers for the decreasing population of the snow leopard. [26] Livestock also cause habitat degradation, which, alongside the increasing use of forests for fuel, reduces snow leopard habitat. [55]

Conservation

Global snow leopard population
CountryYearEstimate
Afghanistan201650–200 [56]
Bhutan201679–112 [57]
China20164,500 [58]
India2016516–524 [59]
Kazakhstan2016100–120 [60]
Kyrgyzstan2016300–400 [61]
Mongolia20161,000 [62]
Nepal2016301–400 [63]
Pakistan2016250-420 [64]
Russia201670–90 [65]
Tajikistan2016250–280 [66]
Uzbekistan201630–120 [67]

The snow leopard is listed in CITES Appendix I. [68] It has been listed as threatened with extinction in Schedule I of the Convention on the Conservation of Migratory Species of Wild Animals since 1985. [51] Hunting snow leopards has been prohibited in Kyrgyzstan since the 1950s. [50] In India, the snow leopard is granted the highest level of protection under the Wildlife Protection Act, 1972, and hunting is sentenced with imprisonment of 3–7 years. [59] In Nepal, it has been legally protected since 1973, with penalties of 5–15 years in prison and a fine for poaching and trading it. [69] Since 1978, it has been listed in the Soviet Union’s Red Book and is still inscribed today in the Red Data Book of the Russian Federation as threatened with extinction. Hunting snow leopards is only permitted for the purposes of conservation and monitoring, and to eliminate a threat to the life of humans and livestock. Smuggling of snow leopard body parts is punished with imprisonment and a fine. [70] Hunting snow leopards has been prohibited in Afghanistan since 1986. [53] In China, it has been protected by law since 1989; hunting and trading snow leopards or their body parts constitute a criminal offence that is punishable by the confiscation of property, a fine and a sentence of at least 10 years in prison. [71] It has been protected in Bhutan since 1995. [57]

At the end of 2020, 35 cameras were installed on the outskirts of Almaty, Kazakhstan in hopes to catch footage of snow leopards. In November 2021, it was announced by the Russian World Wildlife Fund (WWF) that snow leopards were spotted 65 times on these cameras in the Trans-Ili Alatau mountains since the cameras were installed. [72]

Snow leopards inhabit the following protected areas:

Global Snow Leopard Forum

In 2013, government leaders and officials from all 12 countries encompassing the snow leopard's range (Afghanistan, Bhutan, China, India, Kazakhstan, Kyrgyzstan, Mongolia, Nepal, Pakistan, Russia, Tajikistan, and Uzbekistan) came together at the Global Snow Leopard Forum (GSLF) initiated by the then-President of Kyrgyzstan Almazbek Atambayev, and the State Agency on Environmental Protection and Forestry under the government of Kyrgyzstan. The meeting was held in Bishkek, and all countries agreed that the snow leopard and the high mountain habitat need trans-boundary support to ensure a viable future for snow leopard populations, and to safeguard its fragile environment. The event brought together many partners, including NGOs like the Snow Leopard Conservancy, the Snow Leopard Trust, and the Nature and Biodiversity Conservation Union. Also supporting the initiative were the Snow Leopard Network, the World Bank's Global Tiger Initiative, the United Nations Development Programme, the World Wild Fund for Nature, the United States Agency for International Development, and Global Environment Facility. [82]

At the GSLF meeting, the 12 range countries signed the Bishkek Declaration, which stated: "[We] acknowledge that the snow leopard is an irreplaceable symbol of our nations' natural and cultural heritage and an indicator of the health and sustainability of mountain ecosystems; and we recognize that mountain ecosystems inhabited by snow leopards provide essential ecosystem services, including storing and releasing water from the origins of river systems benefitting one-third of the world’s human population; sustaining the pastoral and agricultural livelihoods of local communities which depend on biodiversity for food, fuel, fodder, and medicine; and offering inspiration, recreation, and economic opportunities."

In captivity

Snow leopard in the San Diego Zoo Lightmatter snowleopard.jpg
Snow leopard in the San Diego Zoo

The Moscow Zoo exhibited the first captive snow leopard in 1872 that had been caught in Turkestan. In Kyrgyzstan, 420 live snow leopards were caught between 1936 and 1988 and exported to zoos around the world. The first captive bred snow leopard cubs were born in the 1990s in the Beijing Zoo. [50] The Snow Leopard Species Survival Plan was initiated in 1984; by 1986, American zoos held 234 individuals. [83]

Cultural significance

Snow leopard on the reverse of the old 10,000-Kazakhstani tenge banknote SnowLeopard10000KZT.jpg
Snow leopard on the reverse of the old 10,000-Kazakhstani tenge banknote

