The Green River Formation is a geological formation located in the Intermountain West of the United States, in the states of Colorado, Utah, and Wyoming. It comprises sediments deposited during the Early Eocene in a series of large freshwater lakes: Lake Gosiute, Lake Uinta, and Fossil Lake (the last containing Fossil Butte National Monument). It preserves a high diversity of freshwater fish, birds, reptiles, and mammals, with some sections of the formation (including Fossil Lake and the Parachute Creek member of Lake Uinta) qualifying as Konservat-Lagerstätten due to their extremely well-preserved fossils.
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Asterotrygon | A. maloneyi | Fossil Lake | A stingray of uncertain affinities. [1] | ||
Heliobatis | H. radians | A stingray of uncertain affinities. |
Primarily based on Grande (2001), with changes where necessary: [2]
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Crossopholis | C. magnicaudatus | Fossil Lake | A paddlefish. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Atractosteus | A. atrox | Fossil Lake | A gar related to the alligator gar. A. atrox is the largest of the Green River gar species. | ||
A. simplex | Fossil Lake, possibly Lake Gosiute | ||||
Cuneatus | C. cuneatus | Lake Gosiute, Lake Uinta, possibly Fossil Lake | A gar. | ||
C. wileyi | Lake Uinta | ||||
Lepisosteus | L. bemisi | Fossil Lake | A gar. [3] | ||
Masillosteus | M. janeae | Fossil Lake | A gar. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Amia | A. pattersoni | Fossil Lake, Lake Gosiute | An amiid related to modern bowfins. | ||
Cyclurus | C. gurleyi | Fossil Lake | Only 8 specimens known | An amiid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Eohiodon | E. falcatus | Fossil Lake | A hiodontid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Phareodus | P. encaustus | Fossil Lake, Lake Gosiute | A bonytongue. | ||
P. testis | Fossil Lake |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Diplomystus | D. dentatus | Fossil Lake, Lake Gosiute, & Lake Uinta | An armigatid ellimmichthyiform. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Gosiutichthys | G. parvus | Lake Gosiute | A clupeiform. | ||
Knightia | K. alta | Fossil Lake, Lake Gosiute & Lake Uinta | A clupeid. [4] The most commonly found articulated vertebrate fossil in the world. [5] | ||
K. eocaena | The most common fish of the formation. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Notogoneus | N. osculus | Fossil Lake | A beaked sandfish. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Amyzon | A. gosiutensis | Lake Gosiute | A sucker. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Astephus | A. antiquus | Lake Gosiute, Lake Uinta, Fossil Lake | Only one specimen known from Fossil Lake. | An ictalurid catfish. | |
Hypsidoris | H. farsonensis | Lake Gosiute | A hypsidorid catfish. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Esox | E. kronneri | Fossil Lake | Only a single specimen known. | A pickerel. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Amphiplaga | A. brachyptera | Fossil Lake | A relative of trout-perch. | ||
Erismatopterus | E. levatus | Lake Gosiute, Lake Uinta | A relative of trout-perch. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Mioplosus | M. labracoides | Fossil Lake, Lake Uinta | A relative of lates perches. [5] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Cockerellites | C. liops | Fossil Lake | A temperate bass. [5] | ||
Hypsiprisca | H. hypsacantha | Fossil Lake | A temperate bass. [5] | ||
H. sp. | |||||
Priscacara | P. serrata | Fossil Lake, Lake Uinta | A temperate bass. [5] | ||
P. sp. a | Fossil Lake | ||||
P. sp. b. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Asineops | A. squamifrons | Fossil Lake, Lake Gosiute, Lake Uinta | An acanthomorph of uncertain affinities, possibly a percopsiform, polymixiiform, or perciform. [6] | ||
A. sp. | Fossil Lake |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Aerugoamnis | A. paulus | Fossil Lake | Only two specimens known. | A relative of parsley frogs. [7] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Paleoamphiuma | P. tetradactylum | Fossil Lake | A sirenid salamander. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Afairiguana | A. avius | Fossil Lake | A stem-anole. [8] | ||