The snow leopard is widely used in heraldry and as an emblem in Central Asia. It has long been used as a political symbol, the Aq Bars ('White Leopard'), by Tatars, Kazakhs, and Bulgars. A snow leopard is depicted on the official seal of Almaty and on the former 10,000 Kazakhstani tenge banknote. A mythical winged Aq Bars is depicted on the national coat of arms of Tatarstan, the seal of the city of Samarqand, Uzbekistan and the old coat of arms of Nur-Sultan. In Kyrgyzstan, it has been used in highly stylized form in the modern emblem of the capital Bishkek, and the same art has been integrated into the badge of the Kyrgyzstan Girl Scouts Association. A crowned snow leopard features in the arms of Shushensky District in Russia. It is the state animal of Ladakh and Himachal Pradesh in India. [84] [85]

See also

Related Research Articles

<span class="mw-page-title-main">Felidae</span> Family of mammals

Felidae is the family of mammals in the order Carnivora colloquially referred to as cats. A member of this family is also called a felid. The term "cat" refers both to felids in general and specifically to the domestic cat.

<span class="mw-page-title-main">Jaguar</span> Large cat native to the Americas

The jaguar is a large cat species and the only living member of the genus Panthera native to the Americas. With a body length of up to 1.85 m and a weight of up to 158 kg (348 lb), it is the largest cat species in the Americas and the third largest in the world. Its distinctively marked coat features pale yellow to tan colored fur covered by spots that transition to rosettes on the sides, although a melanistic black coat appears in some individuals. The jaguar's powerful bite allows it to pierce the carapaces of turtles and tortoises, and to employ an unusual killing method: it bites directly through the skull of mammalian prey between the ears to deliver a fatal blow to the brain.

A black panther is the melanistic colour variant of the leopard and the jaguar. Black panthers of both species have excess black pigments, but their typical rosettes are also present. They have been documented mostly in tropical forests, with black leopards in Kenya, India, Sri Lanka, Nepal, Thailand, Peninsular Malaysia and Java, and black jaguars of the Americas in Mexico, Panama, Costa Rica, Brazil and Paraguay. Melanism is caused by a recessive allele in the leopard, and by a dominant allele in the jaguar.

<span class="mw-page-title-main">Leopard</span> Large cat native to Africa and Asia

The leopard is one of the five extant species in the genus Panthera, a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, and on the Indian subcontinent to Southeast and East Asia. It is listed as Vulnerable on the IUCN Red List because leopard populations are threatened by habitat loss and fragmentation, and are declining in large parts of the global range. The leopard is considered locally extinct in Hong Kong, Singapore, South Korea, Jordan, Morocco, Togo, the United Arab Emirates, Uzbekistan, Lebanon, Mauritania, Kuwait, Syria, Libya, Tunisia and most likely in North Korea, Gambia, Laos, Lesotho, Tajikistan, Vietnam and Israel. Contemporary records suggest that the leopard occurs in only 25% of its historical global range.

Panthera is a genus within the family Felidae that was named and described by Lorenz Oken in 1816 who placed all the spotted cats in this group. Reginald Innes Pocock revised the classification of this genus in 1916 as comprising the tiger, lion, jaguar, and leopard on the basis of common cranial features. Results of genetic analysis indicate that the snow leopard also belongs to the genus Panthera, a classification that was accepted by IUCN Red List assessors in 2008. The lion is the only species with anatomical structures that enable them to roar; the snow leopard cannot. The primary reason for this was formerly assumed to be the incomplete ossification of the hyoid bones studies show the ability to roar is due to other morphological features, especially of the larynx.

<span class="mw-page-title-main">Clouded leopard</span> Species of wild cat

The clouded leopard, also called the mainland clouded leopard, is a wild cat inhabiting dense forests from the foothills of the Himalayas through mainland Southeast Asia into South China. In the early 19th century, a clouded leopard was brought to London from China and described in 1821. It has large dusky-grey blotches and irregular spots and stripes reminiscent of clouds. Its head-and-body length ranges from 68.6 to 108 cm with a 61 to 91 cm long tail. It uses its tail for balancing when moving in trees and is able to climb down vertical tree trunks head first. It rests in trees during the day and hunts by night on the forest floor.

The term "big cat" is typically used to refer to any of the five living members of the genus Panthera, namely the tiger, lion, jaguar, leopard, and snow leopard, as well as the non-pantherine cheetah and cougar.

<span class="mw-page-title-main">Pallas's cat</span> Small wild cat species (Otocolobus manul)

The Pallas's cat, also known as the manul, is a small wild cat with long and dense light grey fur, native to mountainous regions across central Asia.

<span class="mw-page-title-main">Pantherinae</span> Subfamily of felids

Pantherinae is a subfamily within the family Felidae; it was named and first described by Reginald Innes Pocock in 1917 as only including the Panthera species. The Pantherinae genetically diverged from a common ancestor between 9.32 to 4.47 million years ago and 10.67 to 3.76 million years ago.

<span class="mw-page-title-main">African leopard</span> Leopard subspecies

The African leopard is the nominate subspecies of the leopard, native to many countries in Africa. It is widely distributed in most of sub-Saharan Africa, but the historical range has been fragmented in the course of habitat conversion. Leopards have also been recorded in North Africa as well.