Bahndwivici | B. ammoskius | A relative of the Chinese crocodile lizard. | |||
Boavus | B. idelmani | A boa. | |||
Dunnophis | D. sp. | Possibly a dwarf boa. [9] | |||
Saniwa | S. ensidens | A stem-monitor lizard. | |||
Shinisauridae indet. | Lake Uinta (Parachute Creek) | A relative of the Chinese crocodile lizard, unusually preserved with a mineralized body without a skeleton. [10] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Borealosuchus | B. wilsoni | Fossil Lake | A eusuchian of uncertain affinities. | ||
Tsoabichi | T. greenriverensis | A caiman. [11] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Axestemys | A. sp. | Fossil Lake | A softshell turtle. | ||
Baptemys | B. wyomingensis | A dermatemydid. | |||
Chisternon | C. undatum | A baenid. | |||
Echmatemys | E. wyomingensis | A geoemydid. [12] | |||
Oliveremys | O. uintaensis | A softshell turtle. | |||
Plastomenus | P. thomasii | A softshell turtle. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Calciavis | C. grandei | Fossil Lake | A lithornithid. [13] | ||
Pseudocrypturus | P. cercanaxius | A lithornithid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Paakniwatavis | P. grandei | Fossil Lake | An anseriform of uncertain affinities. | ||
Presbyornis | P. pervetus | A presbyornithid anseriform. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Gallinuloides | G. wyomingensis | Fossil Lake | A gallinuloid galliform. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Anneavis | A. annae | Fossil Lake | A sandcoleid mousebird. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Plesiocathartes | P. major | Fossil Lake | A stem-cuckoo roller. [14] | ||
P. wyomingensis |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Paracoracias | P. occidentalis | A coraciiform. [15] | |||
Primobucco | P. mcgrewi | Fossil Lake | A primobucconid coraciiform. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Neanis | N. schucherti | Fossil Lake | A possible piciform of uncertain affinities. | ||
Pristineanis | P. kistneri | A possible piciform of uncertain affinities. [16] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Eocypselus | E. rowei | Fossil Lake | A eocypselid apodiform. [17] | ||
Fluvioviridavis | F. platyrhamphus | A basal strisore, possibly a stem-frogmouth. [18] [19] | |||
Prefica | P. nivea | A stem-oilbird. [20] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Foro | F. panarium | Fossil Lake | A stem-turaco. [21] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Juncitarsus | J. merkeli | Fossil Lake | A mirandornithine, possibly an early phoenicopteriform. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Limnofregata | L. azygosternon | Fossil Lake | An early frigatebird. | ||
L. hasegawai |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Vadaravis | V. brownae | Fossil Lake | Potentially a stem-ibis. [22] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
?Recurvirostridae indet. | Lake Uinta | A trackway of three footprints. | A shorebird and potentially the earliest record of recurvirostrids, known from only three footprints. [23] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Messelornis | M. nearctica | Fossil Lake | A messelornithid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Avolatavis | A. tenens | Fossil Lake | A quercypsittid. | ||
Cyrilavis | C. colburnorum | A halcyornithid. | |||
C. olsoni | |||||
Eofringillirostrum | E. boudreauxi | A psittacopedid. [24] | |||
Eozygodactylus | E. americanus | A zygodactylid. [25] | |||
Tynskya | T. eocaena | A messelasturid. | |||
Zygodactylus | Z. grandei | A zygodactylid. [26] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Nahmavis | N. grandei | Fossil Lake | Either a stem-gruiform or a stem-charadriiform. [27] |
Two other genera, Eoeurypyga (a stem-sunbittern) and Wyomingcypselus (an early apodiform) are mentioned only in a 2002 dissertation, [28] and are presently nomina nuda .
Partially based on Grande (1984). Aside from the few well-preserved mammals found in Fossil Lake, a majority of Green River mammals are based on isolated bones and teeth: [29]
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Herpetotherium | H. innominatum | Lake Gosiute & Lake Uinta | An opossum-like basal metatherian. Originally placed in Peratherium . | ||
H. knighti | Lake Gosiute |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Palaeosinopa | P. didelphoides | Fossil Lake | A pantolestid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Icaronycteris | I. gunnelli | Fossil Lake | 2 skeletons. [30] | An icaronycterid bat. | |
I. index | |||||
Onychonycteris | O. finneyi | An onychonycterid bat. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Nyctitherium | N. nitidum | Lake Gosiute | A nyctitheriid. Originally identified as Talpavus nitidus. | ||
N. sp. | Lake Uinta |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
" Miacis " | "M." gracilis | Lake Uinta | A carnivoraform of uncertain affinities. | ||
Sinopa | S. minor | Lake Uinta | A sinopid hyaenodont. | ||
Viverravus | V. minutus | Lake Uinta | A viverravid. | ||
Vulpavus | V. profectus | Lake Gosiute | A miacid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Hyopsodus | H. minusculus | Lake Gosiute & Lake Uinta | A hyopsodontid. | ||
H. paulus (=H. vicarius) | Lake Uinta | ||||
H. wortmani | Fossil Lake | ||||
Lambdotherium | L. popoagicum | Fossil Lake | A small-sized basal brontothere. | ||
Protorohippus | P. venticolus | 2 skeletons. | An equid. | ||
Tapiromorpha indet. | 1 skeleton | A juvenile tapiromorph, the largest mammal specimen known from the formation. Initially identified as a Heptodon calciculus. [31] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Apatemys | A. chardini | Fossil Lake | 1 skeleton. | An apatemyid. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Paramys | P. sp. | Lake Uinta | A ischyromyid rodent. | ||
Sciuravus | S. eucristadens | Lake Uinta | A sciuravid rodent. Originally identified as Viverravus eucristadens. | ||
S. sp. | |||||
Thisbemys | T. sp. | Lake Gosiute | A ischyromyid rodent. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Microsyops | M. elegans | Lake Gosiute | A microsyopid stem-primate. | ||
Notharctus | N. sp. | Lake Gosiute | A notharctid adapiform. | ||
Omomys | O. carteri (=O. pucillus) [32] | Lake Uinta | An omomyid primate. | ||
Smilodectes | S. gracilis | Lake Uinta | A notharctid adapiform. Formerly placed in Notharctus matthewi. | ||
Tetonius | T. sp. | Lake Uinta | An omomyid primate. | ||
Uintasorex | U. parvulus | Lake Gosiute | A microsyopid stem-primate. | ||
Washakius | W. insignis | Lake Gosiute | An omomyid primate. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Bechleja | B. rostrata | Fossil Lake | A palaemonid shrimp. [33] | ||
Cyclestherioides | C. wyomingensis | Lake Gosiute (Laney) | A clam shrimp. [34] | ||
Procambarus | P. primaevus | Fossil Lake | A cambarid crayfish. | ||
Prolynceus | P. laneyensis | Lake Gosiute (Laney) | A clam shrimp. [34] |
Primarily based on Bechly et al (2020): [35]
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Carlea | C. eocenica | Fossil Lake | A carleid damselfly. | ||
Cordulibellula | C. inopinata | Lake Uinta (Parachute Creek) | A cordulibellulid dragonfly, the last member of an ancient lineage. [36] | ||
Dysagrion | D. fredericii | Lake Gosiute (Laney) | A dysagrionid damselfly. | ||
D. integrum | Fossil Lake | ||||
D. lakesii | |||||
Eocalopteryx | E. atavina | A calopterygid damsefly. | |||
Eodysphaea | E. magnifica | Lake Uinta (Parachute Creek) | A euphaeid damselfly. | ||
Eolestes | E. syntheticus | An eolestid damselfly. | |||
Eopodagrion | E. scudderi | A damselfly of uncertain affinities. | |||
Epallagites | E. avus | A euphaeid damselfly. | |||
Kohlslibellula | K. lini | Lake Uinta (Parachute Creek) | A urolibellulid dragonfly | ||
Labandeiraia | L. americaborealis | Lake Uinta (Parachute Creek) | A euphaeid damselfly. | ||
L. riveri | Fossil Lake | ||||
Litheuphaea | L. coloradensis | Lake Uinta (Parachute Creek) | A euphaeid damselfly. | ||
Oreodysagrion | O. tenebris | Fossil Lake | A thaumatoneurid damselfly. | ||
Petrolestes | P. hendersoni | Lake Uinta (Parachute Creek) | A dysagrionid damselfly. | ||
Protamphipteryx | P. basalis | Fossil Lake | An amphipterygid damselfly. | ||