<span class="mw-page-title-main">Arabian leopard</span> Leopard subspecies in the Arabian Peninsula

The Arabian leopard is a leopard subspecies native to the Arabian Peninsula. It has been listed as Critically Endangered on the IUCN Red List since 1996 as fewer than 200 wild individuals were estimated to be alive in 2006. The population is severely fragmented. Subpopulations are isolated and not larger than 50 mature individuals. The population is thought to decline continuously.

<span class="mw-page-title-main">Indian leopard</span> Leopard subspecies

The Indian leopard is a leopard subspecies widely distributed on the Indian subcontinent. The species Panthera pardus is listed as Vulnerable on the IUCN Red List because populations have declined following habitat loss and fragmentation, poaching for the illegal trade of skins and body parts, and persecution due to conflict situations. The Indian leopard is one of the big cats occurring on the Indian subcontinent, along with the Asiatic lion, Bengal tiger, snow leopard and clouded leopard. In 2014, a national census of leopards around tiger habitats was carried out in India except the northeast. 7,910 individuals were estimated in surveyed areas and a national total of 12,000–14,000 speculated.

Afghanistan has long been known for diverse wildlife. Many of the larger mammals in the country are categorized by the International Union for Conservation of Nature (IUCN) as globally threatened. These include the snow leopard, Marco Polo sheep, Siberian musk deer, markhor, urial, and the Asiatic black bear. Other species of interest are the ibex, the gray wolf, and the brown bear, striped hyenas, and numerous bird of prey species. Most of the Marco Polo sheep and ibex are being poached for food, whereas wolves, snow leopards and bears are being killed for damage prevention.

<span class="mw-page-title-main">Sunda clouded leopard</span> Species of carnivore

The Sunda clouded leopard is a medium-sized wild cat native to Borneo and Sumatra. It is listed as Vulnerable on the IUCN Red List since 2015, as the total effective population probably consists of fewer than 10,000 mature individuals, with a decreasing population trend. On both Sunda islands, it is threatened by deforestation. It was classified as a separate species, distinct from the clouded leopard in mainland Southeast Asia based on a study in 2006. Its fur is darker with a smaller cloud pattern.

<i>Panthera pardus tulliana</i> Leopard subspecies

Panthera pardus tulliana is a leopard subspecies native to the Iranian Plateau and surrounding areas encompassing Turkey, the Caucasus, Azerbaijan, Georgia, Armenia, Iraq, Iran, Turkmenistan, Afghanistan and possibly Pakistan. Since 2016, it has been listed as Endangered on the IUCN Red List, as the wild population is estimated at less than 1,000 mature individuals.

<span class="mw-page-title-main">Prusten</span>

Prusten is a form of communicative behaviour exhibited by some members of the family Felidae. Prusten is also referred to as chuffing or chuffle. It is described as a short, low intensity, non-threatening vocalization. In order to vocalize a chuff, the animal's mouth is closed and air is blown through the nostrils, producing a breathy snort. It is typically accompanied by a head bobbing movement. It is often used between two cats as a greeting, during courting, or by a mother comforting her cubs. The vocalization is produced by tigers, jaguars, snow leopards and clouded leopards. Prusten has significance in both the fields of evolution and conservation.

<span class="mw-page-title-main">Snow Leopard Trust</span>

The Snow Leopard Trust is the largest and oldest organization working solely to protect the endangered snow leopard and its habitat in 12 countries of Central Asia. The trust is a non-profit organization with its headquarters in Seattle, Washington. The present total population of snow leopards in the wild is estimated at between 3,920 and 6,390.

<span class="mw-page-title-main">Amur leopard</span> Leopard subspecies in Far East Asia

The Amur leopard is a leopard subspecies native to the Primorye region of southeastern Russia and northern China. It is listed as Critically Endangered on the IUCN Red List, as in 2007, only 19–26 wild leopards were estimated to survive in southeastern Russia and northeastern China. It is considered one of the rarest cats on Earth.

<span class="mw-page-title-main">Indochinese leopard</span> Leopard subspecies

The Indochinese leopard is a leopard subspecies native to mainland Southeast Asia and southern China. In Indochina, leopards are rare outside protected areas and threatened by habitat loss due to deforestation as well as poaching for the illegal wildlife trade. The population trend is suspected to be decreasing. As of 2016, the population is thought to comprise 973–2,503 mature individuals, with only 409–1,051 breeding adults. The historical range has decreased by more than 90%.

References

  1. 1 2 3 4 5 6 McCarthy, T.; Mallon, D.; Jackson, R.; Zahler, P. & McCarthy, K. (2017). "Panthera uncia". IUCN Red List of Threatened Species . 2017: e.T22732A50664030. doi: 10.2305/IUCN.UK.2017-2.RLTS.T22732A50664030.en . Retrieved 19 November 2021.
  2. Allen, E. A. (1908). "English Doublets". Publications of the Modern Language Association of America. New Series 16. 23 (1): 184–239. doi:10.2307/456687. JSTOR   456687. S2CID   251028590.
  3. Liddell, H. G. & Scott, R. (1940). "πάνθηρ". A Greek-English Lexicon (Revised and augmented ed.). Oxford: Clarendon Press.
  4. Schreber, J. C. D. (1777). "Die Unze". Die Säugethiere in Abbildungen nach der Natur mit Beschreibungen. Erlangen: Wolfgang Walther. pp. 386–387.
  5. Gray, J. E. (1854). "The ounces". Annals and Magazine of Natural History. 2. 14: 394.
  6. Ehrenberg, C. G. (1830). "Observations et données nouvelles sur le tigre du nord et la panthère du nord, recueillies dans le voyage de Sibérie fait par M.A. de Humboldt, en l'année 1829". Annales des sciences naturelles, Zoologie. 21: 387–412.
  7. Horsfield, T. (1855). "Brief notices of several new or little-known species of Mammalia, lately discovered and collected in Nepal, by Brian Houghton Hodgson". The Annals and Magazine of Natural History: Including Zoology, Botany, and Geology. 2. 16 (92): 101–114. doi:10.1080/037454809495489.
  8. 1 2 Pocock, R. I. (1930). "The panthers and ounces of Asia. Part II. The panthers of Kashmir, India, and Ceylon". Journal of the Bombay Natural History Society. 34 (2): 307–336.
  9. Medvedev, D. G. (2000). "Morfologicheskie otlichiya irbisa iz Yuzhnogo Zabaikalia" [Morphological differences of the snow leopard from Southern Transbaikalia]. Vestnik Irkutskoi Gosudarstvennoi Sel'skokhozyaistvennoi Akademyi [Proceedings of Irkutsk State Agricultural Academy]. 20: 20–30.
  10. Wozencraft, W. C. (2005). "Species Uncia uncia". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 548. ISBN   978-0-8018-8221-0. OCLC   62265494.
  11. 1 2 3 Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E. & O'Brien, S. J. (2006). "The late Miocene radiation of modern Felidae: a genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID   16400146. S2CID   41672825.
  12. 1 2 3 Davis, B. W.; Li, G. & Murphy, W. J. (2010). "Supermatrix and species tree methods resolve phylogenetic relationships within the big cats, Panthera (Carnivora: Felidae)" (PDF). Molecular Phylogenetics and Evolution. 56 (1): 64–76. doi:10.1016/j.ympev.2010.01.036. PMID   20138224.[ dead link ]
  13. Kitchener, A. C.; Driscoll, C. A. & Yamaguchi, N. (2016). "What is a Snow Leopard? Taxonomy, Morphology, and Phylogeny". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 3–11. ISBN   9780128024966.
  14. Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O’Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.; Nowell, K.; Timmons, Z. & Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Cat News (Special Issue 11): 69.
  15. Janecka, J. E.; Zhang, Y.; Li, D.; Munkhtsog, B.; Bayaraa, M.; Galsandorj, N.; Wangchuk, T. R.; Karmacharya, D.; Li, J.; Lu, Z. & Uulu, K. Z. (2017). "Range-Wide Snow Leopard Phylogeography Supports Three Subspecies". Journal of Heredity. 108 (6): 597–607. doi: 10.1093/jhered/esx044 . PMID   28498961.
  16. Senn, H.; Murray-Dickson, G.; Kitchener, A. C.; Riordan, P. & Mallon, D. (2018). "Response to Janecka et al. 2017". Heredity. 120 (6): 581–585. doi:10.1038/s41437-017-0015-4. PMC   5943311 . PMID   29225352.
  17. Janecka, J. E.; Janecka, M. J.; Helgen, K. M. & Murphy, W. J. (2018). "The validity of three snow leopard subspecies: response to Senn et al". Heredity. 120 (6): 586–590. doi:10.1038/s41437-018-0052-7. PMC   5943360 . PMID   29434338.
  18. Janecka, J. E.; Hacker, C.; Broderick, J.; Pulugulla, S.; Auron, P.; Ringling, M.; Nelson, B.; Munkhtsog, B.; Hussain, S.; Davis, B. & Jackson, R. (2020). "Noninvasive genetics and genomics shed light on the status, phylogeography, and evolution of the elusive Snow Leopard". In Ortega, J. & Maldonado, J. E. (eds.). Conservation Genetics in Mammals. Integrative Research Using Novel Approaches. Basel: Springer International Publishing. pp. 83–120. doi:10.1007/978-3-030-33334-8_5. ISBN   978-3-030-33334-8. S2CID   213437425.