Tenebragrion | T. shermani | Fossil Lake | A dysagrionid damselfly. | ||
Tynskysagrion | T. brookeae | Fossil Lake | A thaumatoneurid damselfly. | ||
Urolibellula | U. eocaenica | Lake Uinta (Parachute Creek) | A urolibellulid dragonfly | ||
Viridiflumineagrion | V. aasei | Lake Uinta (Parachute Creek) | A "megapodagrionid" damselfly. [37] | ||
Zacallites | Z. balli | A zacallitid damselfly. | |||
Z. cockerelli | Fossil Lake |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Arethaea | A. solterae | Lake Uinta (Parachute Creek) | 1 specimen | A roundhead katydid. [38] |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Hersiliola | H. sp. | Lake Uinta (Parachute Creek) | A tree trunk spider. [39] | ||
Miagrammopes | M. sp. | A uloborid spider. [39] | |||
Selenopidae indet. | A selenopid spider. [39] | ||||
Thomisidae indet. | A crab spider. [39] | ||||
Uintascorpio | U. halandrasi | 1 specimen | A buthid scorpion. [40] | ||
Uloboridae indet. | A uloborid spider. [39] |
Based on Grande (1984): [29] [41]
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Plesielliptio | P. priscus | Lake Gosiute | A unionid river mussel. | ||
P. sp. "A" | |||||
Sphaerium | S. sp. | A fingernail clam. |
Genus | Species | Member | Abundance | Notes | Image |
---|---|---|---|---|---|
Biomphalaria | B. aequalis | Lake Gosiute | A ramshorn snail. | ||
B. storchi | |||||
B. pseudoammonius | |||||
? Drepanotrema | ?D. sp. | A ramshorn snail. | |||
Elimia | E. tenera | A pleurocerid, the most common fossil snail of the formation and the source of the famous "Turritella agate". | |||
Gastrocopta | G. sp. | A gastrocoptid land snail. [41] | |||
Gyraulus | G. militaris | A ramshorn snail. | |||
Holospira | H. sp. | A urocoptid land snail. [41] | |||
Hydrobia | H. aff. utaensis | A mud snail. | |||
H. sp. "A." | |||||
Lymnaea | L. similis | A pond snail. | |||
L. sp. "B." | |||||
Omalodiscus | O. cirrus | A ramshorn snail. | |||
Oreoconus | O. sp. "A." | A bulimulid land snail. | |||
Physa | P. bridgerensis | A bladder snail. | |||
P. longiuscula | |||||
P. pleromatis | |||||
P. sp. "A." | |||||
Pleurolimnaea | P. tenuicosta | A pond snail. | |||
Valvata | V. subumbilicata | A valve snail. | |||
V. cf. filosa | |||||
Viviparus | V. paludinaeformis | A river snail. | |||
V. trochiformis |
Vegavis is a genus of extinct bird that lived in Antarctica during the Maastrichtian stage of the Late Cretaceous. The type species is Vegavis iaai. Vegavis was initially considered to be a member of Anseriformes within Galloanserae, but this claim has not been supported by some recent studies.
The Lance (Creek) Formation is a division of Late Cretaceous rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas. The Lance Formation is Late Maastrichtian in age, and shares much fauna with the Hell Creek Formation of Montana and North Dakota, the Frenchman Formation of southwest Saskatchewan, and the lower part of the Scollard Formation of Alberta.
"Precursor" is a controversial prehistoric bird genus from the Early Eocene. It was established based on fossils found in England, including in the famous London Clay deposits. Three species are included in the genus: "P." parvus, the type species, "P." magnus, and "P." litorum, all named by Colin Harrison and Cyril Walker in 1977.
Fluvioviridavis is an extinct genus of bird from the Early Eocene Green River Formation of Wyoming and London Clay Formation of the United Kingdom. There are three known species: F. platyrhamphus from the Green River and F. michaeldanielsi and F. nazensis from the London Clay. Fluvioviridavis is the only genus currently named in the monotypic family Fluvioviridavidae.
Pangalliformes is the scientific name of a provisional clade of birds within the group Galloanserae. It is defined as all birds more closely related to chickens than to ducks, and includes all modern chickens, turkeys, pheasants, and megapodes, as well as extinct species that do not fall within the crown group Galliformes.
Dinosaur coloration is generally one of the unknowns in the field of paleontology, as skin pigmentation is nearly always lost during the fossilization process. However, recent studies of feathered dinosaurs and skin impressions have shown the colour of some species can be inferred through the use of melanosomes, the colour-determining pigments within the feathers.