  19. Korablev, M.; Poyarkov, A. D.; Karnaukhov, A. S.; Zvychaynaya, E. Y.; Kuksin, A. N.; Malykh, S. V.; Istomov, S. V.; Spitsyn, S. V.; Aleksandrov, D. Y.; Hernandez-Blanco, J. A. & Munkhtsog, B. (2021). "Large‑scale and fine‑grain population structure and genetic diversity of snow leopards (Panthera uncia Schreber, 1776) from the northern and western parts of the range with an emphasis on the Russian population" (PDF). Conservation Genetics. 22 (3): 397–410. doi:10.1007/s10592-021-01347-0. S2CID   233480791.
  20. Hemmer, H. (2022). "An intriguing find of an early Middle Pleistocene European snow leopard, Panthera uncia pyrenaica ssp. nov. (Mammalia, Carnivora, Felidae), from the Arago cave (Tautavel, Pyrénées-Orientales, France)". Palaeobiodiversity and Palaeoenvironments. Online edition. doi: 10.1007/s12549-021-00514-y . S2CID   246433218.
  21. 1 2 Werdelin, L.; Yamaguchi, N.; Johnson, W. E. & O'Brien, S. J. (2010). "Phylogeny and evolution of cats (Felidae)". In Macdonald, D. W. & Loveridge, A. J. (eds.). Biology and Conservation of Wild Felids. Oxford, UK: Oxford University Press. pp. 59–82. ISBN   978-0-19-923445-5.
  22. Mazák, J.H.; Christiansen, P.; Kitchener, A.C. & Goswami, A. (2011). "Oldest known pantherine skull and evolution of the tiger". PLOS ONE. 6 (10): e25483. Bibcode:2011PLoSO...625483M. doi: 10.1371/journal.pone.0025483 . PMC   3189913 . PMID   22016768.
  23. Tseng, Z. J.; Wang, X.; Slater, G. J.; Takeuchi, G. T.; Li, Q.; Liu, J. & Xie, G. (2014). "Himalayan fossils of the oldest known pantherine establish ancient origin of big cats". Proceedings of the Royal Society B: Biological Sciences. 281 (1774): 20132686. doi:10.1098/rspb.2013.2686. PMC   3843846 . PMID   24225466.
  24. Li, G.; Davis, B. W.; Eizirik, E. & Murphy, W. J. (2016). "Phylogenomic evidence for ancient hybridization in the genomes of living cats (Felidae)". Genome Research. 26 (1): 1–11. doi:10.1101/gr.186668.114. PMC   4691742 . PMID   26518481.
  25. 1 2 Hemmer, H. (1972). "Uncia uncia" (PDF). Mammalian Species (20): 1–5. doi:10.2307/3503882. JSTOR   3503882. Archived from the original (PDF) on 2012-04-01.
  26. 1 2 3 4 5 6 7 8 Sunquist, M. & Sunquist, F. (2002). "Snow leopard Uncia uncia (Schreber, 1775)". Wild Cats of the World. Chicago: University of Chicago Press. pp. 377–394. ISBN   978-0-226-77999-7.
  27. Christiansen, P. (2007). "Canine morphology in the larger Felidae: implications for feeding ecology". Biological Journal of the Linnean Society. 91 (4): 573–592. doi: 10.1111/j.1095-8312.2007.00819.x .
  28. Torregrosa, V.; Petrucci, M.; Pérez-Claros, J. A. & Palmqvist, P. (2010). "Nasal aperture area and body mass in felids: Ecophysiological implications and paleobiological inferences". Geobios. 43 (6): 653–661. doi:10.1016/j.geobios.2010.05.001.
  29. Janecka, J.E.; Nielsen, S.S.; Andersen, S.D.; Hoffmann, F.G.; Weber, R.E.; Anderson, T.; Storz, J.F. & Fago, A. (2015). "Genetically based low oxygen affinities of felid hemoglobins: lack of biochemical adaptation to high-altitude hypoxia in the snow leopard". Journal of Experimental Biology. 218 (15): 2402–2409. doi: 10.1242/jeb.125369 . PMC   4528707 . PMID   26246610.
  30. Chadwick, D. H. (2008). "Out of the Shadows". National Geographic. Retrieved 2010-01-29.
  31. Hast, M. H. (1989). "The larynx of roaring and non-roaring cats". Journal of Anatomy. 163: 117–121. PMC   1256521 . PMID   2606766.
  32. Weissengruber, G. E.; Forstenpointner, G.; Peters, G.; Kübber-Heiss, A.; Fitch, W. T. (2002). "Hyoid apparatus and pharynx in the lion (Panthera leo), jaguar (Panthera onca), tiger (Panthera tigris), cheetah (Acinonyx jubatus) and domestic cat (Felis silvestris f. catus)". Journal of Anatomy. 201 (3): 195–209. doi:10.1046/j.1469-7580.2002.00088.x. PMC   1570911 . PMID   12363272.
  33. 1 2 Heptner, V. G.; Sludskij, A. A. (1992) [1972]. "Snow Leopard, Ounce [Irbis]". Mlekopitajuščie Sovetskogo Soiuza. Moskva: Vysšaia Škola[Mammals of the Soviet Union. Volume II, Part 2. Carnivora (Hyaenas and Cats)]. Washington DC: Smithsonian Institution and the National Science Foundation. pp. 276–319.
  34. 1 2 McCarthy, T. M. & Chapron, G. (2003). Snow Leopard Survival Strategy (PDF). Seattle, USA: International Snow Leopard Trust and Snow Leopard Network.
  35. Janečka, J. E.; Jackson. R.; Yuquang, Z.; Diqiang, L.; Munkhtsog, B.; Buckley-Beason, V.; Murphy, W. J. (2008). "Population monitoring of snow leopards using noninvasive collection of scat samples: a pilot study". Animal Conservation. 11 (5): 401–411. doi:10.1111/j.1469-1795.2008.00195.x. S2CID   20787622.
  36. Simms, A.; Moheb, Z.; Salahudin; Ali, H.; Ali, I.; Wood, T. (2011). "Saving threatened species in Afghanistan: snow leopards in the Wakhan Corridor". International Journal of Environmental Studies. 68 (3): 299–312. doi:10.1080/00207233.2011.577147. S2CID   96170915.
  37. Jackson, R. & Ahlborn, G. (1988). "Observations on the ecology of snow leopard in west Nepal" (PDF). In Freeman, H. (ed.). Proceedings of the Fifth International Snow Leopard Symposium. India: International Snow Leopard Trust. pp. 65–97.
  38. Jackson, R. (1996). Home Range, Movements and Habitat Use of Snow Leopard in Nepal (PhD). London: University of London.
  39. 1 2 3 4 5 6 7 8 9 10 Johansson, Ö.; Rauset, G. R.; Samelius, G.; McCarthy, T.; Andrén, H.; Tumursukh, L. & Mishra, C. (2016). "Land sharing is essential for snow leopard conservation" (PDF). Biological Conservation. 203 (203): 1–7. doi:10.1016/j.biocon.2016.08.034.
  40. Das, S.; Manna, S.; Ray, S.; Das, P.; Rai, U.; Ghosh, B. & Sarkar, M. P. (2019). "Do Urinary Volatiles Carry Communicative Messages in Himalayan Snow Leopards [Panthera uncia, (Schreber, 1775)]?". In Buesching, C. (ed.). Chemical Signals in Vertebrates. Vol. 14. Cham: Springer. pp. 27–37. doi:10.1007/978-3-030-17616-7_3. ISBN   978-3-030-17615-0. S2CID   200084900.
  41. Fox, J. L. & Chundawat, R. S. (1988). "Observations of snow leopard stalking, killing and feeding behavior" (PDF). Mammalia. 52 (1): 137–140.
  42. Johansson, Ö.; McCarthy, T.; Samelius, G.; Andrén, H.; Tumursukh, L. & Mishra, C. (2015). "Snow leopard predation in a livestock dominated landscape in Mongolia" (PDF). Biological Conservation. 184: 251–258. doi:10.1016/j.biocon.2015.02.003.
  43. Lyngdoh, S.; Shrotriya, S.; Goyal, S. P., Clements, H.; Hayward, M. W. & Habib, B. (2014). "Prey preferences of the snow leopard (Panthera uncia): regional diet specificity holds global significance for conservation". PLOS ONE. 9 (2): e88349. Bibcode:2014PLoSO...988349L. doi: 10.1371/journal.pone.0088349 . PMC   3922817 . PMID   24533080.
  44. 1 2 Shehzad, W.; McCarthy, T. M.; Pompanon, F.; Purevjav, L.; Coissac, E.; Riaz, T. & Taberlet, P. (2012). "Prey Preference of Snow Leopard (Panthera uncia) in South Gobi, Mongolia". PLOS ONE. 7 (2): e32104. Bibcode:2012PLoSO...732104S. doi: 10.1371/journal.pone.0032104 . PMC   3290533 . PMID   22393381.
  45. Khatoon, R.; Hussain, I.; Anwar, M. & Nawaz, M. A. (2017). "Diet selection of snow leopard (Panthera uncia) in Chitral, Pakistan". Turkish Journal of Zoology. 41 (41): 914–923. doi:10.3906/zoo-1604-58.
  46. Pal, R.; Bhattacharya, T. & Sathyakumar, S. (2020). "Woolly flying squirrel Eupetaurus cinereus: A new addition to the diet of snow leopard Panthera uncia" (PDF). Journal of the Bombay Natural History Society. 117. doi:10.17087/jbnhs/2020/v117/142056 (inactive 31 December 2022). Archived from the original (PDF) on 21 September 2020. Retrieved 2 August 2021.{{cite journal}}: CS1 maint: DOI inactive as of December 2022 (link)
  47. Macri, A. M. & Patterson-Kane, E. (2011). "Behavioural analysis of solitary versus socially housed snow leopards (Panthera uncia), with the provision of simulated social contact". Applied Animal Behaviour Science. 130 (3–4): 115–123. doi:10.1016/j.applanim.2010.12.005.