Avolatavis is an extinct genus of stem-parrot (pan-psittaciform) or a member of the stem group of Psittacopasseres, known from the early Eocene Fossil Butte Member of the Green River Formation of Wyoming, United States, and from the London Clay of Walton-on-the-Naze. It was first named by Daniel T. Ksepka and Julia A. Clarke in 2012 and the type species is Avolatavis tenens. Gerald Mayr and Andrew C. Kitchener described the second species, A. europaeus, in 2023. Mayr and Kitchener assigned Avolatavis to the family Vastanavidae, which might be early diverging stem group presentatives of Pan-Psittaciformes or stem group representatives of Psittacopasseres.
The Willwood Formation is a sedimentary sequence deposited during the late Paleocene to early Eocene, or Clarkforkian, Wasatchian and Bridgerian in the NALMA classification.
Primobucco is an extinct genus of bird placed in its own family, Primobucconidae. The type species, Primobucco mcgrewi, lived during the Lower Eocene of North America. It was initially described by American paleo-ornithologist Pierce Brodkorb in 1970, from a fossil right wing, and thought to be an early puffbird. However, the discovery of a further 12 fossils in 2010 indicate that it is instead an early type of roller.
Vastanavidae is an extinct family of birds related to parrots and passerine birds. They are known from fossils from Eocene sites in India, Europe, and North America. The vastanavids resemble parrots and the extinct parrot relative Quercypsitta in their morphology, including the partially zygodactyl foot, in which two toes could face opposite the other two.
Vegaviidae is an extinct family of birds of uncertain phylogenetic placement, which existed during the Late Cretaceous and possibly the Paleocene. Definitive fossils attributed to the family have only been found in Antarctica, though other fossils from the Southern Hemisphere including Chile and New Zealand may represent this group. The putative Campanian vegaviid from Canada known as Maaqwi, more likely belongs to the Procellariformes.
Roger Lansing Grande, more commonly known as Lance Grande, is an evolutionary biologist and curatorial scientist. His research and work is focused on Paleontology, Ichthyology, Systematics and Evolution. He is well known for his work on the paleontology of the Green River Formation and for his detailed monographs on the comparative anatomy and evolution of ray-finned fishes. He has also published books on broader issues, engaging larger audiences on the importance of the natural and the social sciences.
Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.
Zygodactylidae is a family of extinct birds found in Europe and North America from the Eocene epoch to the Middle Miocene. First named in 1971, based on fragmentary remains of two species from Germany, a more complete description of the birds became possible in 2008 when a number of other, better-preserved fossil species were assigned to the family based on a number of shared characteristics.
Morsoravis is an extinct genus of neoavian bird from the Early Eocene Fur Formation of Denmark. It contains a single named species, Morsoravis sedilis. Fossils of Morsoravis have also been found in the Green River Formation of Wyoming and possibly the Nanjemoy Formation of Virginia.
Eocypselus rowei is an extinct bird believed to be ancestral to modern hummingbirds and swifts. It was a small bird, less than 5 in (13 cm) in length, and probably had black feathers. The bird was first described in 2013 and lived approximately 50 million years ago, during the Eocene epoch.
This is an overview of the paleofauna of the Eocene Messel Formation as explored by the Messel Pit excavations in Germany. A former quarry and now UNESCO World Heritage Site, the Messel Formation preserves what once were a series of anoxic lakes surrounded by a sub-tropical rainforest during the Middle Eocene, approximately 47 Ma.
Cyrilavis is an extinct genus of halcyornithid bird from the Early Eocene Fossil Butte Member of the Green River Formation, United States. The genus contains two species, Cyrilavis olsoni and Cyrilavis colburnorum.
Eofringillirostrum is an extinct species of bird known from the Early Eocene Green River Formation of the Western United States and from the Messel Pit in Germany. The genus contains two species, Eofringillirostrum boudreauxi and Eofringillirostrum parvulum. They are the earliest known fossil birds to have a finch-like beak.
Paakniwatavis is a genus of extinct waterfowl from the early Eocene Green River Formation of Wyoming, United States. The genus contains a single species, P. grandei, known from a partial skeleton with a skull.
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