  48. Johansson, Ö.; Ausilio, G.; Low, M.; Lkhagvajav, P.; Weckworth, B. & Sharma, K. (2021). "The timing of breeding and independence for snow leopard females and their cubs". Mammalian Biology. 101 (2): 173–180. doi: 10.1007/s42991-020-00073-3 . S2CID   225114786.
  49. Pacifici, M.; Santini, L.; Di Marco, M.; Baisero, D.; Francucci, L.; Grottolo Marasini, G.; Visconti, P. & Rondinini, C. (2013). "Generation length for mammals". Nature Conservation (5): 87–94.
  50. 1 2 3 Dexel, B. (2002). The Illegal Trade in Snow Leopards – A Global Perspective. Berlin: German Society for Nature Conservation. CiteSeerX   10.1.1.498.7184 .
  51. 1 2 Theile, S. (2003). Fading footprints; the killing and trade of snow leopards (PDF). Cambridge, UK: TRAFFIC International. ISBN   1-85850-201-2.
  52. Nowell, K.; Li, J.; Paltsyn, M. & Sharma, R.K. (2016). An Ounce of Prevention: Snow Leopard Crime Revisited (PDF). Cambridge, UK: TRAFFIC International. ISBN   978-1-85850-409-4.
  53. 1 2 Maheshwari, A.; Niraj, S. K.; Sathyakumar, S.; Thakur, M. & Sharma, L. K. (2016). "Snow leopard illegal trade in Afghanistan: A rapid survey". Cat News (64): 22–23.
  54. Forrest, J. L.; Wikramanayake, E.; Shrestha, R.; Areendran, G.; Gyeltshen, K.; Maheshwari, A.; Mazumdar, S.; Naidoo, R.; Thapa, G. J. & Thapa, K. (2012). "Conservation and climate change: Assessing the vulnerability of snow leopard habitat to treeline shift in the Himalaya" (PDF). Biological Conservation. 150 (1): 129–135. doi:10.1016/j.biocon.2012.03.001.
  55. "What water means to snow leopards". UNDP. 2 June 2022. Retrieved 6 August 2022.
  56. Moheb, Z. & Paley, R. (2016). "Central Asia: Afghanistan". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 409–417. ISBN   9780128024966.
  57. 1 2 Lham, D.; Thinley, P.; Wangchuk,S.; Wangchuk, N.; Lham, K.; Namgay, T.; Tharchen, L. & Wangchuck, T. (2016). National Snow Leopard Survey of Bhutan – Phase II: Camera Trap Survey for Population Estimation (Report). Thimphu, Bhutan: Wildlife Conservation Division, Department of Forests and Park Services.
  58. Liu, Y.; Weckworth, B.; Li, J.; Xiao, L.; Zhao, X. & Lu, Z. (2016). "China: The Tibetan Plateau, Sanjiangyuan Region". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 513–521. ISBN   9780128024966.
  59. 1 2 Bhatnagar, Y. V.; Mathur, V. B.; Sathyakumar, S.; Ghoshal, A.; Sharma, R. K.; Bijoor, A.; Raghunath, R.; Timbadia, R. & Lal, P. (2016). "South Asia: India". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 457–470. ISBN   9780128024966.
  60. Loginov, O. (2016). "Central Asia: Kazakhstan". In McCarthy, T.; Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 427–430. ISBN   9780128024966.
  61. Daveltbakov, A.; Rosen, T.; Anarbaev, M.; Kubanychbekov, Z.; Jumabai uulu, K.; Samanchina, J. & Sharma, K. (2016). "Central Asia: Kyrgyzstan". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 419–425. ISBN   9780128024966.
  62. Munkhtsok, B.; Purevjav, L.; McCarthy, T. & Bayrakçismith, R. (2016). "Northern Range: Mongolia". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 493–500. ISBN   9780128024966.
  63. Ale, S.; Shah, K. B.; Jackson, R. M. & Rosen, T. (2016). "South Asia: Nepal". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 471–479. ISBN   9780128024966.
  64. Khan, A. (2016). "South Asia: Pakistan". In McCarthy, T.; Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 481–491. ISBN   9780128024966.
  65. Paltsyn, M.; Poyarkov, A.; Spitsyn, S.; Kuksin, A.; Istomov, S.; Gibbs, J.P.; Jackson, R. M.; Castner, J.; Kozlova, S.; Karnaukhov, A. & Malykh, S. (2016). "Northern Range: Russia". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 501–511. ISBN   9780128024966.
  66. Saidov, A.; Karimov, K.; Amirov, Z. & Rosen, T. (2016). "Central Asia: Tajikistan". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 433–437. ISBN   9780128024966.
  67. Esipov, A.; Bykova, E.; Protas, Y. & Aromov, B. (2016). "Central Asia: Uzbekistan". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards: Biodiversity of the World: Conservation from Genes to Landscapes. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 439–448. ISBN   9780128024966.
  68. Nowell, K.; Jackson, P. (1996). "Snow leopard, Uncia uncia". Wild cats: Status survey and conservation action plan. Gland, Switzerland: International Union for Conservation of Nature and Natural Resources. pp. 91–95. ISBN   9782831700458.
  69. Kattel, B. & Bajiimaya, S. (1995). "Status and conservation of Snow Leopard in Nepal". In Jackson, R. & Ahmad, A. A. (eds.). Proceedings of the Eighth International Snow Leopard Symposium, 12–16 November 1995, Islamabad, Pakistan. Islamabad, Pakistan: International Snow Leopard Trust. pp. 21–27.
  70. Paltsyn, M.Y.; Spitsyn, S.V.; Kuksin, A.N. & Istomov, S.V. (2012). Snow Leopard Conservation in Russia – Data for Conservation Strategy for Snow Leopard in Russia (Report). Krasnoyarsk: WWF Russia.
  71. Riordan, P. & Kun, S. (2010). "The Snow Leopard in China". Cat News (Special Issue 5): 14–17.
  72. November 2021, Saniya Bulatkulova in Kazakhstan Region Profiles: A. Deep Dive Into the Heart of Central Asia on 21 (2021-11-21). "Snow Leopards Caught On Camera 65 Times and Counting This Year Alone in Almaty Region". The Astana Times. Retrieved 2021-11-23.
  73. Esipov, A.; Bykova, E.; Protas, Y. & Aromov, B. (2016). "Central Asia: Uzbekistan". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 439–447. ISBN   9780128024966.
  74. Jackson, R. (1998). "People-Wildlife Conflict Management in the Qomolangma Nature Preserve, Tibet" (PDF). In Wu Ning; D. Miller; Lhu Zhu; J. Springer (eds.). Tibet's Biodiversity: Conservation and Management. Proceedings of a Conference, August 30 – September 4, 1998. Tibet Forestry Department and World Wide Fund for Nature. pp. 40–46.
  75. Liu, Y.; Weckworth, B.; Li, J.; Xiao, L.; Zhao, X. & Lu, Z. (2016). "China: The Tibetan Plateau, Sanjiangyuan Region". In McCarthy, T. & Mallon, D. (eds.). Snow Leopards. Amsterdam, Boston, Heidelberg, London, New York: Academic Press. pp. 513–521. ISBN   9780128024966.
  76. Ming, M.; Feng, X.; Turghan, M. & Shoujin, Y. (2004). Report on Snow Leopard (Uncia uncia) Surveys in Tomur, Xinjiang, China (PDF). Xinjiang: Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences.
  77. Alexander, J. S.; Shi, K.; Tallents, L. A. & Riordan, P. (2016). "On the high trail: examining determinants of site use by the Endangered snow leopard Panthera uncia in Qilianshan, China" (PDF). Oryx. 50 (2): 231–238. doi:10.1017/S0030605315001027. S2CID   88193382.
  78. Mishra, C. (1997). "Livestock depredation by large carnivores in the Indian trans-Himalaya: conflict perceptions and conservation prospects" (PDF). Environmental Conservation. 24 (4): 338–343. doi:10.1017/S0376892997000441. S2CID   73633974.
  79. Devkota, B. P.; Silwal, T.; Shrestha, B. P.; Sapkota, A. P.; Lakhey, S. P. & Yadav, V. K. (2017). "Abundance of snow leopard (Panthera uncia) and its wild prey in Chhekampar VDC, Manaslu Conservation Area, Nepal". Banko Janakari. 27 (1): 11–20. doi: 10.3126/banko.v27i1.18545 .
  80. Leki; Thinley, P.; Rajaratnam, R. & Shrestha, R. (2018). "Establishing baseline estimates of blue sheep (Pseudois nayaur) abundance and density to sustain populations of the vulnerable snow leopard (Panthera uncia) in Western Bhutan". Wildlife Research. 45 (1): 38–46. doi:10.1071/WR16218. S2CID   89745062.
  81. Jamtsho, Y. & Katel, O. (2019). "Livestock depredation by snow leopard and Tibetan wolf: Implications for herders' livelihoods in Wangchuck Centennial National Park, Bhutan". Pastoralism. 9 (1): 1. doi: 10.1186/s13570-018-0136-2 .
  82. "Global Snow Leopard Conservation Forum". World Bank. Retrieved 2021-04-17.
  83. Wharton, D. & Freeman, H. (1988). "The Snow Leopard in North America: Captive Breeding Under the Species Survival Plan". In Freeman, H. (ed.). Proceedings of the Fifth International Snow Leopard Symposium. Seattle and Dehra Dun: International Snow Leopard Trust and Wildlife Institute of India. pp. 131–136.
  84. "Ladakh adopts State animal and bird". The Hindu. 2021-09-02. Retrieved 2021-10-18.
  85. "Project Snow Leopard". Himachal Pradesh Forest Department, Government of Himachal Pradesh. 2021. Retrieved 2021-10-08.

Further reading